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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L700	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon	Monophyllus plethodon		[MSW2] Includes luciae and frater; see Schwartz and Jones (1967:13). See Homan and Jones (19756, Mammalian Species, 58).; [MSW3] Includes luciae and frater; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones (1975b).; [HMW] Monophyllus plethodon G. S. Miller, 1900 , St. Michael Parish, Barbados , Lesser Antilles. Puerto Rican subspecies frater by H. E. Anthony in 1917 is known only from fossil material. Two subspecies recognized.; [batnames2022] Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975 b ).; [IUCN] Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003).; [batnames2023] Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975 b ).; [batnames2025_1.7] Includes luciae and frater; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975b).				luciae		frater, luciae.	plethodon, luciae	plethodon, frater, luciae		plethodon, luciae		plethodon, frater, luciae		plethodon, luciae, frater	Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003).	plethodon, frater, luciae		plethodon, luciae, frater 	plethodon, luciae, frater	frater, luciae, plethodon 		plethodon G. S. Miller, 1900|luciae G. S. Miller, 1902|frater H. E. Anthony, 1917		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Barbados long-tongued bat	Lesser Antilles, t Puerto Rico	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Monophyllus plethodon	Barbados (Lesser Antilles), St. Michael parish.	Miller	1900	Proc. Wash. Acad. Sei., 2:35.	Distribution: Living populations are confined to the Lesser Antilles, but the species is known fossil from Puerto Rico.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Barbados long-tongued bat (Insular long-tongued Rico (Insular long-tongued bat)	Lesser Antilles, f Puerto Rico	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Miller	1900	Proc. Washington Acad. Sci., 2:35.	Includes luciae and frater; see Schwartz and Jones (1967:13). See Homan and Jones (19756, Mammalian Species, 58).	Lesser Antilles from Anguilla to St. Vincent and Barbados. Fossils known from Puerto Rico.	Barbados (Lesser Antilles), St. Michael Parish.		MILLER	1900	Upper premolars separated by a diastema less than half the length of the anterior upper premolar. Size relatively large (forearm length, 38-46 mm; condylobasal length, 19-23 mm).	Distribution: Living populations are confined to the Lesser Antilles, but the species is known fossil from Puerto Rico.	Two living subspecies:	M. p. plethodon (Barbados), M. p. luciae (remaining Lesser Antilles from St. Vincent to Anguilla).	81	species	M. plethodon	MILLER	1900	Monophyllus	genus	Monophyllus plethodon				Upper premolars separated by a diastema less than half the length of the anterior upper premolar. Size relatively large (forearm length, 38-46 mm; condylobasal length, 19-23 mm).	Two living subspecies:		1. M. plethodon MILLER 1900.	1	_M. p. frater_ Anthony, 1917; _M. p. luciae_ Miller, 1902; _M. p. plethodon_ Miller, 1900			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Glossophagini	Monophyllus plethodon	Monophyllus		plethodon	Miller		1900		Proc. Washington Acad. Sci.	2		35		Insular Single-leaf Bat	Barbados (Lesser Antilles), St. Michael Parish.	Lesser Antilles from Anguilla to St. Vincent and Barbados. Fossils known from Puerto Rico.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	frater Anthony, 1917; luciae Miller, 1902.	Includes luciae and frater; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones (1975b).	03A687BCFFAAFFAA13BCFD8FFAC0FB13	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	511	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFAAFFAA13BCFD8FFAC0FB13.xml	Monophyllus plethodon	Phyllostomidae	Monophyllus	plethodon	G. S. Miller	1900	Monophylle des Petites Antilles @fr | Kleine-Antillen-Blattnase @de | Mondfilode las Pequenas Antillas @es | Other common names @en | nsular Long-tongued Bat @en | Lesser Antillean Long-tongued Bat @en	Monophyllus plethodon G. S. Miller, 1900 , St. Michael Parish, Barbados , Lesser Antilles. Puerto Rican subspecies frater by H. E. Anthony in 1917 is known only from fossil material. Two subspecies recognized.	M.p.plethodonG.S.Miller,1900—Barbados. M. p. luciae G. S. Miller, 1902 — Lesser Antilles, from Anguilla to Saint Vincent Is.	Head-body 67-84 mm, tail 8-16 mm, ear 14-15 mm, hindfoot 12-13 mm, forearm 38-8-45-7 mm; weight 12-5-17-2 g. The Insular Single-leaf Batis is easily distinguished from other Antillean bat species based on long narrow snout, ending in small but clearly defined noseleaf. Tongue is exceedingly long. Ears appear proportional to head and are well separated over it. Fur is brownish to pale buffy tan. Dental formula of all species of Monophyllusis12/2, C1/1,P 2/3, M 3/3 ( x2 ) = 34. Teeth are very small, with distorted W-pattern. Upper premolars are relatively crowded and separated by diastemaless than one-half the length of P°. Testes are 2-9-5 mm.	Xeric areas as well as dense rainforests. Most often captured in or near fruit plantations, especially in banana groves. In Barbados , an individual was caught in a mist net stretched between buttresses under a bridge near a sugarcane field. The Insular Single-leaf Bat is not equally abundant throughout its distribution, perhaps because smaller populations occur on islands with more recent impact from hurricanes.	No information, but probably similar to Leach’s Single-leaf Bat ( M. redmani ), which feeds on nectar.	Pregnant Insular Single-leaf Bats have been captured in March-April, lactating females in April-May and July, and post-lactating females in June-July. Females give birth to one young at a time.	Based on its diet and close relationship to Leach’s Single-leaf Bat, the Insular Single-leaf Bat is expected to be nocturnal, beginning its activity well after sunset. It roosts in caves.	Most colonies of Insular Single-leaf Bats appear to be rather small, numbering hundreds of individuals. Small size of colonies and islands where it is found suggest that it disperses over short distances.	Classified as Least Concern on The IUCNRed List. The Insular Single-leaf Bat is presumably common throughoutits restricted distribution. Its habitat is not considered to be as declining.	Anthony (1917) | Genoways, Larsen, PA. et al. (2007) | Genoways, Larsen, R.J. et al. (2011) | Genoways, Pedersen et al. (2007) | Grindal (2004) | Homan & Jones (1975b) | Larsen et al. (2006) | Masson et al. (1990) | Pedersen, Genoways, Morton, Johnson & Courts (2003) | Pedersen, Genoways, Morton, Kwiecinski & Courts (2005) | Pedersen, Larsen et al. (2018) | Rodriguez-Duran & Davalos (2018) | Timm & Genoways (2003)	https://zenodo.org/record/6458696/files/figure.png	47. Insular Single-leat Bat Monophyllus plethodon French: Monophylle des Petites Antilles / German: Kleine-Antillen-Blattnase / Spanish: Mondfilo de las Pequenas Antillas Other common names: Insular Long-tongued Bat , Lesser Antillean Long-tongued Bat Taxonomy. Monophyllus plethodon G. S. Miller, 1900 , St. Michael Parish, Barbados , Lesser Antilles. Puerto Rican subspecies frater by H. E. Anthony in 1917 is known only from fossil material. Two subspecies recognized. Subspecies and Distribution. M.p.plethodonG.S.Miller,1900—Barbados. M. p. luciae G. S. Miller, 1902 — Lesser Antilles, from Anguilla to Saint Vincent Is. Descriptive notes. Head-body 67-84 mm, tail 8-16 mm, ear 14-15 mm, hindfoot 12-13 mm, forearm 38-8-45-7 mm; weight 12-5-17-2 g. The Insular Single-leaf Batis is easily distinguished from other Antillean bat species based on long narrow snout, ending in small but clearly defined noseleaf. Tongue is exceedingly long. Ears appear proportional to head and are well separated over it. Fur is brownish to pale buffy tan. Dental formula of all species of Monophyllusis12/2, C1/1,P 2/3, M 3/3 ( x2 ) = 34. Teeth are very small, with distorted W-pattern. Upper premolars are relatively crowded and separated by diastemaless than one-half the length of P°. Testes are 2-9-5 mm. Habitat. Xeric areas as well as dense rainforests. Most often captured in or near fruit plantations, especially in banana groves. In Barbados , an individual was caught in a mist net stretched between buttresses under a bridge near a sugarcane field. The Insular Single-leaf Bat is not equally abundant throughout its distribution, perhaps because smaller populations occur on islands with more recent impact from hurricanes. Food and Feeding. No information, but probably similar to Leach’s Single-leaf Bat ( M. redmani ), which feeds on nectar. Breeding. Pregnant Insular Single-leaf Bats have been captured in March-April, lactating females in April-May and July, and post-lactating females in June-July. Females give birth to one young at a time. Activity patterns. Based on its diet and close relationship to Leach’s Single-leaf Bat, the Insular Single-leaf Bat is expected to be nocturnal, beginning its activity well after sunset. It roosts in caves. Movements, Home range and Social organization. Most colonies of Insular Single-leaf Bats appear to be rather small, numbering hundreds of individuals. Small size of colonies and islands where it is found suggest that it disperses over short distances. Status and Conservation. Classified as Least Concern on The IUCNRed List. The Insular Single-leaf Bat is presumably common throughoutits restricted distribution. Its habitat is not considered to be as declining. Bibliography. Anthony (1917), Genoways, Larsen, PA. et al. (2007), Genoways, Larsen, R.J. et al. (2011), Genoways, Pedersen et al. (2007), Grindal (2004), Homan & Jones (1975b), Larsen et al. (2006), Masson et al. (1990), Pedersen, Genoways, Morton, Johnson & Courts (2003), Pedersen, Genoways, Morton, Kwiecinski & Courts (2005), Pedersen, Larsen et al. (2018), Rodriguez-Duran & Davalos (2018), Timm & Genoways (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Monophyllus plethodon	Monophyllus		plethodon	Miller	1900	0	Proc. Washington Acad. Sci.	2:35	Insular Single-leaf Bat	<b> frater </b> Anthony, 1917; <b> luciae </b> Miller, 1902.	Barbados (Lesser Antilles), St. Michael Parish.	Lesser Antilles from Anguilla to St. Vincent and Barbados. Fossils known from Puerto Rico.	Not listed.	Least Concern	Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975 b ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Monophyllus plethodon	23	Insular Single-leaf Bat	Insular Long-tongued Bat|Lesser Antillean Long-tongued Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLOSSOPHAGINAE	GLOSSOPHAGINI	Monophyllus	NA	plethodon	G. S. Miller	1900	0	Monophyllus_plethodon	Miller, G. S. , Jr. (1900). The Bats of the Genus Monophyllus. Proceedings of the Washington Academy of Sciences, 2, 35.	https://repository.si.edu/bitstream/handle/10088/74399/Miller%20Bats%20Genus%20Monophyllus.pdf?sequence=1&isAllowed=y	USNM 101530		St. Michael Parish, Barbados, Lesser Antilles.			plethodon G. S. Miller, 1900|luciae G. S. Miller, 1902|frater Anthony, 1917	NA	NA	Anguilla|Antigua & Barbuda|Saint Barthélemy|Saint Kitts & Nevis|Saint Martin|Montserrat|Guadeloupe|Dominica|Martinique|Saint Lucia|Saint Vincent & the Grenadines|Barbados	North America	Neotropic	LC	0	0	0	Monophyllus_plethodon	0	sciname match	Monophyllus_plethodon	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13719	Monophyllus plethodon	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Monophyllus	plethodon	Miller, 1900	Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003).	20000000	Monophyllus plethodon	Least Concern		2018	2018-03-01 00:00:00 UTC	3.1	English	Listed as Least Concern because its presumed large population extended over several small islands, and because its habitat is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species has been collected in a variety of situations. The area where it occurs was described as a steep and wooded ravine near sugar cane fields; also it was trapped in banana groves, across a trail that separated woodlands from a Theobroma (cacao) ;grove, and in dense rainforest. It roosts in caves. Females taken on Dominica between March and April were gravid (Homan and Jones 1975); pregnant females found in January (Pedersen et al . 1996) on Montserrat.	Threats to this species include mining and tourism in the caves. Severe weather (hurricanes) can affect small populations on very small islands.	There is no detailed information on the status of its populations. Pedersen et al . (1996) compared the number of captures of M. plethodon in Montserrat before (1984) and after (1986) the hurricane Hugo, finding a decrease of over 60% of bats netted. In general, frugivores were the most affected guild on the island. No similar data exist for other islands, but this points out to the effect of severe weather on small populations of bats on small Caribbean islands.	Unknown	This species is known from Lesser Antilles from Anguilla to St. Vincent and Barbados (Pedersen et al . 2013). There are fossils records from Puerto Rico (Simmons 2005), but no recent records on the island (A. Rodriguez-Herrera pers. comm.). It is known from sea level to 550 m (Schwartz and Jones 1967).		Terrestrial	The species is found in nature reserves and protected areas.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Monophyllus		plethodon	Miller	1900	0	Proc. Washington Acad. Sci.	2:35	Insular Single-leaf Bat	<b> frater </b> Anthony, 1917; <b> luciae </b> Miller, 1902.	Barbados (Lesser Antilles), St. Michael Parish.	Lesser Antilles from Anguilla to St. Vincent and Barbados. Fossils known from Puerto Rico.	Not listed.	Least Concern	Includes luciae and frater ; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975 b ).	Monophyllus plethodon	1004917	23	Insular Single-leaf Bat	Insular Long-tongued Bat|Lesser Antillean Long-tongued Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLOSSOPHAGINAE	GLOSSOPHAGINI	Monophyllus	NA	plethodon	G. S. Miller	1900	0	Monophyllus_plethodon	Miller, G. S. , Jr. (1900). The Bats of the Genus Monophyllus. Proceedings of the Washington Academy of Sciences, 2, 35.	https://repository.si.edu/bitstream/handle/10088/74399/Miller%20Bats%20Genus%20Monophyllus.pdf?sequence=1&isAllowed=y	USNM 101530		St. Michael Parish, Barbados, Lesser Antilles.			plethodon G. S. Miller, 1900|luciae G. S. Miller, 1902|frater Anthony, 1917	NA	NA				Anguilla|Antigua & Barbuda|Saint Barthélemy|Saint Kitts & Nevis|Saint Martin|Montserrat|Guadeloupe|Dominica|Martinique|Saint Lucia|Saint Vincent & the Grenadines|Barbados	North America	Neotropic	LC	0	0	0	Monophyllus_plethodon	0	sciname match	Monophyllus_plethodon	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Monophyllus_plethodon	1004917	23	Eastern Antillean Single-leaf Bat	Insular Long-tongued Bat|Lesser Antillean Long-tongued Bat|Insular Single-leaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glossophaginae	Glossophagini	Monophyllus	NA	plethodon	G. S. Miller	0	Monophyllus plethodon	Miller, G.S., Jr. 1900-03-30. The bats of the genus _Monophyllus_. Proceedings of the Washington Academy of Sciences 2:31-38.	https://www.biodiversitylibrary.org/page/8872039	USNM:MAMM:101530	holotype	http://n2t.net/ark:/65665/3d9b8f53d-8f50-4edf-b424-5cf6a9d50d3e	St. Michael Parish, Barbados, Lesser Antilles.			NA	NA				Anguilla|Antigua and Barbuda|Saint Barthélemy|Saint Kitts and Nevis|Sint Maarten|Montserrat|Guadeloupe|Dominica|Martinique|Saint Lucia|Saint Vincent and the Grenadines|Barbados	North America	Neotropic	LC	0	0	0	Monophyllus_plethodon	0	sciname match	Monophyllus_plethodon	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Monophyllus		plethodon	Miller	1900	0	Proc. Washington Acad. Sci.	2:35	Insular Single-leaf Bat	frater Anthony, 1917; luciae Miller, 1902.	Barbados (Lesser Antilles), St. Michael Parish.	Lesser Antilles from Anguilla to St. Vincent and Barbados. Fossils known from Puerto Rico.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13719/22112320/' target='_blank'>Least Concern</a>	Includes luciae and frater; see Schwartz and Jones (1967), Hall (1981), and Timm and Genoways (2003). See Homan and Jones(1975b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Monophyllus plethodon; Monophyllus plethodon; Monophyllus plethodon; Monophyllus plethodon; Monophyllus plethodon; Monophyllus plethodon; plethodon; frater; luciae; plethodon; luciae; frater; luciae; plethodon; luciae; frater; Monophylle des Petites Antilles; Kleine-Antillen-Blattnase; Mondfilode las Pequenas Antillas; Other common names; nsular Long-tongued Bat; Lesser Antillean Long-tongued Bat; Insular Single-leaf Bat; Insular Long-tongued Bat; Lesser Antillean Long-tongued Bat; Insular Single-leaf Bat; Insular Single-leaf Bat; M. plethodon