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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L658	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Miniopterus minor [synonym of]	Miniopterus minor newtoni	Miniopterus minor newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni	Miniopterus newtoni		[HMW] Miniopterus newtoni Bocage, 1889 , “St. Thomé [= Sao Tomé Island ].” Miniopterus newton : was previously considered a subspecies of M. minor . Monotypic.; [batnames2022] Distinct from minor; see Juste et al. (2007).; [MDD2022] split from M. minor; moved from Vespertilionidae to Miniopteridae; [IUCN] Recently elevated from synonymy with Miniopterus minor by Juste et al. (2007).; [batnames2023] Distinct from minor; see Juste et al. (2007).; [MDD2023] split from M. minor; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] split from M. minor; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] Distinct from minor; see Juste et al. (2007).; [MDD2025_2.2] split from M. minor; moved from Vespertilionidae to Miniopteridae														newtoni	Recently elevated from synonymy with Miniopterus minor by Juste et al. (2007).			newtoni 	newtoni 			newtoni du Bocage, 1889|newtonii de Seabra, 1900 [incorrect subsequent spelling]						N/A																																								NA																											E84887F9FFDAD6550F2FF66C159D359D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	703	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFDAD6550F2FF66C159D359D.xml	Miniopterus newtoni	Miniopteridae	Miniopterus	newtoni	Bocage	1889	Minioptére de Sao Tomé @fr | Newton-Langflligelfledermaus @de | Minidptero de Newton @es | Newton's Bent-winged Bat @en	Miniopterus newtoni Bocage, 1889 , “St. Thomé [= Sao Tomé Island ].” Miniopterus newton : was previously considered a subspecies of M. minor . Monotypic.	Endemic to Sao Tomé I.	Head—body 45 mm , tail 43 mm , ear 9-5-10 mm, hindfoot 9 mm , forearm 38-40 mm; weight 7 g . Newton's Long-fingered Bat has blackish gray dorsal and ventral pelage. There is also a reddish morph, although less abundant. It has narrow hairy edge along dorsal side of wing membrane in area between femur and humerus. Lower part of this membrane has few and sparse hairs, and uropatagium is hairless on both sides. Ears are small, and tragusis relatively long (4: 5 mm ) and narrow, with rounded tip. On following pages: 21. Least Long-fingered Bat ( Miniopterus minon; 22. Sandy Long-fingered Bat ( Miniopterus arenarius); 23. African Long-fingered Bat ( Miniopterus africanus ); 24. Natal Long-fingered Bat ( Miniopterus natalensis ); 25. Mozambican Long-fingered Bat ( Miniopterus mossambicus ); 26. Lesser Long-fingered Bat ( Miniopterus fraterculus ); 27. Aellen’s Long-fingered Bat ( Miniopterus aelleni ); 28. Griveaud’s Long-fingered Bat ( Miniopterus griveaud)); 29. Glen's Long-fingered Bat ( Miniopterus gleni ); 30. Madagascar Long-fingered Bat ( Miniopterus brachytragos ); 31. Montagne dAmbre Long-fingered Bat ( Miniopterus ambohitrensis); 32. Major's Long-fingered Bat ( Miniopterus majori ); 33. Eger’s Long-fingered Bat ( Miniopterus egeri ); 34. Manavil Long-fingered Bat ( Miniopterus manavi ); 35. Sororcula Long-fingered Bat ( Miniopterus sororculus ); 36. Peterson's Long-fingered Bat ( Miniopterus petersoni ); 37. Mahafaly Long-fingered Bat ( Miniopterus mahafaliensis ); 38.	Various habitats including non-altered forests and anthropogenic environments (e.g. cocoa plantations and around streetlights in urban areas) at elevations of 0-1300 m.	There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight.	No information.	Presumably nocturnal. It mainly roosts in caves but also mines and tunnels. Echolocation calls have downward FM signals, with maximum frequencies of 69-6-119 kHz, minimum frequencies of 47-8-52-2 kHz, peak frequencies of 53-2— 61 kHz, durations of 2-3-7-8 milliseconds, and intervals of 31-8-105-7 milliseconds.	Newton's LLong-fingered Bats form colonies with thousands of individuals and share roosts with other species of bats such as Noack’s Leaf-nosed Bat (Hipposideros ruber) and Temminck’s Myotis ( Myotis tricolor).	Classified as Data Deficient on The IUCN Red List. There is little information about population status, ecology, and distribution of Newton's Longfingered Bat, and conservation threats are unknown.	Bocage (1889) | Juste & Ibanez (1992) | Juste et al. (2007) | Rainho et al. (2010)	https://zenodo.org/record/5735262/files/figure.png	20. Newton's Long-fingered Bat Miniopterus newtoni French: Minioptére de Sao Tomé / German: Newton-Langflligelfledermaus / Spanish: Minidptero de Newton Other common names: Newton's Bent-winged Bat Taxonomy. Miniopterus newtoni Bocage, 1889 , “St. Thomé [= Sao Tomé Island ].” Miniopterus newton : was previously considered a subspecies of M. minor . Monotypic. Distribution. Endemic to Sao Tomé I. Descriptive notes. Head—body 45 mm , tail 43 mm , ear 9-5-10 mm, hindfoot 9 mm , forearm 38-40 mm; weight 7 g . Newton's Long-fingered Bat has blackish gray dorsal and ventral pelage. There is also a reddish morph, although less abundant. It has narrow hairy edge along dorsal side of wing membrane in area between femur and humerus. Lower part of this membrane has few and sparse hairs, and uropatagium is hairless on both sides. Ears are small, and tragusis relatively long (4: 5 mm ) and narrow, with rounded tip. On following pages: 21. Least Long-fingered Bat ( Miniopterus minon; 22. Sandy Long-fingered Bat ( Miniopterus arenarius); 23. African Long-fingered Bat ( Miniopterus africanus ); 24. Natal Long-fingered Bat ( Miniopterus natalensis ); 25. Mozambican Long-fingered Bat ( Miniopterus mossambicus ); 26. Lesser Long-fingered Bat ( Miniopterus fraterculus ); 27. Aellen’s Long-fingered Bat ( Miniopterus aelleni ); 28. Griveaud’s Long-fingered Bat ( Miniopterus griveaud)); 29. Glen's Long-fingered Bat ( Miniopterus gleni ); 30. Madagascar Long-fingered Bat ( Miniopterus brachytragos ); 31. Montagne dAmbre Long-fingered Bat ( Miniopterus ambohitrensis); 32. Major's Long-fingered Bat ( Miniopterus majori ); 33. Eger’s Long-fingered Bat ( Miniopterus egeri ); 34. Manavil Long-fingered Bat ( Miniopterus manavi ); 35. Sororcula Long-fingered Bat ( Miniopterus sororculus ); 36. Peterson's Long-fingered Bat ( Miniopterus petersoni ); 37. Mahafaly Long-fingered Bat ( Miniopterus mahafaliensis ); 38. Griffiths's Long-fingered Bat ( Miniopterus griffithsi ) Habitat. Various habitats including non-altered forests and anthropogenic environments (e.g. cocoa plantations and around streetlights in urban areas) at elevations of 0-1300 m. Food and Feeding. There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight. Breeding. No information. Activity patterns. Presumably nocturnal. It mainly roosts in caves but also mines and tunnels. Echolocation calls have downward FM signals, with maximum frequencies of 69-6-119 kHz, minimum frequencies of 47-8-52-2 kHz, peak frequencies of 53-2— 61 kHz, durations of 2-3-7-8 milliseconds, and intervals of 31-8-105-7 milliseconds. Movements, Home range and Social organization. Newton's LLong-fingered Bats form colonies with thousands of individuals and share roosts with other species of bats such as Noack’s Leaf-nosed Bat (Hipposideros ruber) and Temminck’s Myotis ( Myotis tricolor). Status and Conservation. Classified as Data Deficient on The IUCN Red List. There is little information about population status, ecology, and distribution of Newton's Longfingered Bat, and conservation threats are unknown. Bibliography. Bocage (1889), Juste & Ibanez (1992), Juste et al. (2007), Rainho et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Miniopteridae	Miniopterus newtoni	Miniopterus		newtoni	Bocage	1889	0	Jornal de Sciencias Mathematicas, Physicas e Naturaes, Lisboa	0.2208	Newton's Long-fingered Bat	None.		Sao Tom&eacute;.	Not listed.	Data Deficient	Distinct from minor; see Juste et al. (2007).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus newtoni	23	Newton's Long-fingered Bat	Newton's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	newtoni	Bocage	1889	0	Miniopterus_Newtoni	du Bocage, J. V. B. (1889). ChiroptÃ¨res de l'Ã®le St. ThomÃ©. Jornal de sciencias mathematicas, physicas e naturaes, ser. 2, 1, 198.	https://www.biodiversitylibrary.org/item/23044#page/212/mode/1up	EBD 17350		"St. ThomÃ© [= SÃ£o TomÃ© Island]."			newtoni Bocage, 1889	split from M. minor; moved from Vespertilionidae to Miniopteridae	Juste, J., FerrÃ¡ndez, A., Fa, J. E., Masefield, W., & IbÃ¡Ã±ez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao TomÃ©, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.	SÃ£o TomÃ© & PrÃ­ncipe	Africa	Afrotropic	DD	0	0	0	Miniopterus_newtoni	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136310	Miniopterus newtoni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MINIOPTERIDAE	Miniopterus	newtoni	Bocage, 1889	Recently elevated from synonymy with Miniopterus minor by Juste et al. (2007).	20000000	Miniopterus newtoni	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	The species is confirmed as Data Deficient in view of the absence of sufficient information on its extent of occurrence, natural history, threats and conservation status.	The species appears to be quite common both in forest and plantations (Javier Juste pers. comm. 2008). A particular cave in Santa Catarina (north) had a quite impressive population (several thousands) that was near the 'city' and could be particularly sensitive to human disturbances (Javier Juste pers. comm. 2008).	The threats to this species are not well known.	It is a common species (Juste et al. 1994).	Unknown	This species is endemic to the island of SÃ£o TomÃ© (SÃ£o TomÃ© and PrÃ­ncipe) where it is widespread (Juste et al. 2007).		Terrestrial	It is not known if the species is present in any protected areas. Further studies are needed into the natural history and threats to this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		newtoni	Bocage	1889	0	Jornal de Sciencias Mathematicas, Physicas e Naturaes, Lisboa	0.220833	Newton's Long-fingered Bat	None.		Sao Tom&eacute;.	Not listed.	Data Deficient	Distinct from minor; see Juste et al. (2007).	Miniopterus newtoni	1005127	23	Newton's Long-fingered Bat	Newton's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	newtoni	Bocage	1889	0	Miniopterus_Newtoni	du Bocage, J. V. B. (1889). ChiroptÃ¨res de l'Ã®le St. ThomÃ©. Jornal de sciencias mathematicas, physicas e naturaes, ser. 2, 1, 198.	https://www.biodiversitylibrary.org/item/23044#page/212/mode/1up	EBD 17350		"St. ThomÃ© [= SÃ£o TomÃ© Island]."			newtoni Bocage, 1889	split from M. minor; moved from Vespertilionidae to Miniopteridae	Juste, J., FerrÃ¡ndez, A., Fa, J. E., Masefield, W., & IbÃ¡Ã±ez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao TomÃ©, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				SÃ£o TomÃ© & PrÃ­ncipe	Africa	Afrotropic	DD	0	0	0	Miniopterus_newtoni	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_newtoni	1005127	23	Newton's Long-fingered Bat	Newton's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	newtoni	du Bocage	0	Miniopterus Newtoni	Bocage, J.V.B. du. 1889-12. ChiroptÃ¨res de l'Ã®le St. ThomÃ©. Jornal de sciencias mathematicas, physicas e naturaes (2)1(3):197-199.	https://www.biodiversitylibrary.org/page/3418951	EBD 17.350	neotype		"St. ThomÃ© [= SÃ£o TomÃ© Island]."	0.266667	6.483333	split from M. minor; moved from Vespertilionidae to Miniopteridae	Juste, J., FerrÃ¡ndez, A., Fa, J. E., Masefield, W., & IbÃ¡Ã±ez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao TomÃ©, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				SÃ£o TomÃ© and PrÃ­ncipe	Africa	Afrotropic	DD	0	0	0	Miniopterus_newtoni	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		newtoni	Bocage	1889	0	Jornal de Sciencias Mathematicas, Physicas e Naturaes, Lisboa	0.220833	Newton's Long-fingered Bat	None.		Sao Tom&eacute;.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136310/22019007/' target='_blank'>Data Deficient</a>	Distinct from minor; see Juste et al. (2007).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus newtoni; Miniopterus newtoni; Miniopterus newtoni; Miniopterus newtoni; Miniopterus newtoni; newtoni; Minioptére de Sao Tomé; Newton-Langflligelfledermaus; Minidptero de Newton; Newton's Bent-winged Bat; Newton's Long-fingered Bat; Newton's Bent-winged Bat; Newton's Long-fingered Bat; M. newtoni
