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authorityParentheses originalNameCombination authoritySpeciesCitation authoritySpeciesLink typeVoucher typeKind typeVoucherURIs typeLocality typeLocalityLatitude typeLocalityLongitude taxonomyNotes taxonomyNotesCitation distributionNotes distributionNotesCitation subregionDistribution countryDistribution continentDistribution biogeographicRealm iucnStatus extinct domestic flagged CMW_sciName diffSinceCMW MSW3_matchtype MSW3_sciName diffSinceMSW3 docOrigin_batnames2025 Family Genus Subgenus Species Author Date Parentheses (1=author & date in parentheses) Citation Pages Common Name Synonyms Type Locality Distribution CITES IUCN Comments column3781 column3791 subtribe CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L653 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A Miniopterus minor [synonym of] Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi Miniopterus manavi [MSW3] Formerly included in minor, but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Includes griveaudi; see Peterson et al. (1995).; [HMW] Miniopterus manavi Thomas, 1906 , “Imasindrary, N.E. Betsileo,” Madagascar . Miniopterus manavi has been considered the common species of Miniopterus , with the widest distribution in Madagascar . Nevertheless, recent genetic studies have shown it to represent a species complex with at least six species, several of them not even phylogenetically related. Some recognized characteristics of the life history of M. manavi date back to before the group was split, which makes it difficult to assign them to specific lineages so they are omitted here. Monotypic.; [batnames2022] Formerly included in minor , but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi ; see Juste et al. (2007), but see also Peterson et al. (1995).; [MDD2022] previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae; [batnames2023] Formerly included in minor , but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi ; see Juste et al. (2007), but see also Peterson et al. (1995).; [MDD2023] previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] Formerly included in minor, but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi; see Juste et al. (2007), but see also Peterson et al. (1995).; [MDD2025_2.2] previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae manavi, grivaudi manavi, griveaudi manavi manavi manavi manavi O. Thomas, 1906 N/A Distribution: Confined to Madagascar and the Comoro islands. THOMAS 1906 Size relatively small (forearm length, 32-40 mm; condylobasal length, 12-14 mm). Braincase not markedly inflated. Rostrum relatively slender. Distribution: Confined to Madagascar and the Comoro islands. Two subspecies are recognized: M. m. manavi (Madagascar), M. m. grivaudi (Comoros). 133 species M. manavi THOMAS 1906 Miniopterus genus Miniopterus manavi Size relatively small (forearm length, 32-40 mm; condylobasal length, 12-14 mm). Braincase not markedly inflated. Rostrum relatively slender. Two subspecies are recognized: 1. M. manavi THOMAS 1906 [australis group], 1 NA Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Vespertilionidae Miniopterinae Miniopterus manavi Miniopterus manavi Thomas 1906 Ann. Mag. Nat. Hist., ser. 7 17 176 Manavi Long-fingered Bat Madagascar, E/NE of Betsileo, 20°17’S, 47°31’E [Fandriana region]. Madagascar, Comoro Isls. IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient as M. menavi (misspelled). griveaudi Harrison, 1959. Formerly included in minor, but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Includes griveaudi; see Peterson et al. (1995). E84887F9FFC7D6490FD7FBD717103DEF Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Miniopteridae_674.pdf.imf hash://md5/1471ff81ffd6d6580a4affec112f3619 708 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFC7D6490FD7FBD717103DEF.xml Miniopterus manavi Miniopteridae Miniopterus manavi Thomas 1906 Minioptére des Betsileo @fr | Zentral-Madagaskar-Langflligelfledermaus @de | Minidptero de Betsileo @es | Manavil Bent-winged Bat @en Miniopterus manavi Thomas, 1906 , “Imasindrary, N.E. Betsileo,” Madagascar . Miniopterus manavi has been considered the common species of Miniopterus , with the widest distribution in Madagascar . Nevertheless, recent genetic studies have shown it to represent a species complex with at least six species, several of them not even phylogenetically related. Some recognized characteristics of the life history of M. manavi date back to before the group was split, which makes it difficult to assign them to specific lineages so they are omitted here. Monotypic. Restricted to highlands of C Madagascar from Fandriana S to Ranomafana (Ifanadiana) and Vinanitelo. Head—body c. 51 mm , tail 39 mm , ear 10 mm , hindfoot 6 mm , forearm 37-6-39-2 mm; weight 6-4 g (single individual except for forearm length). Dorsal hairs of the Manavil Long-fingered Bat are of medium length (c. 5 mm ). General color is blackish, with hairs blackish along two-thirds their length and having dull brown tips. Underparts are similar but rather paler. Some specimens are dull reddish throughout, representing a rufous morph, but others have coal black backs and slightly lighter bellies, which is best considered a reddish brown red morph. Wing and tail membranes are dark brownish black. Uropatagium and plagiopatagium are attached to femur at same level above anklejoint. Fur extends thinly over nearly one-half of upper surface of uropatagium, being sparser on proximal ventral surface. Tragus ( 6 mm ) is relatively thin along distal two-thirds ofits length, medial margin has a flange, distal lateral portion is slightly enlarged and rounded, and distal medial tip terminates with an angular straight edge. Continuous forests, irrigated lowland rice paddies, secondary vegetation, and hillside rice at elevations of ¢.900-1500 m. In Ranomafana National Park, according to bat ultrasound recorders, the Manavil Long-fingered Bat seems to occur more frequently in forest fragments. Diet mainly contains Lepidoptera , Hymenoptera, Coleoptera, and Hemiptera , with lesser amounts of Blattodea, Diptera, Ephemeroptera , and Neuroptera . Prey also included mosquitoes that serve as vectors of diseases such as Anopheles squamosus (Malaria, Rift Valley fever virus [RVFV]) and Wuchereria bancrofti and Coquillettidia sp. (RVFV, West Nile virus [WNV]). Other prey is possibly fruit pests (e.g. Pandemis sp.). No information. The Manavil Long-fingered Bat is nocturnal. It uses caves as daytime shelters. Echolocation calls have downward FM signals, with maximum frequencies of 89-110 kHz, minimum frequency of 53 kHz, peak frequencies of 55-5-58-2 kHz, durations of 2-1-3 milliseconds, and intervals of 54-1-84-7 milliseconds. No information. Classified as Least Concern on The IUCN Red Lust. Goodman, Maminirina, Bradman et al. (2009) | Goodman, Ratrimomanarivo et al. (2008) | Kemp et al. (2019) | Monadjem, Razafimanahaka et al. (2017f) | Ramasindrazana et al. (2011) | Schoeman et al. (2015) | Thomas (1906) https://zenodo.org/record/5735302/files/figure.png 34. Manavil Long-fingered Bat Miniopterus manavi French: Minioptére des Betsileo / German: Zentral-Madagaskar-Langflligelfledermaus / Spanish: Minidptero de Betsileo Other common names: Manavil Bent-winged Bat Taxonomy. Miniopterus manavi Thomas, 1906 , “Imasindrary, N.E. Betsileo,” Madagascar . Miniopterus manavi has been considered the common species of Miniopterus , with the widest distribution in Madagascar . Nevertheless, recent genetic studies have shown it to represent a species complex with at least six species, several of them not even phylogenetically related. Some recognized characteristics of the life history of M. manavi date back to before the group was split, which makes it difficult to assign them to specific lineages so they are omitted here. Monotypic. Distribution. Restricted to highlands of C Madagascar from Fandriana S to Ranomafana (Ifanadiana) and Vinanitelo. Descriptive notes. Head—body c. 51 mm , tail 39 mm , ear 10 mm , hindfoot 6 mm , forearm 37-6-39-2 mm; weight 6-4 g (single individual except for forearm length). Dorsal hairs of the Manavil Long-fingered Bat are of medium length (c. 5 mm ). General color is blackish, with hairs blackish along two-thirds their length and having dull brown tips. Underparts are similar but rather paler. Some specimens are dull reddish throughout, representing a rufous morph, but others have coal black backs and slightly lighter bellies, which is best considered a reddish brown red morph. Wing and tail membranes are dark brownish black. Uropatagium and plagiopatagium are attached to femur at same level above anklejoint. Fur extends thinly over nearly one-half of upper surface of uropatagium, being sparser on proximal ventral surface. Tragus ( 6 mm ) is relatively thin along distal two-thirds ofits length, medial margin has a flange, distal lateral portion is slightly enlarged and rounded, and distal medial tip terminates with an angular straight edge. Habitat. Continuous forests, irrigated lowland rice paddies, secondary vegetation, and hillside rice at elevations of ¢.900-1500 m. In Ranomafana National Park, according to bat ultrasound recorders, the Manavil Long-fingered Bat seems to occur more frequently in forest fragments. Food and Feeding. Diet mainly contains Lepidoptera , Hymenoptera, Coleoptera, and Hemiptera , with lesser amounts of Blattodea, Diptera, Ephemeroptera , and Neuroptera . Prey also included mosquitoes that serve as vectors of diseases such as Anopheles squamosus (Malaria, Rift Valley fever virus [RVFV]) and Wuchereria bancrofti and Coquillettidia sp. (RVFV, West Nile virus [WNV]). Other prey is possibly fruit pests (e.g. Pandemis sp.). Breeding. No information. Activity patterns. The Manavil Long-fingered Bat is nocturnal. It uses caves as daytime shelters. Echolocation calls have downward FM signals, with maximum frequencies of 89-110 kHz, minimum frequency of 53 kHz, peak frequencies of 55-5-58-2 kHz, durations of 2-1-3 milliseconds, and intervals of 54-1-84-7 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Goodman, Maminirina, Bradman et al. (2009), Goodman, Ratrimomanarivo et al. (2008), Kemp et al. (2019), Monadjem, Razafimanahaka et al. (2017f), Ramasindrazana et al. (2011), Schoeman et al. (2015), Thomas (1906). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Miniopteridae Miniopterus manavi Miniopterus manavi Thomas 1906 0 Ann. Mag. Nat. Hist. ser. 7, 17: 176 Manavi Long-fingered Bat None. Madagascar, E/NE of Betsileo, 20°17'S, 47°31'E [Fandriana region]. Madagascar, Comoro Isls. Not listed. Least Concern Formerly included in minor , but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi ; see Juste et al. (2007), but see also Peterson et al. (1995). Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Miniopterus manavi 23 Manavil Long-fingered Bat Manavil Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA MINIOPTERIDAE NA NA Miniopterus NA manavi O. Thomas 1906 0 Miniopterus_manavi Thomas, O. (1906). New African mammals of the genera Cercopithecus, Scotophilus, Miniopterus, Crocidura, Georychus, and Heliophobius. Annals and Magazine of Natural History, ser. 7, 17, 176. https://www.biodiversitylibrary.org/item/209983#page/202/mode/1up BM 1897.9.1.37 "Imasindrary, N.E. Betsileo," Madagascar. manavi O. Thomas, 1906 previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae Juste, J., Ferrández, A., Fa, J. E., Masefield, W., & Ibáñez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao Tomé, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Goodman, S. M., Maminirina, C. P., Weyeneth, N., Bradman, H. M., Christidis, L., Ruedi, M., & Appleton, B. (2009). The use of molecular and morphological characters to resolve the taxonomic identity of cryptic species: the case of Miniopterus manavi (Chiroptera, Miniopteridae). Zoologica Scripta, 38(4), 339-363.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561. Madagascar Africa Afrotropic LC 0 0 0 Miniopterus_manavi 0 sciname match Miniopterus_manavi 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 80000000 Miniopterus manavi ANIMALIA CHORDATA MAMMALIA CHIROPTERA MINIOPTERIDAE Miniopterus manavi Thomas, 1906 20000000 Miniopterus manavi Least Concern 2017 2016-08-31 00:00:00 UTC 3.1 English This species is listed as Least Concern in view of its widespread distribution (near suitable caves) across Madagascar. While it is locally threatened by hunting, this is not thought to be a major threat of this species across its range. This species occurs in wide variety of vegetation types, such as dry deciduous forest, humid rainforest, degraded forest, plantations and agricultural areas (Goodman 1999, Eger and Mitchell 2003, Goodman et al. 2005, Rakotoarivelo and Randrianandriananina 2007), but requires the presence of suitable roost sites. It is not thought to be forest dependent, but is unlikely to forage in completely open areas (Goodman et al . 2005). There are no major threats to the species. In Makira, this species is eaten by people (Golden 2005), but it is not known if this species is hunted in other parts of its range. It is probably not forest dependent, but is rarely netted in areas without vegetation stands. It is locally abundant in cavities or rocky overhangs that provide suitable roosting features. A colony of approximately 4,000 was found in Makira (Bayliss and Hayes 1999). Unknown This species is found on Madagascar and the Comoros islands (Peterson et al. 1995). In Madagascar, it is widely distributed (Eger and Mitchell 2003, Goodman et al . 2005) and has a wide elevational range, from 20 to 1,500 m asl. Terrestrial This species occurs in many of the forest protected areas in western and eastern Madagascar and as such does not require specific conservation measures. Additional information on its ecology would be helpful to better understand how it responds to deforestation and roost disturbance. There is some uncertainty about the taxonomy of M. manavi and that this taxon is actually a complex of new species (F. Ratromomanarivo pers. comm). Further studies on morphology, acoustics and genetics are therefore needed. Afrotropical FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Miniopteridae Miniopterus manavi Thomas 1906 0 Ann. Mag. Nat. Hist. ser. 7, 17: 176 Manavi Long-fingered Bat None. Madagascar, E/NE of Betsileo, 20°17'S, 47°31'E [Fandriana region]. Madagascar, Comoro Isls. Not listed. Least Concern Formerly included in minor , but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi ; see Juste et al. (2007), but see also Peterson et al. (1995). Miniopterus manavi 1005122 23 Manavil Long-fingered Bat Manavil Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA Miniopteridae NA NA Miniopterus NA manavi O. Thomas 1906 0 Miniopterus_manavi Thomas, O. (1906). New African mammals of the genera Cercopithecus, Scotophilus, Miniopterus, Crocidura, Georychus, and Heliophobius. Annals and Magazine of Natural History, ser. 7, 17, 176. https://www.biodiversitylibrary.org/item/209983#page/202/mode/1up BM 1897.9.1.37 "Imasindrary, N.E. Betsileo," Madagascar. manavi O. Thomas, 1906 previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae Juste, J., Ferrández, A., Fa, J. E., Masefield, W., & Ibáñez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao Tomé, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Goodman, S. M., Maminirina, C. P., Weyeneth, N., Bradman, H. M., Christidis, L., Ruedi, M., & Appleton, B. (2009). The use of molecular and morphological characters to resolve the taxonomic identity of cryptic species: the case of Miniopterus manavi (Chiroptera, Miniopteridae). Zoologica Scripta, 38(4), 339-363.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561. Madagascar Africa Afrotropic LC 0 0 0 Miniopterus_manavi 0 sciname match Miniopterus_manavi 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Miniopterus_manavi 1005122 23 Manavil Long-fingered Bat Manavil Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yangochiroptera NA NA Vespertilionoidea Miniopteridae NA NA Miniopterus NA manavi O. Thomas 0 Miniopterus manavi Thomas, O. 1906-02-01. New African mammals of the genera _Cercopithecus_, _Scotophilus_, _Miniopterus_, _Crocidura_, _Georychus_, and _Heliophobius_. Annals and Magazine of Natural History (7)17(98):173-179. https://www.biodiversitylibrary.org/page/51472219 BMNH:Mamm:1897.9.1.37 holotype https://data.nhm.ac.uk/object/05f34bb0-1130-4dae-aa4b-0b03ac22b941 "Imasindrary, N.E. Betsileo," Madagascar. -20.2833 47.51667 previously included M. griveaudi and populations now attributed to M. aelleni, M. brachytragos, and M. mahafaliensis; moved from Vespertilionidae to Miniopteridae Juste, J., Ferrández, A., Fa, J. E., Masefield, W., & Ibáñez, C. (2007). Taxonomy of little bent-winged bats (Miniopterus, Miniopteridae) from the African islands of Sao Tomé, Grand Comoro and Madagascar, based on mtDNA. Acta Chiropterologica, 9(1), 27-37.|Goodman, S. M., Maminirina, C. P., Weyeneth, N., Bradman, H. M., Christidis, L., Ruedi, M., & Appleton, B. (2009). The use of molecular and morphological characters to resolve the taxonomic identity of cryptic species: the case of Miniopterus manavi (Chiroptera, Miniopteridae). Zoologica Scripta, 38(4), 339-363.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561. Madagascar Africa Afrotropic LC 0 0 0 Miniopterus_manavi 0 sciname match Miniopterus_manavi 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Miniopteridae Miniopterus manavi Thomas 1906 0 Ann. Mag. Nat. Hist. ser. 7, 17: 176 Manavi Long-fingered Bat None. Madagascar, E/NE of Betsileo, 20°17'S, 47°31'E [Fandriana region]. Madagascar, Comoro Isls. Not Listed Least Concern Formerly included in minor, but see Peterson et al. (1995); also see Juste and Ibáñez (1992). Does not include griveaudi; see Juste et al. (2007), but see also Peterson et al. (1995). Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Miniopterus manavi; Miniopterus manavi; Miniopterus manavi; Miniopterus manavi; Miniopterus manavi; Miniopterus manavi; manavi; griveaudi; manavi; Minioptére des Betsileo; Zentral-Madagaskar-Langflligelfledermaus; Minidptero de Betsileo; Manavil Bent-winged Bat; Manavil Long-fingered Bat; Manavil Bent-winged Bat; Manavi Long-fingered Bat; Manavi Long-fingered Bat; M. manavi