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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L66	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus societatis	Pipistrellus societatis	Pipistrellus societatis	Pipistrellus societatis	Pipistrellus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis	Arielulus societatis		[MSW2] Subgenus Arielulus. Synonymized with P. circumdatus by Heller and Volleth (1984:66-68), but see Hill and Francis (1984:314-315).; [MSW3] Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).; [HMW] Pipistrellus societatis Hill, 1972 , Malaysia , “Base Camp, Gunong Benom, Pahang , Malaya [= Malaysia ], 3°51'N , 102°11'E , 800 ft. [= 244 m ].” Previously considered merely a lowland form of A. circumdatus , but this opinion was rejected on the basis of morphological characteristics. Monotypic..; [batnames2022] Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).; [IUCN] Heller and Volleth (1984) treated this as a junior synonym of P. circumdatus but Hill and Francis (1984) and subsequent authors recognize it as a distinct species. Originally described in the genus Pipistrellus , it was placed in the subgenus Arielulus by Hill and Harrison (1987), a name which Csorba and Lee (1999) subsequently elevated to generic rank.; [batnames2023] Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).; [batnames2025_1.7] Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).											circumdatus?			societatis	Heller and Volleth (1984) treated this as a junior synonym of P. circumdatus but Hill and Francis (1984) and subsequent authors recognize it as a distinct species. Originally described in the genus Pipistrellus , it was placed in the subgenus Arielulus by Hill and Harrison (1987), a name which Csorba and Lee (1999) subsequently elevated to generic rank.			societatis	societatis			societatis (J. Edwards Hill, 1972)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Malaya; ref. 4.61	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus societatis	Malaysia, Pahang, Gunong Benom.	Hill	1972	Bull. Br. Mus. (Nat. Hist.) Zool., 23:34.	Distribution: Known only from Malaya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Malaya	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill	1972	Bull. Brit. Mus. (Nat. Hist.) Zool., 23:34.	Subgenus Arielulus. Synonymized with P. circumdatus by Heller and Volleth (1984:66-68), but see Hill and Francis (1984:314-315).	W Malaysia.	Malaysia, Pahang, Gunong Benom.		HILL	1972	Size medium (forearm length, 37-40 mm). Inner upper incisor faintly bicuspid. Anterior upper premolar may be absent. Supraorbital ridges and median sulcus of rostrum poorly developed. Braincase greatly inflated, but with the basicranial pits poorly developed. Last upper molar reduced.	Distribution: Known only from Malaya.	No subspecies.		115	species	P. societatis	HILL	1972	Arielulus	subgenus	Pipistrellus societatis				Size medium (forearm length, 37-40 mm). Inner upper incisor faintly bicuspid. Anterior upper premolar may be absent. Supraorbital ridges and median sulcus of rostrum poorly developed. Braincase greatly inflated, but with the basicranial pits poorly developed. Last upper molar reduced.	No subspecies.		41. P. societatis HILL 1972.	41	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Eptesicini	Arielulus societatis	Arielulus		societatis	Hill	y	1972		Bull. Brit. Mus. (Nat. Hist.) Zool.	23		34		Social Sprite	Malaysia, Pahang, Gunong Benom, Base Camp (03°51'N, 102°11'E), 800 ft. (266 m).	W Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).	4C3D87E8FFBE6A01FA8B933A1835B7FC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	827	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFBE6A01FA8B933A1835B7FC.xml	Arielulus societatis	Vespertilionidae	Arielulus	societatis		1972	Farfadet des sociétés @fr | Soziale Elfenfledermaus @de | Arielulus social @es | Benom-gilded Pipistrelle @en | Social Pipistrelle @en	Pipistrellus societatis Hill, 1972 , Malaysia , “Base Camp, Gunong Benom, Pahang , Malaya [= Malaysia ], 3°51'N , 102°11'E , 800 ft. [= 244 m ].” Previously considered merely a lowland form of A. circumdatus , but this opinion was rejected on the basis of morphological characteristics. Monotypic..	Known only from Peninsular Malaysia , including Selangor (Ulu Gombak) and Pahang (Sungai Tekam Forest Reserve, Gunung Benom, and Krau Wildlife Reserve) provinces. Also listed from Fraser’s Hill Forest Reserve, Pahang ; but this record needs verification.	Head-body 41-44 mm, tail 33-37 mm, ear 8-5-9 mm, forearm 36-6—40-6 mm; weight 4-3-8-5 g. The Social Sprite closely resembles the Bronze Sprite ( A. circumdatus ), but with smaller forearms, post-palatal extension, tooth row, and rostrum. Dorsal pelage is blackish brown with orange or bronze tipping (hairs basally brown, blackish brown for most of length, and tipped orange or bronze); crown pelage is orange-tipped but with a small area of straw-color; underparts are dark brown, tipped grayish white. Ears are large and bluntly rounded, with posteriorly directed basal lobe; tragus is large, and ears have yellowish-white rims, as in congeners. Wings are uniformly black. Baculum is Y-shaped with paired basal lobes and short shaft. Skull is short and wide, with rounded, rather globular braincase, inflated and elevated in frontal region; interorbital region wide; supraorbital region crests only faintly defined, terminating in small tubercles; rostrum is shorter, and narrower than in the Bronze Sprite, with more inflated braincase. Dentition is similar to that of the Bronze Sprite, less massive, with M® reduced, and posterior cusp virtually absent. Condylo-basal lengths are 13-9-14-8 mm; maxillary tooth row lengths are 5-2-5-6 mm. Chromosomal complement has 2n= 50 and FN= 48.	Recorded in primary lowland and hill forests, and secondary forests. The type specimen was found near a stream at 266 m .	No information.	No information.	Nocturnal. It was mist-netted above small ford on a tributary of the Gombak River . In Selangor , found roosting in a tree-trunk hole beside a forest stream.	No information.	Classified as Vulnerable on The IUCN Red List because its extent of occurrence is less than 20,000 km ?, its range is fragmented across forest patches, and there is continuing decline in the extent and quality of its habitat due to rapid deforestation. It occurs in Krau Wildlife Reserve.	Bates, Francis & Csorba (2008) | Boitani et al. (2006) | Corbet & Hill (1992) | Francis (2008a) | Heller & Volleth (1984, 1989) | Hill (1972, 1983) | Hill & Francis (1984) | Hill & Harrison (1987) | Kingston , Francis et al. (2003) | Kingston , Lim & Zubaid (2006) | Lim et al. (2017) | Medway (1983) | Mohd-Hanif et a/.(2015) | Simmons (2005)	https://zenodo.org/record/6398071/files/figure.png	142. Social Sprite Arielulus societatis French: Farfadet des sociétés / German: Soziale Elfenfledermaus / Spanish: Arielulus social Other common names: Benom-gilded Pipistrelle , Social Pipistrelle Taxonomy. Pipistrellus societatis Hill, 1972 , Malaysia , “Base Camp, Gunong Benom, Pahang , Malaya [= Malaysia ], 3°51'N , 102°11'E , 800 ft. [= 244 m ].” Previously considered merely a lowland form of A. circumdatus , but this opinion was rejected on the basis of morphological characteristics. Monotypic. Distribution Known only from Peninsular Malaysia , including Selangor (Ulu Gombak) and Pahang (Sungai Tekam Forest Reserve, Gunung Benom, and Krau Wildlife Reserve) provinces. Also listed from Fraser’s Hill Forest Reserve, Pahang ; but this record needs verification. Descriptive notes. Head-body 41-44 mm, tail 33-37 mm, ear 8-5-9 mm, forearm 36-6—40-6 mm; weight 4-3-8-5 g. The Social Sprite closely resembles the Bronze Sprite ( A. circumdatus ), but with smaller forearms, post-palatal extension, tooth row, and rostrum. Dorsal pelage is blackish brown with orange or bronze tipping (hairs basally brown, blackish brown for most of length, and tipped orange or bronze); crown pelage is orange-tipped but with a small area of straw-color; underparts are dark brown, tipped grayish white. Ears are large and bluntly rounded, with posteriorly directed basal lobe; tragus is large, and ears have yellowish-white rims, as in congeners. Wings are uniformly black. Baculum is Y-shaped with paired basal lobes and short shaft. Skull is short and wide, with rounded, rather globular braincase, inflated and elevated in frontal region; interorbital region wide; supraorbital region crests only faintly defined, terminating in small tubercles; rostrum is shorter, and narrower than in the Bronze Sprite, with more inflated braincase. Dentition is similar to that of the Bronze Sprite, less massive, with M® reduced, and posterior cusp virtually absent. Condylo-basal lengths are 13-9-14-8 mm; maxillary tooth row lengths are 5-2-5-6 mm. Chromosomal complement has 2n= 50 and FN= 48. Habitat. Recorded in primary lowland and hill forests, and secondary forests. The type specimen was found near a stream at 266 m . Food and Feeding. No information. Breeding. No information. Activity patterns. Nocturnal. It was mist-netted above small ford on a tributary of the Gombak River . In Selangor , found roosting in a tree-trunk hole beside a forest stream. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCN Red List because its extent of occurrence is less than 20,000 km ?, its range is fragmented across forest patches, and there is continuing decline in the extent and quality of its habitat due to rapid deforestation. It occurs in Krau Wildlife Reserve. Bibliography. Bates, Francis & Csorba (2008), Boitani et al. (2006), Corbet & Hill (1992), Francis (2008a), Heller & Volleth (1984, 1989), Hill (1972, 1983), Hill & Francis (1984), Hill & Harrison (1987), Kingston , Francis et al. (2003), Kingston , Lim & Zubaid (2006), Lim et al. (2017), Medway (1983), Mohd-Hanif et a/.(2015), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Arielulus societatis	Arielulus		societatis	Hill	1972	1	Bull. Brit. Mus. Nat. Hist. Zool.	23:34	Social Sprite	None.	Malaysia, Pahang, Gunong Benom, Base Camp (03°51'N, 102°11'E), 800 ft. (266 m).	W Malaysia.	Not listed.	Least Concern	Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Arielulus societatis	23	Social Sprite	Benom-gilded Pipistrelle|Social Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Arielulus	NA	societatis	J. Edwards Hill	1972	1	Pipistrellus_societatis	Hill, J. E. (1972). The Gunong Benom Expedition 1967. 4. New records of Malayan bats, with taxonomic notes and the description of a new Pipistrellus. Bulletin of the British Museum (Natural History). Zoology, 23, 34.	https://www.biodiversitylibrary.org/item/85291#page/82/mode/1up	BM 1967.1605		Malaysia, "Base Camp, Gunong Benom, Pahang, Malaya [= Malaysia], 3°51'N, 102°11'E, 800 ft. [= 244 m]."	3.85	102.18	societatis (J. Edwards Hill, 1972)	NA	NA	Malaysia	Asia	Indomalaya	LC	0	0	0	Arielulus_societatis	0	sciname match	Arielulus_societatis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	40776	Arielulus societatis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Arielulus	societatis	(Hill, 1972)	Heller and Volleth (1984) treated this as a junior synonym of P. circumdatus but Hill and Francis (1984) and subsequent authors recognize it as a distinct species. Originally described in the genus Pipistrellus , it was placed in the subgenus Arielulus by Hill and Harrison (1987), a name which Csorba and Lee (1999) subsequently elevated to generic rank.	20000000	Arielulus societatis	Least Concern		2020	2020-09-30 00:00:00 UTC	3.1	English	Listed as Least Concern, based on insufficient data to indicate that it qualifies in a higher threat category.	This species has been found in both primary and disturbed lowland and low hill forest at a number of localities, ranging from sea level to 400m above sea level, although it may occur at higher altitudes. It has also been caught in peat swamp forest. Most records are from near streams, with individuals generally being caught in mist nets set across small streams or rivers. However, it is likely that the species forages more widely in or above forest canopy, where it would be difficult to capture. Kingston et al. (2006) report that an individual was found roosting in a tree hole near a stream in Selangor, though the precise locality of that record was not provided.	Logging, agriculture, tree plantations, especially oil palm and rubber and forest fires leading to loss of lowland forest are likely the main threats to this species.	No estimates are available of the population size. The species has rarely been caught, but this may reflect mainly low capture probabilities, except at times when it comes low to the ground over rivers and streams.	Decreasing	The species is known from Peninsular Malaysia and peninsular Thailand. ; Reported distributional records include the following localities in Malaysia: (type locality) lowland rainforest at Gunung Benom base camp (3° 51' N, 102° 11' E, 250m asl; Hill 1972); regenerating hilly lowland forest near Fraser’s Hill, Selangor (3° 37' N, 101° 44' E, altitude 400m; Mohd-Hanif et al. 2015); disturbed hilly lowland forest at Ulu Gombak field station, Selangor (3° 19' N, 101° 45' E, 250m asl; Heller and Volleth 1984); lowland peatswamp forest at Sungai Bebar, Pekan forest reserve, Pahang (3° 15' N, 103° 20' E, 40m asl; Shukor et al. 2005). ; It has also been found at two localities in Thailand: lowland forest at Sirindhorn waterfall, Hala Bala Wildlife Sanctuary, Narathiwat (5° 48' N, 101° 49' E, 160m asl; Karapan et al. 2016); lowland forest in Khao Nam Khang National Park, Songkhla (6° 36' N, 100° 35' E, 120m asl) (Pipat Soisook and Charles M Francis, unpublished data). Likely it occurs in suitable patches of lowland forest throughout the Malay peninsula.		Terrestrial	This species has been reported from a number of protected areas including Krau Wildlife Reserve, Khao Nam Khang Wildlife Sanctuary and Hala Bala Wildlife Sanctuary. Further research is needed to determine the true range, population status and ecology of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Arielulus		societatis	Hill	1972	1	Bull. Brit. Mus. Nat. Hist. Zool.	23:34	Social Sprite	None.	Malaysia, Pahang, Gunong Benom, Base Camp (03°51'N, 102°11'E), 800 ft. (266 m).	W Malaysia.	Not listed.	Least Concern	Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).	Arielulus societatis	1005510	23	Social Sprite	Benom-gilded Pipistrelle|Social Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Arielulus	NA	societatis	J. Edwards Hill	1972	1	Pipistrellus_societatis	Hill, J. E. (1972). The Gunong Benom Expedition 1967. 4. New records of Malayan bats, with taxonomic notes and the description of a new Pipistrellus. Bulletin of the British Museum (Natural History). Zoology, 23, 34.	https://www.biodiversitylibrary.org/item/85291#page/82/mode/1up	BM 1967.1605		Malaysia, "Base Camp, Gunong Benom, Pahang, Malaya [= Malaysia], 3°51'N, 102°11'E, 800 ft. [= 244 m]."	3.85	102.1833	societatis (J. Edwards Hill, 1972)	NA	NA				Malaysia	Asia	Indomalaya	LC	0	0	0	Arielulus_societatis	0	sciname match	Arielulus_societatis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Arielulus_societatis	1005510	23	Social Sprite	Benom-gilded Pipistrelle|Social Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Arielulus	NA	societatis	J. Edwards Hill	1	Pipistrellus societatis	Hill, J.E. 1972. New records of Malayan bats, with taxonomic notes and the description of a new _Pipistrellus_. Bulletin of the British Museum (Natural History) (Zoology)23(3):21-42.	https://www.biodiversitylibrary.org/page/26496860	BMNH:Mamm:1967.1605	holotype	https://data.nhm.ac.uk/object/db461f0e-1ff4-4404-b36b-2e44c9b0be4a	Malaysia, "Base Camp, Gunong Benom, Pahang, Malaya [= Malaysia], 3°51'N, 102°11'E, 800 ft. [= 244 m]."	3.85	102.1833	NA	NA				Malaysia	Asia	Indomalaya	LC	0	0	0	Arielulus_societatis	0	sciname match	Arielulus_societatis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Arielulus		societatis	Hill	1972	1	Bull. Brit. Mus. Nat. Hist. Zool.	23:34	Social Sprite	None.	Malaysia, Pahang, Gunong Benom, Base Camp (03°51'N, 102°11'E), 800 ft. (266 m).	W Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/40776/22134204/' target='_blank'>Least Concern</a>	Synonymized with circumdatus by Heller and Volleth (1984), but see Hill and Francis (1984) and Corbet and Hill (1992).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Arielulus societatis; Arielulus societatis; Arielulus societatis; Arielulus societatis; Arielulus societatis; circumdatus; societatis; Farfadet des sociétés; Soziale Elfenfledermaus; Arielulus social; Benom-gilded Pipistrelle; Social Pipistrelle; Social Sprite; Benom-gilded Pipistrelle; Social Pipistrelle; Social Sprite; Social Sprite; Arielulus societatis circumdatus; Arielulus circumdatus; A. societatis