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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L649	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis	Miniopterus maghrebensis		[HMW] Miniopterus maghrebensis Puechmaille et al, 2014 , “Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °0I’ 46.6" N, 03° 47° 16.7" W, 1060 ma.s. 1.).” Miniopterus maghrebensis was originally referred to as M. schreibersii until a recent molecular study confirmed existence of genetic differences at mitochondrial and nuclearlevels to allow their separation. All previous records of M. schreibersii from the Maghreb need to be reevaluated because they could potentially belong to M. maghrebensis ; both species are found together in the region and can even share roosts. Monotypic.; [batnames2022] None.; [MDD2022] recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae; [IUCN] Previously this species was included within Miniopterus schreibersii ; it is now recognised as a separate species (Puechmaille et al . 2014).; [batnames2023] None.; [MDD2023] recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] None.; [MDD2025_2.2] recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae														maghrebensis	Previously this species was included within Miniopterus schreibersii ; it is now recognised as a separate species (Puechmaille et al . 2014).			maghrebensis	maghrebensis			maghrebensis Puechmaille, Allegrini, Benda, Bilgin, Ibáñez, & Juste in Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibáñez, Juste, & Bilgin, 2014						N/A																																								NA																											E84887F9FFDFD6520FEEF7791787359A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	700	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFDFD6520FEEF7791787359A.xml	Miniopterus maghrebensis	Miniopteridae	Miniopterus	maghrebensis	Puechmaille	2014	Minioptére du Maghreb @fr | Maghreb-Langfllgelfledermaus @de | Miniéptero de Maghreb @es | Maghrebian Bent-winged Bat @en	Miniopterus maghrebensis Puechmaille et al, 2014 , “Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °0I’ 46.6" N, 03° 47° 16.7" W, 1060 ma.s. 1.).” Miniopterus maghrebensis was originally referred to as M. schreibersii until a recent molecular study confirmed existence of genetic differences at mitochondrial and nuclearlevels to allow their separation. All previous records of M. schreibersii from the Maghreb need to be reevaluated because they could potentially belong to M. maghrebensis ; both species are found together in the region and can even share roosts. Monotypic.	So far known only from C & SW Morocco (Middle and High Atlas) and N Tunisia . Possibly occurs continuously across most of the Mediterranean part of N Africa, including Algeria .	Head-body 60 mm ,tail 63 mm , ear 13-4 mm, forearm 45-48 mm (holotype). The Maghrebian Long-fingered Bat is similar in most aspects to Schreibers’s Long-fingered Bat (M. schreibersit), from which it can only be taxonomically confirmed with genetic analyses. Dorsal pelage of the Maghrebian Long-fingered Bat is chestnut-brown to dark grayish brown, and ventral pelage is pale brown. Ventral hairs are bicolored, dark brown on proximal parts and pale brown to buffy on distal parts. Dorsal skin of ears and naked parts of face are pale grayish brown, and ventral skin of ears is almost without pigments, fleshy pale to pinkish. Wing membranes are dark brown to dark grayish brown. Ears are small, and tragus (6-3 mm) is slender and slightly curved forward.	Mediterranean scrublands and dry forests, steppes, and other semi-desert habitats at elevations of 300-1200 m. In Morocco , the Maghrebian Long-fingered Bat seems to prefer southern and more arid environments, and in northern one-half of Morocco , Schreibers’s Long-fingered Bat is found, although with large overlap between species.	There is no specific information available for this species, but based on wing morphology, the Maghrebian Long-fingered Bat probably forages for flying insects by moderately fast hawking in open, uncluttered places including clearings, above trees, and over water.	It is difficult to assign available information on reproduction of Miniopterus in the Maghreb to either the Maghrebian Long-fingered Bat or Schreibers’s L.ongfingered Bat. In any case, the pattern seems to be the typical for a temperate zone species of Miniopterus . An unusually wide range of dates of births has been reported for these bats in Algeria from mid-April to the end ofJune, but again it is not possible to taxonomically assign these records, and it was suggested that this wide range was due to the fact that the two species were present but had differentiated breeding periods.	The Maghrebian Long-fingered Bat is presumably nocturnal. As other temperate species of Miniopterus , it is expected to enter torpor in autumn, and it has been found to store fat in autumn for winter hibernation. It roosts in caves and other underground areas (e.g. irrigation channels). Echolocation calls have downward FM signals, with initial frequencies of 63-6-101-1 kHz, end frequencies of 48-9-51-8 kHz, peak frequencies of 51-54-8 kHz, durations of 3-2-5-6 milliseconds, and intervals of 67-2-239-2 milliseconds.	Colonies of Maghrebian Long-fingered Bats can include up to 400 individuals.	Classified as Near Threatened on The IUCN Red List because actual levels of disturbance are likely to lead to habitat declines of at least 30% in the next 15 years. This species is known from very few sites only, all representing underground spaces threatened by human activities.	Benda & Piraccini (2017) | Benda, Cerveny et al. (2010) | Benda, Ruedi & Aulagnier (2004) | Benda, Spitzenberger et al. (2014) | Bilgin et al. (2016) | Kowalski & Rzebik-Kowalska (1991) | Puechmaille, Allegrini et al. (2014) | Sramek et al. (2013)	https://zenodo.org/record/5735252/files/figure.png	17. Neghrohion Long-fingered Bat Miniopterus maghrebensis French: Minioptére du Maghreb / German: Maghreb-Langfllgelfledermaus / Spanish: Miniéptero de Maghreb Other common names: Maghrebian Bent-winged Bat Taxonomy. Miniopterus maghrebensis Puechmaille et al, 2014 , “Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °0I’ 46.6" N, 03° 47° 16.7" W, 1060 ma.s. 1.).” Miniopterus maghrebensis was originally referred to as M. schreibersii until a recent molecular study confirmed existence of genetic differences at mitochondrial and nuclearlevels to allow their separation. All previous records of M. schreibersii from the Maghreb need to be reevaluated because they could potentially belong to M. maghrebensis ; both species are found together in the region and can even share roosts. Monotypic. Distribution. So far known only from C & SW Morocco (Middle and High Atlas) and N Tunisia . Possibly occurs continuously across most of the Mediterranean part of N Africa, including Algeria . Descriptive notes. Head-body 60 mm ,tail 63 mm , ear 13-4 mm, forearm 45-48 mm (holotype). The Maghrebian Long-fingered Bat is similar in most aspects to Schreibers’s Long-fingered Bat (M. schreibersit), from which it can only be taxonomically confirmed with genetic analyses. Dorsal pelage of the Maghrebian Long-fingered Bat is chestnut-brown to dark grayish brown, and ventral pelage is pale brown. Ventral hairs are bicolored, dark brown on proximal parts and pale brown to buffy on distal parts. Dorsal skin of ears and naked parts of face are pale grayish brown, and ventral skin of ears is almost without pigments, fleshy pale to pinkish. Wing membranes are dark brown to dark grayish brown. Ears are small, and tragus (6-3 mm) is slender and slightly curved forward. Habitat. Mediterranean scrublands and dry forests, steppes, and other semi-desert habitats at elevations of 300-1200 m. In Morocco , the Maghrebian Long-fingered Bat seems to prefer southern and more arid environments, and in northern one-half of Morocco , Schreibers’s Long-fingered Bat is found, although with large overlap between species. Food and Feeding. There is no specific information available for this species, but based on wing morphology, the Maghrebian Long-fingered Bat probably forages for flying insects by moderately fast hawking in open, uncluttered places including clearings, above trees, and over water. Breeding. It is difficult to assign available information on reproduction of Miniopterus in the Maghreb to either the Maghrebian Long-fingered Bat or Schreibers’s L.ongfingered Bat. In any case, the pattern seems to be the typical for a temperate zone species of Miniopterus . An unusually wide range of dates of births has been reported for these bats in Algeria from mid-April to the end ofJune, but again it is not possible to taxonomically assign these records, and it was suggested that this wide range was due to the fact that the two species were present but had differentiated breeding periods. Activity patterns. The Maghrebian Long-fingered Bat is presumably nocturnal. As other temperate species of Miniopterus , it is expected to enter torpor in autumn, and it has been found to store fat in autumn for winter hibernation. It roosts in caves and other underground areas (e.g. irrigation channels). Echolocation calls have downward FM signals, with initial frequencies of 63-6-101-1 kHz, end frequencies of 48-9-51-8 kHz, peak frequencies of 51-54-8 kHz, durations of 3-2-5-6 milliseconds, and intervals of 67-2-239-2 milliseconds. Movements, Home range and Social organization. Colonies of Maghrebian Long-fingered Bats can include up to 400 individuals. Status and Conservation. Classified as Near Threatened on The IUCN Red List because actual levels of disturbance are likely to lead to habitat declines of at least 30% in the next 15 years. This species is known from very few sites only, all representing underground spaces threatened by human activities. Bibliography. Benda & Piraccini (2017), Benda, Cerveny et al. (2010), Benda, Ruedi & Aulagnier (2004), Benda, Spitzenberger et al. (2014), Bilgin et al. (2016), Kowalski & Rzebik-Kowalska (1991), Puechmaille, Allegrini et al. (2014), Sramek et al. (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Miniopteridae	Miniopterus maghrebensis	Miniopterus		maghrebensis	Puechmaille, Allegrini, Benda, Bilgin, Ib&aacute;&ntilde;ez, & Juste	2014	0	Zootaxa	3794(1): 112	Mahgrebian Bent-wing Bat	None.	Morocco, Er Rachidiyah Province, 19 km W-NW of Boudenib, 5.7 km S of Ksar Tazougart, Kef Azigza Cave (or Tazzouguert Cave)	N Morocco and into Tunisia	Not listed.	Near Threatened	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus maghrebensis	23	Maghrebian Long-fingered Bat	Maghrebian Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	maghrebensis	Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibañez, Juste, & Bilgin	2014	0	Miniopterus_maghrebensis	Puechmaille, S. J., Allegrini, B., Benda, P., Gürün, K., Å rà mek, J., Ibanez, C., Juste, J. & Bilgin, R. (2014). A new species of the Miniopterus schreibersii species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa, 3794(1), 112.	https://www.mapress.com/j/zt/article/view/zootaxa.3794.1.4	NMP 94426		"Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °01' 46.6" N, 03° 47' 16.7" W, 1060 m a. s. l.)."	32.03	-3.788	maghrebensis Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibañez, Juste, & Bilgin, 2014	recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae	Puechmaille, S. J., Allegrini, B., Benda, P., Gürün, K., Å rà mek, J., Ibanez, C., ... & Bilgin, R. (2014). A new species of the Miniopterus schreibersii species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa, 3794(1), 108-124.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.	Morocco|Algeria|Tunisia	Africa	Afrotropic	NT	0	0	0	Miniopterus_maghrebensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Miniopterus maghrebensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MINIOPTERIDAE	Miniopterus	maghrebensis	Puechmaille, Allegrini, Benda, Gurun, Sramek, Ibanez, Juste &; Bilgin, 2014	Previously this species was included within Miniopterus schreibersii ; it is now recognised as a separate species (Puechmaille et al . 2014).	100000000	Miniopterus maghrebensis	Near Threatened	A3c	2017	2016-04-27 00:00:00 UTC	3.1	English	This species is known from very few sites only, all representing underground spaces threatened by human activities. Currently it is listed as Near Threatened because actual levels of disturbance are likely to lead the habitat of this species to decline of at least 30% in the next 15 years.	<p>The Maghrebian Bent-wing Bat inhabits the Mediterranean arboreal (subtropical dry ;forest, Mediterranean-type shrubland) vegetation zone. It is known to roost in caves. The species stores fat in autumn for the winter months.</p>	Threats to the species are mainly human disturbance of roost sites, and perhaps the use of pesticides against insects.	This species has been found in the Mediterranean and steppe habitats of the Maghreb.	Unknown	This species possibly occurs across the Mediterranean part of North Africa, but it is known only from central and south-western Morocco and northern Tunisia.		Terrestrial	No specific measures are known to be in place for this species, but it occurs in protected areas across its range.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		maghrebensis	Puechmaille, Allegrini, Benda, Bilgin, Ib&aacute;&ntilde;ez, & Juste	2014	0	Zootaxa	3794(1): 112	Mahgrebian Bent-wing Bat	None.	Morocco, Er Rachidiyah Province, 19 km W-NW of Boudenib, 5.7 km S of Ksar Tazougart, Kef Azigza Cave (or Tazzouguert Cave)	N Morocco and into Tunisia	Not listed.	Near Threatened	None.	Miniopterus maghrebensis	1005118	23	Maghrebian Long-fingered Bat	Maghrebian Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	maghrebensis	Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibañez, Juste, & Bilgin	2014	0	Miniopterus_maghrebensis	Puechmaille, S. J., Allegrini, B., Benda, P., Gürün, K., Å rà mek, J., Ibanez, C., Juste, J. & Bilgin, R. (2014). A new species of the Miniopterus schreibersii species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa, 3794(1), 112.	https://www.mapress.com/j/zt/article/view/zootaxa.3794.1.4	NMP 94426		"Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °01' 46.6" N, 03° 47' 16.7" W, 1060 m a. s. l.)."	32.02961	-3.78797	maghrebensis Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibañez, Juste, & Bilgin, 2014	recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae	Puechmaille, S. J., Allegrini, B., Benda, P., Gürün, K., Å rà mek, J., Ibanez, C., ... & Bilgin, R. (2014). A new species of the Miniopterus schreibersii species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa, 3794(1), 108-124.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Morocco|Algeria|Tunisia	Africa	Afrotropic	NT	0	0	0	Miniopterus_maghrebensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_maghrebensis	1005118	23	Maghrebian Long-fingered Bat	Maghrebian Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	maghrebensis	Puechmaille, Allegrini, Benda, Bilgin, Ibáñez, & Juste in Puechmaille, Allegrini, Benda, Gürün, Å rámek, Ibáñez, Juste, & Bilgin	0	Miniopterus maghrebensis	Puechmaille, S.J., Allegrini, B., Benda, P., Gürün, K., Å rámek, J., Ibáñez, C., Juste, J. and Bilgin, R. 2014-05-05. A new species of the _Miniopterus schreibersii_ species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa 3794(1):108-124.	https://doi.org/10.11646/zootaxa.3794.1.4	NMPr 94426	holotype		"Kef Azigza Cave (or Tazzouguert Cave), 5.7 km S of Ksar Tazougart, 19 km W-NW of Boudenib, Er Rachidiyah Province, Morocco (32 °01' 46.6" N, 03° 47' 16.7" W, 1060 m a. s. l.)."	32.02961	-3.78797	recently described based on populations of M. schreibersii; moved from Vespertilionidae to Miniopteridae	Puechmaille, S. J., Allegrini, B., Benda, P., Gürün, K., Å rà mek, J., Ibanez, C., ... & Bilgin, R. (2014). A new species of the Miniopterus schreibersii species complex (Chiroptera: Miniopteridae) from the Maghreb Region, North Africa. Zootaxa, 3794(1), 108-124.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Morocco|Algeria|Tunisia	Africa	Afrotropic	NT	0	0	0	Miniopterus_maghrebensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		maghrebensis	Puechmaille, Allegrini, Benda, Bilgin, Ib&aacute;&ntilde;ez & Juste	2014	0	Zootaxa	3794(1): 112	Mahgrebian Bent-wing Bat	None.	Morocco, Er Rachidiyah Province, 19 km W-NW of Boudenib, 5.7 km S of Ksar Tazougart, Kef Azigza Cave (or Tazzouguert Cave)	N Morocco and into Tunisia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81633156/95642265/' target='_blank'>Near Threatened</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus maghrebensis; Miniopterus maghrebensis; Miniopterus maghrebensis; Miniopterus maghrebensis; Miniopterus maghrebensis; maghrebensis; Minioptére du Maghreb; Maghreb-Langfllgelfledermaus; Miniéptero de Maghreb; Maghrebian Bent-winged Bat; Maghrebian Long-fingered Bat; Maghrebian Bent-winged Bat; Mahgrebian Bent-wing Bat; M. maghrebensis