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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L645	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi	Miniopterus griffithsi		[HMW] Miniopterus griffithst Goodman et al., 2010 , “ Madagascar : Province de Toliara , Grotte d’Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda , 24°39.012°S, 45°57.797°E, at 110 m above sea-level.” Miniopterus griffithsi was formerly included in M. glen: and now considered as its sister species. Monotypic.; [batnames2022] None.; [MDD2022] recently described; moved from Vespertilionidae to Miniopteridae; [IUCN] Peterson et al. (1995) described Miniopterus gleni based on specimens collected in different portions of Madagascar. ; A phylogeographic study of this species found little genetic variation across its range, with the exception of a population in the extreme southwest and southeast, which was named M . griffithsi (Goodman et al. 2010).; [batnames2023] Although published online in 2009, this paper failed to provide a ZooBank identifier for griffithsi within the paper and therefore the name is unavailable until the print version appeared in February 2010, following section 8.5.3 of the Code.; [MDD2023] recently described; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] recently described; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] Although published online in 2009, this paper failed to provide a ZooBank identifier for griffithsi within the paper and therefore the name is unavailable until the print version appeared in February 2010, following section 8.5.3 of the Code.; [MDD2025_2.2] recently described; moved from Vespertilionidae to Miniopteridae														griffithsi	Peterson et al. (1995) described Miniopterus gleni based on specimens collected in different portions of Madagascar. ; A phylogeographic study of this species found little genetic variation across its range, with the exception of a population in the extreme southwest and southeast, which was named M . griffithsi (Goodman et al. 2010).			griffithsi	griffithsi			griffithsi S. M. Goodman, Maminirina, Bradman, Christidis, & B. R. Appleton, 2010						N/A																																								NA																											E84887F9FFC4D64A0FC8F960189C3B9A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	709	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFC4D64A0FC8F960189C3B9A.xml	Miniopterus griffithsi	Miniopteridae	Miniopterus	griffithsi	Goodman, Maminirina, Bradman, Christidis & Appleton	2010	Minioptére de Griffiths @fr | Griffiths-Landfligelfledermaus @de | Miniéptero de Griffiths @es | Griffiths’'s Bent-winged Bat @en	Miniopterus griffithst Goodman et al., 2010 , “ Madagascar : Province de Toliara , Grotte d’Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda , 24°39.012°S, 45°57.797°E, at 110 m above sea-level.” Miniopterus griffithsi was formerly included in M. glen: and now considered as its sister species. Monotypic.	S Madagascar , known only from three localities in the former Toliara Province S of the Onilahy River.	Head-body ¢.65-68 mm, tail 54-63 mm, ear 13-14 mm, hindfoot 8 mm , forearm 48-50 mm; weight 12-15-5 g. Griffiths’s Long-fingered Batis large, being among the largest species of Miniopterus in Madagascar together with Glen’s Long-fingered Bat (M. glen?) Anterior part of dorsum is distinctly lighter brown than middle and posterior parts that approach medium brown. Ventrum is a mix of predominantly lighter brown (particularly toward wing membrane), with interspersed medium brown fur. Wing membrane and uropatagium are dark brown, with notable light spotting on uropatagium. Tragusis relatively long, consistently 8 mm in type series, and thickset, with distinct flange on distal one-half of external surface and slightly tapered downward deflection on inner edge; distal end is not particularly rounded, giving a slightly squarish terminal edge.	Presumably open habitats or close to forest edges at elevations below 110 m . It is unclear if Griffiths’s Long-fingered Bat is at least partially forest dependent.	Griffiths’s Long-fingered Bat is expected to feed mainly on soft insects captured in flight as do other long-fingered bats.	A male Griffiths’s Long-fingered Bat captured in late February had scrotal testes. A single female captured in late February had no signs of reproduction.	Griffiths’s Long-fingered Bat is nocturnal. Two individuals, caught in May at start of the long dry season, had considerable subcutaneous fat and weighed 15-5 g and 16-5 g. Accumulation offat deposits is presumably an adaptation to survive long dry seasons when food resources are reduced and with probably associated daily torpors. All known individuals were captured in caves or their surroundings. Echolocation calls have downward FM signals, with maximum frequencies of 61-99 kHz, minimum frequency of 40 kHz, peak frequencies of 43-5—45-3 kHz, durations of 2-9-3-6 milliseconds, and intervals of 56-4—128-5 milliseconds.	No information.	Classified as Data Deficient on The IUCN Red List. All localities where Griffiths’s Long-fingered Bat was found have degraded habitats by human activities such forest logging for charcoal production or livestock (goats and sheep) grazing.	Goodman (2017f) | Goodman, Maminirina et al. (2010) | Ramasindrazana et al. (2011)	https://zenodo.org/record/5735314/files/figure.png	38. Griffiths’s Long-fingered Bat Miniopterus griffithsi French: Minioptére de Griffiths / German: Griffiths-Landfligelfledermaus / Spanish: Miniéptero de Griffiths Other common names: Griffiths’'s Bent-winged Bat Taxonomy. Miniopterus griffithst Goodman et al., 2010 , “ Madagascar : Province de Toliara , Grotte d’Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda , 24°39.012°S, 45°57.797°E, at 110 m above sea-level.” Miniopterus griffithsi was formerly included in M. glen: and now considered as its sister species. Monotypic. Distribution. S Madagascar , known only from three localities in the former Toliara Province S of the Onilahy River. Descriptive notes. Head-body ¢.65-68 mm, tail 54-63 mm, ear 13-14 mm, hindfoot 8 mm , forearm 48-50 mm; weight 12-15-5 g. Griffiths’s Long-fingered Batis large, being among the largest species of Miniopterus in Madagascar together with Glen’s Long-fingered Bat (M. glen?) Anterior part of dorsum is distinctly lighter brown than middle and posterior parts that approach medium brown. Ventrum is a mix of predominantly lighter brown (particularly toward wing membrane), with interspersed medium brown fur. Wing membrane and uropatagium are dark brown, with notable light spotting on uropatagium. Tragusis relatively long, consistently 8 mm in type series, and thickset, with distinct flange on distal one-half of external surface and slightly tapered downward deflection on inner edge; distal end is not particularly rounded, giving a slightly squarish terminal edge. Habitat. Presumably open habitats or close to forest edges at elevations below 110 m . It is unclear if Griffiths’s Long-fingered Bat is at least partially forest dependent. Food and Feeding. Griffiths’s Long-fingered Bat is expected to feed mainly on soft insects captured in flight as do other long-fingered bats. Breeding. A male Griffiths’s Long-fingered Bat captured in late February had scrotal testes. A single female captured in late February had no signs of reproduction. Activity patterns. Griffiths’s Long-fingered Bat is nocturnal. Two individuals, caught in May at start of the long dry season, had considerable subcutaneous fat and weighed 15-5 g and 16-5 g. Accumulation offat deposits is presumably an adaptation to survive long dry seasons when food resources are reduced and with probably associated daily torpors. All known individuals were captured in caves or their surroundings. Echolocation calls have downward FM signals, with maximum frequencies of 61-99 kHz, minimum frequency of 40 kHz, peak frequencies of 43-5—45-3 kHz, durations of 2-9-3-6 milliseconds, and intervals of 56-4—128-5 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. All localities where Griffiths’s Long-fingered Bat was found have degraded habitats by human activities such forest logging for charcoal production or livestock (goats and sheep) grazing. Bibliography. Goodman (2017f), Goodman, Maminirina et al. (2010), Ramasindrazana et al. (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Miniopteridae	Miniopterus griffithsi	Miniopterus		griffithsi	Goodman, Maminirina, Bradman, Christidis & Appleton	2010	0	J. Zool. Syst. Evol. Res.	48(1): 81	Griffiths&#39; Long-fingered Bat	None.	Madagascar, Province de Toliara, Grotte d&#39;Androimpano, NE Itampolo	S Madagascar	Not listed.	Data Deficient	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus griffithsi	23	Griffiths's Long-fingered Bat	Griffiths's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	griffithsi	Goodman, Raminirina, Bradman, Christidis, & Appleton	2010	0	Miniopterus_griffithsi	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. R. (2010). Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi. Journal of Zoological Systematics and Evolutionary Research, 48(1), 81.	https://onlinelibrary.wiley.com/doi/pdf/10.1111/j.1439-0469.2009.00524.x?casa_token=TYk3AhA2B0QAAAAA:jtsR3B18_S-l9uWF-5rUy7NmO95eKeFjZmvSDhvmOZAr7P3glK_KIFHpdEUK-KTq6luPU5eRo-uDGA	FMNH 184214		"Madagascar: Province de Toliara, Grotte d'Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda, 24°39.012'S, 43°57.797'E, at 110 m above sea-level."	-24.65	43.963	griffithsi Goodman, Raminirina, Bradman, Christidis, & Appleton, 2010	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. R. (2010). Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi. Journal of Zoological Systematics and Evolutionary Research, 48(1), 75-86.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.	Madagascar	Africa	Afrotropic	DD	0	0	0	Miniopterus_griffithsi	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Miniopterus griffithsi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MINIOPTERIDAE	Miniopterus	griffithsi	Goodman, Maminirina, Bradman, Christidis &; Appleton, 2010	Peterson et al. (1995) described Miniopterus gleni based on specimens collected in different portions of Madagascar. ; A phylogeographic study of this species found little genetic variation across its range, with the exception of a population in the extreme southwest and southeast, which was named M . griffithsi (Goodman et al. 2010).	100000000	Miniopterus griffithsi	Data Deficient		2017	2016-05-02 00:00:00 UTC	3.1	English	This newly described species is known from few localities in the southern portion of Madagascar (Goodman and Ramasindrazana 2013). Research is needed to better define the geographical distribution of Miniopterus griffithsi and new field data are necessary to assess properly this taxon.	At the two sites along the Mahafaly Plateau in the southwest where this species has been collected, it was captured while exiting cave systems and the site in the southeast where it was netted, presumably while foraging, also contains numerous cave systems. ;Hence, it is presumed to make its day roost sites in cave systems. ;It presumably forages in areas that are more open or close to the forest edge and it is unclear if it is at least partially forest dependant.	Threats to this species are not known.	Population size and trends are not known for this species.	Unknown	This bat is known from three different sites in southern Madagascar and across an elevational range from 15 to 110 m above sea level (Goodman and Ramasindrazana 2013).		Terrestrial	This species is not currently known from any protected area. Two of the three sites it has been documented are in degraded forested ecosystems (Goodman et al. 2010). If indeed it is not forest dependent, these human pressures may not directly impact this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		griffithsi	Goodman, Maminirina, Bradman, Christidis & Appleton	2010	0	J. Zool. Syst. Evol. Res.	48(1): 81	Griffiths&#39; Long-fingered Bat	None.	Madagascar, Province de Toliara, Grotte d&#39;Androimpano, NE Itampolo	S Madagascar	Not listed.	Data Deficient	Although published online in 2009, this paper failed to provide a ZooBank identifier for griffithsi within the paper and therefore the name is unavailable until the print version appeared in February 2010, following section 8.5.3 of the Code.	Miniopterus griffithsi	1005114	23	Griffiths's Long-fingered Bat	Griffiths's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	griffithsi	Goodman, Raminirina, Bradman, Christidis, & Appleton	2010	0	Miniopterus_griffithsi	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. R. (2010). Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi. Journal of Zoological Systematics and Evolutionary Research, 48(1), 81.	https://onlinelibrary.wiley.com/doi/pdf/10.1111/j.1439-0469.2009.00524.x?casa_token=TYk3AhA2B0QAAAAA:jtsR3B18_S-l9uWF-5rUy7NmO95eKeFjZmvSDhvmOZAr7P3glK_KIFHpdEUK-KTq6luPU5eRo-uDGA	FMNH 184214		"Madagascar: Province de Toliara, Grotte d'Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda, 24°39.012'S, 43°57.797'E, at 110 m above sea-level."	-24.6502	43.96328	griffithsi Goodman, Raminirina, Bradman, Christidis, & Appleton, 2010	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. R. (2010). Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi. Journal of Zoological Systematics and Evolutionary Research, 48(1), 75-86.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	DD	0	0	0	Miniopterus_griffithsi	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_griffithsi	1005114	23	Griffiths's Long-fingered Bat	Griffiths's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	griffithsi	S. M. Goodman, Maminirina, Bradman, Christidis, & B. R. Appleton	0	Miniopterus griffithsi	Goodman, S.M., Maminirina, C.P., Bradman, H.M., Christidis, L.L. and Appleton, B.R. 2010-02. Patterns of morphological and genetic variation in the endemic Malagasy bat _Miniopterus gleni_ (Chiroptera: Miniopteridae), with the description of a new species, _M. griffithsi_. Journal of Zoological Systematics and Evolutionary Research 48(1):75-86.	https://doi.org/10.1111/j.1439-0469.2009.00524.x	FMNH:Mamm:184214	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2682982	"Madagascar: Province de Toliara, Grotte d'Androimpano, 4.2 km NE Itampolo (village), on old road to Ejeda, 24°39.012'S, 43°57.797'E, at 110 m above sea-level."	-24.6502	43.96328	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. R. (2010). Patterns of morphological and genetic variation in the endemic Malagasy bat Miniopterus gleni (Chiroptera: Miniopteridae), with the description of a new species, M. griffithsi. Journal of Zoological Systematics and Evolutionary Research, 48(1), 75-86.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	DD	0	0	0	Miniopterus_griffithsi	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		griffithsi	Goodman, Maminirina, Bradman, Christidis & Appleton	2010	0	J. Zool. Syst. Evol. Res.	48(1): 81	Griffiths&#39; Long-fingered Bat	None.	Madagascar, Province de Toliara, Grotte d&#39;Androimpano, NE Itampolo	S Madagascar	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81633105/95642250/' target='_blank'>Data Deficient</a>	Although published online in 2009, this paper failed to provide a ZooBank identifier for griffithsi within the paper and therefore the name is unavailable until the print version appeared in February 2010, following section 8.5.3 of the Code.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus griffithsi; Miniopterus griffithsi; Miniopterus griffithsi; Miniopterus griffithsi; Miniopterus griffithsi; griffithsi; Minioptére de Griffiths; Griffiths-Landfligelfledermaus; Miniéptero de Griffiths; Griffiths’'s Bent-winged Bat; Griffiths's Long-fingered Bat; Griffiths's Bent-winged Bat; Griffiths&#39; Long-fingered Bat; M. griffithsi