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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L639	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Miniopterus schreibersii [synonym of]	Miniopterus schreibersii eschscholtzii	Miniopterus schreibersii eschscholtzii	Miniopterus eschscholtzii	Miniopterus fuliginosus eschscholtzii	Miniopterus eschscholtzii	N/A	Miniopterus eschscholtzii	Miniopterus eschscholtzii	Miniopterus eschscholtzii	Miniopterus eschscholtzii	Miniopterus eschscholtzii		[HMW] Vespertilio eschscholtzii Water-house, 1845 , “ Philippine Islands .” Miniopterus eschscholtzii was traditionally considered as M. schreibersii in the Philippines , generally at subspecific rank. Genetic studies of the genus Miniopterus in which specimens of the schreibersii species complex of the Philippines have been included indicate that they are different from Asian (fuliginosus, magnater , and blepotis ), and Australian ( orianae ) representatives of this complex. Apparently, M. eschscholtzii is closer to M. fuliginosus. There are also some cranial differences in size that supportits recognition as a distinct species. In any case, their taxonomic situation is pending a thorough revision of Asian forms of Miniopterus . Monotypic.; [MDD2022] split from M. schreibersii; moved from Vespertilionidae to Miniopteridae; [batnames2023] Formerly included in schreibersii or fuliginosus , but likely distinct; see Kusuminda et al. (2022), which strongly suggests that - once the holotype is examined - eschscholtzii will be recognized as a distinct species.; [MDD2023] split from M. schreibersii; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] split from M. schreibersii; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] Formerly included in schreibersii or fuliginosus, but likely distinct; see Kusuminda et al. (2022), which strongly suggests that - once the holotype is examined - eschscholtzii will be recognized as a distinct species.; [MDD2025_2.2] split from M. schreibersii; moved from Vespertilionidae to Miniopteridae														eschscholtzii				eschscholtzii 	eschscholtzii 			eschscholtzii (G. R. Waterhouse, 1845)|eschholtzii (Fitzinger, 1870) [incorrect subsequent spelling]						N/A																																								NA																											E84887F9FFD2D65C0ADAFAC610813B9C	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	695	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFD2D65C0ADAFAC610813B9C.xml	Miniopterus eschscholtzii	Miniopteridae	Miniopterus	eschscholtzii		1845	Minioptéere d'Eschscholtz @fr | Eschscholtz-Langflligelfledermaus @de | Minidptero de Eschscholtz @es | Eschscholtz’'s Bent-winged Bat @en	Vespertilio eschscholtzii Water-house, 1845 , “ Philippine Islands .” Miniopterus eschscholtzii was traditionally considered as M. schreibersii in the Philippines , generally at subspecific rank. Genetic studies of the genus Miniopterus in which specimens of the schreibersii species complex of the Philippines have been included indicate that they are different from Asian (fuliginosus, magnater , and blepotis ), and Australian ( orianae ) representatives of this complex. Apparently, M. eschscholtzii is closer to M. fuliginosus. There are also some cranial differences in size that supportits recognition as a distinct species. In any case, their taxonomic situation is pending a thorough revision of Asian forms of Miniopterus . Monotypic.	Widely distributed across the Philippines .	Head-body c¢. 55 mm , tail 46-57 mm, ear 11-13 mm, hindfoot 8-10 mm, forearm 42-46 mm; weight 8-14 g. Pelage of Eschscholtz’s Long-fingered Bat is dense and very dark brown and longer over shoulders and on top of head than elsewhere on body. Ears are small. Tragus is narrow, curved, rounded at its tip, and indistinctly emarginated on outer margin. Wing membranes are dark.	Wide variety of habitats from primary lowland forests and montane forests to disturbed forests and agricultural areas from near sea level up to elevations of c. 1500 m .	There is no specific information available for this species, but other long-fingered bats mainly eat soft insects ( Lepidoptera and Diptera ) captured in flight.	Some observations of Eschscholtz’s Long-fingered Bats from different locations suggest that only one young is born per pregnancy. They are not reproductively active in December (only one pregnant female out of 214 captured), but females with small embryos were captured in March, and ¢.50% was pregnant and none were lactating in mid-May. No female was pregnant at the beginning ofJuly, and only four out of 421 were lactating.	Eschscholtz’s Long-fingered Bat is assumed to be mainly nocturnal. It roosts In caves but occasionally uses man-made tunnels. It is a common species in areas with appropriate roosts. Echolocation calls have downward FM signals, and mean minimum frequency is 45-6 kHz.	Colonies of hundreds Eschscholtz’s Long-fingered Bats have been reported sharing roosts with other species of bats such as: the Lesser Dawn Bat (Eonycteris spelaea), the Dusky Leaf-nosed Bat (Hipposideros ater), the Diadem Leaf-nosed Bat (H. diadema), the Acuminate Horseshoe Bat (Rhinolophus acuminatus), the Arcuate Horseshoe Bat (R. arcuatus), Creagh’s Horseshoe Bat (R. creaghi), the Philippine Long-fingered Bat ( M. paululus ), the Great Long-fingered Bat (M. tnistis), and the Pallid Large-footed Myotis ( Myotis macrotarsus).	Not assessed on The IUCN Red List. Eschscholtz’s Long-fingered Bat is now a separate species from Schreibers’s Long-fingered Bat (M. schreibers11), which is classified as Near Threatened.	Akmali et al. (2015) | Corbet & Hill (1992) | Esselstyn, Widmann & Heaney (2004) | Furman, Oztunc & Coraman (2010) | Heaney, Balete et al. (1998) | Heaney, Dolar et al. (2010) | Heaney, Gonzales et al. (1991) | Heaney, Tabaranza et al. (2006) | Hill (1983) | Maeda (1982) | Rickart et al. (1993) | Sedlock (2001) | Sramek et al. (2013) | Waterhouse (1845)	https://zenodo.org/record/5735218/files/figure.png	6. Eschscholtz’s Long-fingered Bat Miniopterus eschscholtzii French: Minioptéere d'Eschscholtz / German: Eschscholtz-Langflligelfledermaus / Spanish: Minidptero de Eschscholtz Other common names: Eschscholtz’'s Bent-winged Bat Taxonomy. Vespertilio eschscholtzii Water-house, 1845 , “ Philippine Islands .” Miniopterus eschscholtzii was traditionally considered as M. schreibersii in the Philippines , generally at subspecific rank. Genetic studies of the genus Miniopterus in which specimens of the schreibersii species complex of the Philippines have been included indicate that they are different from Asian (fuliginosus, magnater , and blepotis ), and Australian ( orianae ) representatives of this complex. Apparently, M. eschscholtzii is closer to M. fuliginosus. There are also some cranial differences in size that supportits recognition as a distinct species. In any case, their taxonomic situation is pending a thorough revision of Asian forms of Miniopterus . Monotypic. Distribution. Widely distributed across the Philippines . Descriptive notes. Head-body c¢. 55 mm , tail 46-57 mm, ear 11-13 mm, hindfoot 8-10 mm, forearm 42-46 mm; weight 8-14 g. Pelage of Eschscholtz’s Long-fingered Bat is dense and very dark brown and longer over shoulders and on top of head than elsewhere on body. Ears are small. Tragus is narrow, curved, rounded at its tip, and indistinctly emarginated on outer margin. Wing membranes are dark. Habitat. Wide variety of habitats from primary lowland forests and montane forests to disturbed forests and agricultural areas from near sea level up to elevations of c. 1500 m . Food and Feeding. There is no specific information available for this species, but other long-fingered bats mainly eat soft insects ( Lepidoptera and Diptera ) captured in flight. Breeding. Some observations of Eschscholtz’s Long-fingered Bats from different locations suggest that only one young is born per pregnancy. They are not reproductively active in December (only one pregnant female out of 214 captured), but females with small embryos were captured in March, and ¢.50% was pregnant and none were lactating in mid-May. No female was pregnant at the beginning ofJuly, and only four out of 421 were lactating. Activity patterns. Eschscholtz’s Long-fingered Bat is assumed to be mainly nocturnal. It roosts In caves but occasionally uses man-made tunnels. It is a common species in areas with appropriate roosts. Echolocation calls have downward FM signals, and mean minimum frequency is 45-6 kHz. Movements, Home range and Social organization. Colonies of hundreds Eschscholtz’s Long-fingered Bats have been reported sharing roosts with other species of bats such as: the Lesser Dawn Bat (Eonycteris spelaea), the Dusky Leaf-nosed Bat (Hipposideros ater), the Diadem Leaf-nosed Bat (H. diadema), the Acuminate Horseshoe Bat (Rhinolophus acuminatus), the Arcuate Horseshoe Bat (R. arcuatus), Creagh’s Horseshoe Bat (R. creaghi), the Philippine Long-fingered Bat ( M. paululus ), the Great Long-fingered Bat (M. tnistis), and the Pallid Large-footed Myotis ( Myotis macrotarsus). Status and Conservation. Not assessed on The IUCN Red List. Eschscholtz’s Long-fingered Bat is now a separate species from Schreibers’s Long-fingered Bat (M. schreibers11), which is classified as Near Threatened. Bibliography. Akmali et al. (2015), Corbet & Hill (1992), Esselstyn, Widmann & Heaney (2004), Furman, Oztunc & Coraman (2010), Heaney, Balete et al. (1998), Heaney, Dolar et al. (2010), Heaney, Gonzales et al. (1991), Heaney, Tabaranza et al. (2006), Hill (1983), Maeda (1982), Rickart et al. (1993), Sedlock (2001), Sramek et al. (2013), Waterhouse (1845).		Miniopteridae	Miniopterus fuliginosus eschscholtzii	Miniopterus		fuliginosus													Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus eschscholtzii	23	Eschscholtz's Long-fingered Bat	Eschscholtz's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	eschscholtzii	Waterhouse	1845	1						"Philippine Islands."			eschscholtzii (Waterhouse, 1845)	split from M. schreibersii; moved from Vespertilionidae to Miniopteridae	Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.	Philippines	Asia	Indomalaya	NA	0	0	0	Miniopterus_eschscholtzii	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		eschscholtzii	Waterhouse	1845	1	Proc. Zool. Soc. London	13 (1845): 3	Eschscholtz's Bent-winged Bat	None.	Philippine Islands	Philippines	Not listed.	Vulnerable under Miniopterus schreibersii 	Formerly included in schreibersii or fuliginosus , but likely distinct; see Kusuminda et al. (2022), which strongly suggests that - once the holotype is examined - eschscholtzii will be recognized as a distinct species.	Miniopterus eschscholtzii	1005109	23	Eschscholtz's Long-fingered Bat	Eschscholtz's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	eschscholtzii	Waterhouse	1845	1						"Philippine Islands."			eschscholtzii (Waterhouse, 1845)	split from M. schreibersii; moved from Vespertilionidae to Miniopteridae	Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Philippines	Asia	Indomalaya	NA	0	0	0	Miniopterus_eschscholtzii	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_eschscholtzii	1005109	23	Eschscholtz's Long-fingered Bat	Eschscholtz's Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	eschscholtzii	G. R. Waterhouse	1	Vespertilio Eschscholtzii	Waterhouse, G.R. 1845-04. Descriptions of species of Bats collectd in the Philippine Islands, and presented to the Society by H. Cuming, Esq. Proceedings of the Zoological Society of London 1845:3-10.	https://www.biodiversitylibrary.org/page/12862500				"Philippine Islands."			split from M. schreibersii; moved from Vespertilionidae to Miniopteridae	Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Philippines	Asia	Indomalaya	NE	0	0	0	Miniopterus_eschscholtzii	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		eschscholtzii	Waterhouse	1845	1	Proc. Zool. Soc. London	13 (1845): 3	Eschscholtz's Bent-winged Bat	None.	Philippine Islands	Philippines	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Formerly included in schreibersii or fuliginosus, but likely distinct; see Kusuminda et al. (2022), which strongly suggests that - once the holotype is examined - eschscholtzii will be recognized as a distinct species.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus eschscholtzii; Miniopterus fuliginosus eschscholtzii; Miniopterus eschscholtzii; Miniopterus eschscholtzii; eschscholtzii; Minioptéere d'Eschscholtz; Eschscholtz-Langflligelfledermaus; Minidptero de Eschscholtz; Eschscholtz’'s Bent-winged Bat; Eschscholtz's Long-fingered Bat; Eschscholtz's Bent-winged Bat; Eschscholtz's Bent-winged Bat; M. eschscholtzii
