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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L637	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos	Miniopterus brachytragos		[HMW] Miniopterus brachytragos Goodman et al., 2009 , “ Madagascar : Province de Mahajanga , Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National ], Forét d’Ambovonomby, 9% km NW Andranomavo , 16°28.2°S, 45°20.9’E, 200 m above sea level.” Formerly included in M. manavi . Monotypic.; [batnames2022] None.; [MDD2022] recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae; [IUCN] Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus brachytragos is one of these recently described species (Goodman et al. 2009).; [batnames2023] None.; [MDD2023] recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] None.; [MDD2025_2.2] recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae														brachytragos	Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus brachytragos is one of these recently described species (Goodman et al. 2009).			brachytragos	brachytragos			brachytragos S. M. Goodman, Maminirina, Bradman, Christidis, & B. Appleton, 2009						N/A																																								NA																											E84887F9FFC6D6480FC9FE281B1D31BC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	707	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFC6D6480FC9FE281B1D31BC.xml	Miniopterus brachytragos	Miniopteridae	Manaopterus	brachytragos	Goodman	2009	Minioptére a oreillons courts @fr | Nordwest-Madagaskar-Langfligelfledermaus @de | Miniopterode Madagascar @es | Broad-tragused Long-fingered Bat @en | Madagascar Bent-winged Bat @en	Miniopterus brachytragos Goodman et al., 2009 , “ Madagascar : Province de Mahajanga , Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National ], Forét d’Ambovonomby, 9% km NW Andranomavo , 16°28.2°S, 45°20.9’E, 200 m above sea level.” Formerly included in M. manavi . Monotypic.	Scattered localities on N & W Madagascar (including Nosy Komba).	Head-body c.45-49 mm, tail 38-43 mm, ear 9-11 mm, hindfoot 5-6 mm, forearm 35-38 mm; weight 2:9-6-3 g. Pelage of the Madagascar Long-fingered Bat is relatively short and not particularly dense. Dorsal fur is medium to slightly dark brown, and ventral hairs are tipped with dark buff, giving slightly mottled appearance. Wing membranes are medium brown, grading into slightly lighter brown on plagiopatagium and uropatagium; these membranes are attached to femur above ankle joint. Uropatagium has relatively short dense fur on most of its dorsal surface, being more obvious on proximal onehalf. Tragusis notably short (3-4 mm) and thick, with rounded to slightly pointed tip; distal part has a few long hairs, difficult to see with the naked eye.	Near native dry deciduous forests and disturbed gallery lowland humid forests from sea level up to elevations of c. 600 m .	The Madagascar Long-fingered Bat is expected to mainly eat soft insects captured in flight.	No information.	The Madagascar Long-fingered Batis nocturnal. It uses caves as daytime roosts. Echolocation calls have downward FM signals, with maximum frequencies of 85-128 kHz, minimum frequencies of 54-57 kHz, peak frequencies of 57-3— 61-7 kHz, durations of 2-6—4-3 milliseconds, and intervals of 56-1-122-5 milliseconds.	No information.	Classified as Least Concern on The IUCN Red Lust.	Goodman (2017d) | Goodman, Maminirina, Bradman et al. (2009) | Ramasindrazana et al. (2011)	https://zenodo.org/record/5735292/files/figure.png	30. Madagascar Long-fingered Bat Manaopterus brachytragos French: Minioptére a oreillons courts / German: Nordwest-Madagaskar-Langfligelfledermaus / Spanish: Minioptero de Madagascar Other common names: Broad-tragused Long-fingered Bat , Madagascar Bent-winged Bat Taxonomy. Miniopterus brachytragos Goodman et al., 2009 , “ Madagascar : Province de Mahajanga , Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National ], Forét d’Ambovonomby, 9% km NW Andranomavo , 16°28.2°S, 45°20.9’E, 200 m above sea level.” Formerly included in M. manavi . Monotypic. Distribution. Scattered localities on N & W Madagascar (including Nosy Komba). Descriptive notes. Head-body c.45-49 mm, tail 38-43 mm, ear 9-11 mm, hindfoot 5-6 mm, forearm 35-38 mm; weight 2:9-6-3 g. Pelage of the Madagascar Long-fingered Bat is relatively short and not particularly dense. Dorsal fur is medium to slightly dark brown, and ventral hairs are tipped with dark buff, giving slightly mottled appearance. Wing membranes are medium brown, grading into slightly lighter brown on plagiopatagium and uropatagium; these membranes are attached to femur above ankle joint. Uropatagium has relatively short dense fur on most of its dorsal surface, being more obvious on proximal onehalf. Tragusis notably short (3-4 mm) and thick, with rounded to slightly pointed tip; distal part has a few long hairs, difficult to see with the naked eye. Habitat. Near native dry deciduous forests and disturbed gallery lowland humid forests from sea level up to elevations of c. 600 m . Food and Feeding. The Madagascar Long-fingered Bat is expected to mainly eat soft insects captured in flight. Breeding. No information. Activity patterns. The Madagascar Long-fingered Batis nocturnal. It uses caves as daytime roosts. Echolocation calls have downward FM signals, with maximum frequencies of 85-128 kHz, minimum frequencies of 54-57 kHz, peak frequencies of 57-3— 61-7 kHz, durations of 2-6—4-3 milliseconds, and intervals of 56-1-122-5 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Goodman (2017d), Goodman, Maminirina, Bradman et al. (2009), Ramasindrazana et al. (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Miniopteridae	Miniopterus brachytragos	Miniopterus		brachytragos	Goodman, Maminirina, Bradman, Christidis, & Appleton	2009	0	Am. Mus. Novitates	######	Broad-tragused Long-fingered Bat	None.	Madagascar, Province de Hahajanga, Reserve Naturelle Integrale de Namoroka	Madagascar.	Not listed.	Least Concern	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus brachytragos	23	Madagascar Long-fingered Bat	Broad-tragused Long-fingered Bat|Madagascar Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	brachytragos	Goodman, Maminirina, Bradman, Christidis, & Appleton	2009	0	Miniopterus_brachytragos	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. (2009). The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates, 2009(3669), 9.	http://digitallibrary.amnh.org/handle/2246/6028	FMNH 175840		"Madagascar: Province de Mahajanga, Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National], Forêt d'Ambovonomby, 26 km NW Andranomavo, 16°28.2'S, 45°20.9'E, 200 m above sea level."	-16.47	45.348	brachytragos Goodman, Maminirina, Bradman, Christidis, & Appleton, 2009	recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. (2009). The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates, 2009(3669), 1-34.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.	Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_brachytragos	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Miniopterus brachytragos	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MINIOPTERIDAE	Miniopterus	brachytragos	Goodman, Maminirina, Bradman, Christidis &; Appleton, 2009	Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus brachytragos is one of these recently described species (Goodman et al. 2009).	100000000	Miniopterus brachytragos	Least Concern		2017	2016-05-02 00:00:00 UTC	3.1	English	This newly described species is known from seven sites in the northern and western portions of Madagascar (Goodman &; Ramasindrazana 2013; Goodman unpublished data), many of which are within existing protected areas. This species is apparently not forest-dependant and by extrapolation not subjected to any important habitat degradation or other human pressures.	A species that at least in the west uses caves for day roost sites, many of these sites are in vicinity of native forests, specifically within or adjacent to dry deciduous forest. ;It is known from two localities in more mesic humid forests. ;It presumably forages in areas that are more open or close to the forest edge and it is suspected that this species is not forest dependant.		Not known.	Unknown	Known from different sites in northern and western Madagascar and across an elevational range from 5 to 600 m above sea level (Goodman &; Ramasindrazana 2013).		Terrestrial	This species is known from several protected areas: Bemaraha, Loky-Manambato, Masoala and Namoroka.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		brachytragos	Goodman, Maminirina, Bradman, Christidis, & Appleton	2009	0	Am. Mus. Novitates	########	Broad-tragused Long-fingered Bat	None.	Madagascar, Province de Hahajanga, Reserve Naturelle Integrale de Namoroka	Madagascar.	Not listed.	Least Concern	None.	Miniopterus brachytragos	1005107	23	Madagascar Long-fingered Bat	Broad-tragused Long-fingered Bat|Madagascar Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	brachytragos	Goodman, Maminirina, Bradman, Christidis, & Appleton	2009	0	Miniopterus_brachytragos	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. (2009). The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates, 2009(3669), 9.	http://digitallibrary.amnh.org/handle/2246/6028	FMNH 175840		"Madagascar: Province de Mahajanga, Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National], Forêt d'Ambovonomby, 26 km NW Andranomavo, 16°28.2'S, 45°20.9'E, 200 m above sea level."	-16.47	45.34833	brachytragos Goodman, Maminirina, Bradman, Christidis, & Appleton, 2009	recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. (2009). The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates, 2009(3669), 1-34.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_brachytragos	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_brachytragos	1005107	23	Madagascar Long-fingered Bat	Broad-tragused Long-fingered Bat|Madagascar Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	brachytragos	S. M. Goodman, Maminirina, Bradman, Christidis, & B. Appleton	0	Miniopterus brachytragos	Goodman, S.M., Maminirina, C.P., Bradman, H.M., Christidis, L.L. and Appleton, B. 2009-11-30. The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: _Miniopterus_) on Madagascar. American Museum Novitates 3669:1-34.	https://www.biodiversitylibrary.org/page/59795850	FMNH:Mamm:175840	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2674608	"Madagascar: Province de Mahajanga, Réserve Naturelle Intégrale de Namoroka [status subsequently changed to Parc National], Forêt d'Ambovonomby, 26 km NW Andranomavo, 16°28.2'S, 45°20.9'E, 200 m above sea level."	-16.47	45.34833	recently described for populations previously included under M. manavi; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Maminirina, C. P., Bradman, H. M., Christidis, L., & Appleton, B. (2009). The use of molecular phylogenetic and morphological tools to identify cryptic and paraphyletic species: Examples from the diminutive long-fingered bats (Chiroptera: Miniopteridae: Miniopterus) on Madagascar. American Museum Novitates, 2009(3669), 1-34.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_brachytragos	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		brachytragos	Goodman, Maminirina, Bradman, Christidis & Appleton	2009	0	Am. Mus. Novitates	########	Broad-tragused Long-fingered Bat	None.	Madagascar, Province de Hahajanga, Reserve Naturelle Integrale de Namoroka	Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81629758/95642235/' target='_blank'>Least Concern</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus brachytragos; Miniopterus brachytragos; Miniopterus brachytragos; Miniopterus brachytragos; Miniopterus brachytragos; brachytragos; Minioptére a oreillons courts; Nordwest-Madagaskar-Langfligelfledermaus; Miniopterode Madagascar; Broad-tragused Long-fingered Bat; Madagascar Bent-winged Bat; Madagascar Long-fingered Bat; Broad-tragused Long-fingered Bat; Madagascar Bent-winged Bat; Broad-tragused Long-fingered Bat; M. brachytragos