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authorityParentheses originalNameCombination authoritySpeciesCitation authoritySpeciesLink typeVoucher typeKind typeVoucherURIs typeLocality typeLocalityLatitude typeLocalityLongitude taxonomyNotes taxonomyNotesCitation distributionNotes distributionNotesCitation subregionDistribution countryDistribution continentDistribution biogeographicRealm iucnStatus extinct domestic flagged CMW_sciName diffSinceCMW MSW3_matchtype MSW3_sciName diffSinceMSW3 docOrigin_batnames2025 Family Genus Subgenus Species Author Date Parentheses (1=author & date in parentheses) Citation Pages Common Name Synonyms Type Locality Distribution CITES IUCN Comments column3781 column3791 subtribe CONCAT_ALTNAMES line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L635 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A Miniopterus schreibersii [synonym of] Miniopterus natalensis arenarius Miniopterus natalensis arenarius Miniopterus arenarius Miniopterus natalensis Miniopterus arenarius N/A Miniopterus arenarius Miniopterus arenarius Miniopterus arenarius Miniopterus arenarius Miniopterus arenarius [HMW] is still unsolved until material from the type locality is studied genetically. Monotypic.; [MDD2022] split from M. natalensis; moved from Vespertilionidae to Miniopteridae; [batnames2023] Formerly included in schreibersii (e.g., ) and subsequently natalensis (see Koopman, 1994; Simmons, 2005), but apparently distinct (pers. comm. Webala and Demos, 2023); see Å rÃ¡mek et al. (2013), Demos et al. (2020), and Monadjem et al., (2020). Note that Å rÃ¡mek et al. (2013) did not compare the type material of arenarius to the specimens of cf. arenarius in their study. Additional research is needed.; [MDD2023] split from M. natalensis; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] split from M. natalensis; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] Formerly included in schreibersii (e.g., ) and subsequently natalensis (see Koopman, 1994; Simmons, 2005), but apparently distinct (pers. comm. Webala and Demos, 2023); see Å rÃ¡mek et al. (2013), Demos et al. (2020), and Monadjem et al., (2020). Note that Å rÃ¡mek et al. (2013) did not compare the type material of arenarius to the specimens of cf. arenarius in their study. Additional research is needed.; [MDD2025_2.2] split from M. natalensis; moved from Vespertilionidae to Miniopteridae arenarius arenarius arenarius arenarius E. Heller, 1912 N/A NA E84887F9FFDBD6550FF1FE1C183A3F09 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Miniopteridae_674.pdf.imf hash://md5/1471ff81ffd6d6580a4affec112f3619 704 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFDBD6550FF1FE1C183A3F09.xml Miniopterus arenarius Miniopteridae Miniopterus arenarius Heller 1912 Minioptére des sables @fr | Sandfarbene Langfligelfledermaus @de | Miniéptero delas arenas @es | Sandy Bent-winged Bat @en is still unsolved until material from the type locality is studied genetically. Monotypic. SW Arabian Peninsula in SW Saudi Arabia and W Yemen and E Africa in Ethiopia , E South Sudan , Somalia , and Kenya . This distribution is temporary although and pending of genetic studies including a complete representation of the species in the analysis. Head—body 55 mm , tail 47 mm , ear 11 mm , hindfoot 9 mm , forearm 45 mm (42-46 mm in South Sudan ; larger than 45 mm in Ethiopia ). Size of the Sandy Long-fingered Bat is similar to the Natal Long-fingered Bat ( M. natalensis ), but itis paler, with shorter pelage, and its skull has broader braincase, with I* greatly exceeding I’ in size. Dorsal pelage is seal-brown, with darker clove-brown head and lighter walnut-brown rump. Hairs are only slightly darker (dark slate) at bases, underparts are washed drab gray, and ears and wing membranes are blackish, the latter narrowly bordered in white. Mainly Acacia ( Fabaceae )— Commiphora ( Burseraceae ) brushlands and thickets ( Kenya ), possibly forest-savanna mosaics and Afromontane vegetation, up to elevations of ¢. 1400 m . The type specimen of the Sandy Long-fingered Bat was caught in a large crevice among granite boulders on the summit of a small hill; another specimen was shot at dusk among some acacia trees in the same vicinity. In Ethiopia ,it was found at ¢. 1300 m in a cave on the steep, rocky side of a river gorge covered by dense mixed Acacia and broadleaved scrubs and a narrow zone of more luxuriant vegetation on the riverbanks and also in an area dominated by Acacia scrub but near a river with welldeveloped riverine forest with large Ficus ( Moraceae ) trees at 1400 m . There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight. No information. Sandy Long-fingered Bats are thought to be nocturnal. They roost in caves. No information. Not assessed on The [UCN Red List. The Sandy Long-fingered Bat was recently separated from Schreibers’s Long-fingered Bat ( M. schreibersii ), which is classified as Near Threatened. Benda, Al-Jumaily et al. (2011) | Heller (1912) | Koopman (1975) | Largen et al. (1974) | Lavrenchenko et al. (2004) | Musila et al. (2019) | Nader & Kock (1987) | Sramek et al. (2013) https://zenodo.org/record/5735268/files/figure.png 22. Sandy Long-fingered Bat Miniopterus arenarius French: Minioptére des sables / German: Sandfarbene Langfligelfledermaus / Spanish: Miniéptero de las arenas Other common names: Sandy Bent-winged Bat Taxonomy. Miniopterus natalensis arenarius Heller, 1912 , “Guaso Nyuki (= Nanyuki), Northern Guaso Nyiro [= Ewaso Ng’iro] River, British East Africa [= Kenya ].” Previously a subspecies of M. schreibersii or M. natalensis , but recent genetic studies have shown that specimens from the Arabian Peninsula and Ethiopia (which were assigned provisionally to M. arenarius) are quite differentiated from M. schreibersii and M. natalensis . Taxonomy is still unsolved until material from the type locality is studied genetically. Monotypic. Distribution. SW Arabian Peninsula in SW Saudi Arabia and W Yemen and E Africa in Ethiopia , E South Sudan , Somalia , and Kenya . This distribution is temporary although and pending of genetic studies including a complete representation of the species in the analysis. Descriptive notes. Head—body 55 mm , tail 47 mm , ear 11 mm , hindfoot 9 mm , forearm 45 mm (42-46 mm in South Sudan ; larger than 45 mm in Ethiopia ). Size of the Sandy Long-fingered Bat is similar to the Natal Long-fingered Bat ( M. natalensis ), but itis paler, with shorter pelage, and its skull has broader braincase, with I* greatly exceeding I’ in size. Dorsal pelage is seal-brown, with darker clove-brown head and lighter walnut-brown rump. Hairs are only slightly darker (dark slate) at bases, underparts are washed drab gray, and ears and wing membranes are blackish, the latter narrowly bordered in white. Habitat. Mainly Acacia ( Fabaceae )— Commiphora ( Burseraceae ) brushlands and thickets ( Kenya ), possibly forest-savanna mosaics and Afromontane vegetation, up to elevations of ¢. 1400 m . The type specimen of the Sandy Long-fingered Bat was caught in a large crevice among granite boulders on the summit of a small hill; another specimen was shot at dusk among some acacia trees in the same vicinity. In Ethiopia ,it was found at ¢. 1300 m in a cave on the steep, rocky side of a river gorge covered by dense mixed Acacia and broadleaved scrubs and a narrow zone of more luxuriant vegetation on the riverbanks and also in an area dominated by Acacia scrub but near a river with welldeveloped riverine forest with large Ficus ( Moraceae ) trees at 1400 m . Food and Feeding. There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight. Breeding. No information. Activity patterns. Sandy Long-fingered Bats are thought to be nocturnal. They roost in caves. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCN Red List. The Sandy Long-fingered Bat was recently separated from Schreibers’s Long-fingered Bat ( M. schreibersii ), which is classified as Near Threatened. Bibliography. Benda, Al-Jumaily et al. (2011), Heller (1912), Koopman (1975), Largen et al. (1974), Lavrenchenko et al. (2004), Musila et al. (2019), Nader & Kock (1987), Sramek et al. (2013). Miniopteridae synonym of Miniopterus natalensis Miniopterus natalensis Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Miniopterus arenarius 23 Sandy Long-fingered Bat Sandy Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA MINIOPTERIDAE NA NA Miniopterus NA arenarius E. Heller 1912 0 Miniopterus_natalensis_arenarius Heller, E. (1912). New Races of Insectivores, Bats, and Lemurs from British East Africa. Smithsonian Miscellaneous Collections, 60(12), 2. https://repository.si.edu/bitstream/handle/10088/23473/SMC_60_Heller_1912_12_1-13.pdf?sequence=1&isAllowed=y USNM 181811 "Guaso Nyuki (= Nanyuki), Northern Guaso Nyiro [= Ewaso Ng'iro] River, British East Africa [= Kenya]." arenarius Heller, 1912 split from M. natalensis; moved from Vespertilionidae to Miniopteridae Monadjem, A., Goodman, S. M., Stanley, W. T., & Appleton, B. (2013). A cryptic new species of Miniopterus from south-eastern Africa based on molecular and morphological characters. Zootaxa, 3746(1), 123-142.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561. Saudi Arabia|Yemen|Ethiopia|South Sudan|Somalia|Kenya Africa|Asia Afrotropic NA 0 0 0 Miniopterus_arenarius 0 unmatched NA 1 Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Miniopteridae Miniopterus arenarius Heller 1912 0 Smithsonian Misc. Collect. 60(12): 2 Sandy Long-fingered Bat None. British East Africa [Kenya], Northern Guaso Nyiro [Ewaso Ng'iro] River, Guaso Nyuki [Nanyuki] SW Saudi Arabia, W Yemen, and E Africa from E South Sudan, Ethiopia, through Somalia and Kenya Not listed. Least Concern under Miniopterus natalensis Formerly included in schreibersii (e.g., ) and subsequently natalensis (see Koopman, 1994; Simmons, 2005), but apparently distinct (pers. comm. Webala and Demos, 2023); see Å rÃ¡mek et al. (2013), Demos et al. (2020), and Monadjem et al., (2020). Note that Å rÃ¡mek et al. (2013) did not compare the type material of arenarius to the specimens of cf. arenarius in their study. Additional research is needed. Miniopterus arenarius 1005104 23 Sandy Long-fingered Bat Sandy Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA Miniopteridae NA NA Miniopterus NA arenarius E. Heller 1912 0 Miniopterus_natalensis_arenarius Heller, E. (1912). New Races of Insectivores, Bats, and Lemurs from British East Africa. Smithsonian Miscellaneous Collections, 60(12), 2. https://repository.si.edu/bitstream/handle/10088/23473/SMC_60_Heller_1912_12_1-13.pdf?sequence=1&isAllowed=y USNM 181811 "Guaso Nyuki (= Nanyuki), Northern Guaso Nyiro [= Ewaso Ng'iro] River, British East Africa [= Kenya]." arenarius Heller, 1912 split from M. natalensis; moved from Vespertilionidae to Miniopteridae Monadjem, A., Goodman, S. M., Stanley, W. T., & Appleton, B. (2013). A cryptic new species of Miniopterus from south-eastern Africa based on molecular and morphological characters. Zootaxa, 3746(1), 123-142.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561. Saudi Arabia|Yemen|Ethiopia|South Sudan|Somalia|Kenya Africa|Asia Afrotropic NA 0 0 0 Miniopterus_arenarius 0 unmatched NA 1 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Miniopterus_arenarius 1005104 23 Sandy Long-fingered Bat Sandy Bent-winged Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yangochiroptera NA NA Vespertilionoidea Miniopteridae NA NA Miniopterus NA arenarius E. Heller 0 Miniopterus natalensis arenarius Heller, E. 1912-11-04. New races of insectivores, bats, and lemurs from British East Africa. Smithsonian Miscellaneous Collections 60(12):1-13. https://www.biodiversitylibrary.org/page/8910532 USNM:MAMM:181811 holotype http://n2t.net/ark:/65665/35dc9f22a-9942-4267-a00e-5fbf33495615 "Guaso Nyuki (= Nanyuki), Northern Guaso Nyiro [= Ewaso Ng'iro] River, British East Africa [= Kenya]." split from M. natalensis; moved from Vespertilionidae to Miniopteridae Monadjem, A., Goodman, S. M., Stanley, W. T., & Appleton, B. (2013). A cryptic new species of Miniopterus from south-eastern Africa based on molecular and morphological characters. Zootaxa, 3746(1), 123-142.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.|Demos, T. C., Webala, P. W., Goodman, S. M., Peterhans, J. C. K., Lutz, H. L., Agwanda, B. R., ... & Patterson, B. D. (2023). Ultraconserved elements resolve phylogenetic relationships and biogeographic history of African-Malagasy bent-winged bats (Miniopterus). Molecular Phylogenetics and Evolution, 188, 107890. Saudi Arabia|Yemen|Ethiopia|South Sudan|Somalia|Kenya Africa|Asia Afrotropic NE 0 0 0 Miniopterus_arenarius 0 unmatched NA 1 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Miniopteridae Miniopterus arenarius Heller 1912 0 Smithsonian Misc. Collect. 60(12): 2 Sandy Long-fingered Bat None. British East Africa [Kenya], Northern Guaso Nyiro [Ewaso Ng'iro] River, Guaso Nyuki [Nanyuki] SW Saudi Arabia, W Yemen, and E Africa from E South Sudan, Ethiopia, through Somalia and Kenya Not Listed Not Evaluated Formerly included in schreibersii (e.g., ) and subsequently natalensis (see Koopman, 1994; Simmons, 2005), but apparently distinct (pers. comm. Webala and Demos, 2023); see Å rÃ¡mek et al. (2013), Demos et al. (2020), and Monadjem et al., (2020). Note that Å rÃ¡mek et al. (2013) did not compare the type material of arenarius to the specimens of cf. arenarius in their study. Additional research is needed. Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Miniopterus arenarius; Miniopterus natalensis; Miniopterus arenarius; ; arenarius; Minioptére des sables; Sandfarbene Langfligelfledermaus; Miniéptero delas arenas; Sandy Bent-winged Bat; Sandy Long-fingered Bat; Sandy Bent-winged Bat;; M. arenarius