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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L634	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis	Miniopterus ambohitrensis		[HMW] Miniopterus ambohitrensis Goodman et al, 2015 , “ Madagascar : Province d’Antsiranana, Parc National de la Montagne d’Ambre, 5.5 km SWJoffreville ( Ambohitra ), Station Forestiere des Roussettes , 12°31'37.978, 49° 10'19.1E, 1600'm.” Some specimens belonging to M. ambohitrensis were included in the original description of M. aelleni , which is its sister species. Monotypic.; [batnames2022] None.; [MDD2022] recently described; moved from Vespertilionidae to Miniopteridae; [IUCN] Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus ambohitrensis is one of these recently described species (Goodman et al. 2015). It was originally separated from the M. manavi under the name M. aelleni (Goodman et al. 2009), which was subsequently found to be composed of two divergent taxa (Goodman et al. 2015).; [batnames2023] None.; [MDD2023] recently described; moved from Vespertilionidae to Miniopteridae; [MDD2025_2.0] recently described; moved from Vespertilionidae to Miniopteridae; [batnames2025_1.7] None.; [MDD2025_2.2] recently described; moved from Vespertilionidae to Miniopteridae														ambohitrensis	Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus ambohitrensis is one of these recently described species (Goodman et al. 2015). It was originally separated from the M. manavi under the name M. aelleni (Goodman et al. 2009), which was subsequently found to be composed of two divergent taxa (Goodman et al. 2015).			ambohitrensis	ambohitrensis			ambohitrensis S. M. Goodman, Ramasindrazana, K. M. Naughton, & B. Appleton, 2015						N/A																																								NA																											E84887F9FFC6D6490FCCF80314203471	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Miniopteridae_674.pdf.imf	hash://md5/1471ff81ffd6d6580a4affec112f3619	707	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/E8/48/87/E84887F9FFC6D6490FCCF80314203471.xml	Miniopterus ambohitrensis	Miniopteridae	Miniopterus	ambohitrensis	Bonaparte	2015	Minioptére d Ambohitra @fr | Montagne-d Ambre-Langfllgelfledermaus @de | Miniéptero de Ambohitra @es | Ambohitra Long-fingered Bat @en | Montagne d Ambre Bent-winged Bat @en	Miniopterus ambohitrensis Goodman et al, 2015 , “ Madagascar : Province d’Antsiranana, Parc National de la Montagne d’Ambre, 5.5 km SWJoffreville ( Ambohitra ), Station Forestiere des Roussettes , 12°31'37.978, 49° 10'19.1E, 1600'm.” Some specimens belonging to M. ambohitrensis were included in the original description of M. aelleni , which is its sister species. Monotypic.	Known only from four sites on N & C Madagascar (Marojejy and Montagne d’Ambre national parks, Ambohitantely Reserve, and Bemanevika).	Head-body ¢.53-55 mm, tail 40-47 mm, ear 10-12 mm, hindfoot 5-7 mm, forearm 37-42 mm; weight 5-3-7-7 g. The Montagne d’Ambre Long-fingered Bat has mixed medium and dark brown pelage. Pectoral area and head often have varying amounts of rufous brown fur. Tragus is moderately long (6-7 mm), slightly broader at base, with slight constriction along mid-shaft; distal portion is medially deflected; and tip is slightly rounded. It is very similar to Aellen’s Long-fingered Bat ( M. aelleni ), except for its rufous brown areas of pelage, slightly larger size, and slightly shorter tragus.	Upland mesic or montane humid forests at elevations of ¢.800-1600 m.	There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight.	Evidence of recent reproductive activity of Montagne d’Ambre Long-fingered Bats was found in early April, with one female lactating and having single placental scar, another female with large mammae and showing signs of estrus, severalfemales with large mammae and missing fur, and a male with testes measuring 4 x 3 mm and epididymides partially convoluted. The female record of late seasonal breeding and apparent postpartum estrus might indicate a period of reproductive delayafter mating, as known for some African species of Miniopterus .	The Montagne d’Ambre Long-fingered Bat is thought to be nocturnal. It was found roosting in caves and is probably dependent on caves and rock shelters for day roosts. Echolocation calls have downward FM signals, with maximum frequencies of 64-78-2 kHz, minimum frequencies of 47-7-51-6 kHz, peak frequencies of 53-1-55-4 kHz, durations of 2-5-3-4 milliseconds, and intervals of 61-4-73-4 milliseconds.	There is evidence to suggest that some populations migrate elevationally to some degree.	Classified as Least Concern on The IUCN Red Last.	Goodman (2017¢c) | Goodman, Ramasindrazana et al. (2015) | Ramasindrazana et al. (2011)	https://zenodo.org/record/5735294/files/figure.png	31. Montagne d’Ambre Long-fingered Bat Miniopterus ambohitrensis French: Minioptére dAmbohitra / German: Montagne-dAmbre-Langfllgelfledermaus / Spanish: Miniéptero de Ambohitra Other common names: Ambohitra Long-fingered Bat , Montagne dAmbre Bent-winged Bat Taxonomy. Miniopterus ambohitrensis Goodman et al, 2015 , “ Madagascar : Province d’Antsiranana, Parc National de la Montagne d’Ambre, 5.5 km SWJoffreville ( Ambohitra ), Station Forestiere des Roussettes , 12°31'37.978, 49° 10'19.1E, 1600'm.” Some specimens belonging to M. ambohitrensis were included in the original description of M. aelleni , which is its sister species. Monotypic. Distribution. Known only from four sites on N & C Madagascar (Marojejy and Montagne d’Ambre national parks, Ambohitantely Reserve, and Bemanevika). Descriptive notes. Head-body ¢.53-55 mm, tail 40-47 mm, ear 10-12 mm, hindfoot 5-7 mm, forearm 37-42 mm; weight 5-3-7-7 g. The Montagne d’Ambre Long-fingered Bat has mixed medium and dark brown pelage. Pectoral area and head often have varying amounts of rufous brown fur. Tragus is moderately long (6-7 mm), slightly broader at base, with slight constriction along mid-shaft; distal portion is medially deflected; and tip is slightly rounded. It is very similar to Aellen’s Long-fingered Bat ( M. aelleni ), except for its rufous brown areas of pelage, slightly larger size, and slightly shorter tragus. Habitat. Upland mesic or montane humid forests at elevations of ¢.800-1600 m. Food and Feeding. There is no specific information available for this species, but other long-fingered bats mainly eat soft insects captured in flight. Breeding. Evidence of recent reproductive activity of Montagne d’Ambre Long-fingered Bats was found in early April, with one female lactating and having single placental scar, another female with large mammae and showing signs of estrus, severalfemales with large mammae and missing fur, and a male with testes measuring 4 x 3 mm and epididymides partially convoluted. The female record of late seasonal breeding and apparent postpartum estrus might indicate a period of reproductive delayafter mating, as known for some African species of Miniopterus . Activity patterns. The Montagne d’Ambre Long-fingered Bat is thought to be nocturnal. It was found roosting in caves and is probably dependent on caves and rock shelters for day roosts. Echolocation calls have downward FM signals, with maximum frequencies of 64-78-2 kHz, minimum frequencies of 47-7-51-6 kHz, peak frequencies of 53-1-55-4 kHz, durations of 2-5-3-4 milliseconds, and intervals of 61-4-73-4 milliseconds. Movements, Home range and Social organization. There is evidence to suggest that some populations migrate elevationally to some degree. Status and Conservation. Classified as Least Concern on The IUCN Red Last. Bibliography. Goodman (2017¢c), Goodman, Ramasindrazana et al. (2015), Ramasindrazana et al. (2011).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Miniopteridae	Miniopterus ambohitrensis	Miniopterus		ambohitrensis	Goodman, Ramasindrazana, Naughton & Appleton	2015	0	Zootaxa	3936(4): 543	Ambohitra Long-fingered Bat	None.	Madagascar: Province d&#39;Antsiranana, Parc National de la Montagne d&#39;Ambre, 5.5 km SW Joffreville (Ambohitra), Station Foresti&egrave;re des Roussettes, 12 &deg; 31 &#39; 37.3 &quot; S, 49 &deg; 10 &#39; 19.1 &quot; E, 1000 m	N and C Madagascar	Not listed.	Least Concern	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Miniopterus ambohitrensis	23	Montagne d'Ambre Long-fingered Bat	Ambohitra Long-fingered Bat|Montagne d'Ambre Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MINIOPTERIDAE	NA	NA	Miniopterus	NA	ambohitrensis	Goodman, Ramasindrazana, Naughton, & Appleton	2015	0	Miniopterus_ambohitrensis	Goodman, S. M., Ramasindrazana, B., Naughton, K. M., & Appleton, B. (2015). Description of a new species of the Miniopterus aelleni group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa, 3936(4), 538.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3936.4.4	FMNH 202450		"Madagascar: Province d'Antsiranana, Parc National de la Montagne d'Ambre, 5.5 km SW Joffreville (Ambohitra), Station Forestière des Roussettes, 12°31'37.3"S, 49°10'19.1"E, 1000 m."	-12.53	49.172	ambohitrensis Goodman, Ramasindrazana, Naughton, & Appleton, 2015	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Ramasindrazana, B., Naughton, K. M., & Appleton, B. (2015). Description of a new species of the Miniopterus aelleni group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa, 3936(4), 538-58.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.	Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_ambohitrensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Miniopterus ambohitrensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MINIOPTERIDAE	Miniopterus	ambohitrensis	Goodman, Ramasindrazana, Naughton &; Appleton, 2015	Peterson et al. (1995) noted considerable morphological variation in the widespread Malagasy species Miniopterus manavi . Recent work using morphological and molecular characters found that animals previously assigned to this species are distinctly more speciose than previously understood (Christidis et al. 2014). Further, not all of the "manavi " like taxa are closely related to M . manavi. Miniopterus ambohitrensis is one of these recently described species (Goodman et al. 2015). It was originally separated from the M. manavi under the name M. aelleni (Goodman et al. 2009), which was subsequently found to be composed of two divergent taxa (Goodman et al. 2015).	100000000	Miniopterus ambohitrensis	Least Concern		2017	2016-05-02 00:00:00 UTC	3.1	English	This newly described species is known from several sites in the northern portion of Madagascar (Goodman et al. 2015), most of which are montane and apparently not subjected to any important habitat degradation or other human pressures. It is notably common at several of these sites.	A species that uses caves for day roost sites, many of these caves are in close proximity to native forests. ;In certain cases day roosting sites are in isolated forest fragments. ;Most of these sites are within or adjacent to montane humid forest, although one is found in the upper end of lowland humid forest. It is suspected that this species is not forest dependant.		Not known.	Unknown	On the basis of the recent species description, it occurs in montane areas of northern Madagascar and across an elevational range from 810 to 1570 m above sea level (Goodman et al. 2015).		Terrestrial	This species is known from several protected areas, which include Marojejy, Montagne d’Ambre, Ambohitantely, and Bemanevika.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Miniopteridae	Miniopterus		ambohitrensis	Goodman, Ramasindrazana, Naughton & Appleton	2015	0	Zootaxa	3936(4): 543	Ambohitra Long-fingered Bat	None.	Madagascar: Province d&#39;Antsiranana, Parc National de la Montagne d&#39;Ambre, 5.5 km SW Joffreville (Ambohitra), Station Foresti&egrave;re des Roussettes, 12 &deg; 31 &#39; 37.3 &quot; S, 49 &deg; 10 &#39; 19.1 &quot; E, 1000 m	N and C Madagascar	Not listed.	Least Concern	None.	Miniopterus ambohitrensis	1005103	23	Montagne d'Ambre Long-fingered Bat	Ambohitra Long-fingered Bat|Montagne d'Ambre Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Miniopteridae	NA	NA	Miniopterus	NA	ambohitrensis	Goodman, Ramasindrazana, Naughton, & Appleton	2015	0	Miniopterus_ambohitrensis	Goodman, S. M., Ramasindrazana, B., Naughton, K. M., & Appleton, B. (2015). Description of a new species of the Miniopterus aelleni group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa, 3936(4), 538.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3936.4.4	FMNH 202450		"Madagascar: Province d'Antsiranana, Parc National de la Montagne d'Ambre, 5.5 km SW Joffreville (Ambohitra), Station Forestière des Roussettes, 12°31'37.3"S, 49°10'19.1"E, 1000 m."	-12.527	49.17197	ambohitrensis Goodman, Ramasindrazana, Naughton, & Appleton, 2015	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Ramasindrazana, B., Naughton, K. M., & Appleton, B. (2015). Description of a new species of the Miniopterus aelleni group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa, 3936(4), 538-58.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_ambohitrensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Miniopterus_ambohitrensis	1005103	23	Montagne d'Ambre Long-fingered Bat	Ambohitra Long-fingered Bat|Montagne d'Ambre Bent-winged Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Miniopteridae	NA	NA	Miniopterus	NA	ambohitrensis	S. M. Goodman, Ramasindrazana, K. M. Naughton, & B. Appleton	0	Miniopterus ambohitrensis	Goodman, S.M., Ramasindrazana, B., Naughton, K.M. and Appleton, B. 2015-03-23. Description of a new species of the _Miniopterus aelleni_ group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa 3936(4):538-558.	https://doi.org/10.11646/zootaxa.3936.4.4	FMNH:Mamm:202450	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2701337	"Madagascar: Province d'Antsiranana, Parc National de la Montagne d'Ambre, 5.5 km SW Joffreville (Ambohitra), Station Forestière des Roussettes, 12°31'37.3"S, 49°10'19.1"E, 1000 m."	-12.527	49.17197	recently described; moved from Vespertilionidae to Miniopteridae	Goodman, S. M., Ramasindrazana, B., Naughton, K. M., & Appleton, B. (2015). Description of a new species of the Miniopterus aelleni group (Chiroptera: Miniopteridae) from upland areas of central and northern Madagascar. Zootaxa, 3936(4), 538-58.|Miller-Butterworth, C. M., Murphy, W. J., O'Brien, S. J., Jacobs, D. S., Springer, M. S., & Teeling, E. C. (2007). A family matter: conclusive resolution of the taxonomic position of the long-fingered bats, Miniopterus. Molecular Biology and Evolution, 24(7), 1553-1561.				Madagascar	Africa	Afrotropic	LC	0	0	0	Miniopterus_ambohitrensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Miniopteridae	Miniopterus		ambohitrensis	Goodman, Ramasindrazana, Naughton & Appleton	2015	0	Zootaxa	3936(4): 543	Ambohitra Long-fingered Bat	None.	Madagascar: Province d&#39;Antsiranana, Parc National de la Montagne d&#39;Ambre, 5.5 km SW Joffreville (Ambohitra), Station Foresti&egrave;re des Roussettes, 12 &deg; 31 &#39; 37.3 &quot; S, 49 &deg; 10 &#39; 19.1 &quot; E, 1000 m	N and C Madagascar	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/81633128/95642255/' target='_blank'>Least Concern</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Miniopterus ambohitrensis; Miniopterus ambohitrensis; Miniopterus ambohitrensis; Miniopterus ambohitrensis; Miniopterus ambohitrensis; ambohitrensis; Minioptére d Ambohitra; Montagne-d Ambre-Langfllgelfledermaus; Miniéptero de Ambohitra; Ambohitra Long-fingered Bat; Montagne d Ambre Bent-winged Bat; Montagne d'Ambre Long-fingered Bat; Ambohitra Long-fingered Bat; Montagne d'Ambre Bent-winged Bat; Ambohitra Long-fingered Bat; M. ambohitrensis