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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L601	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum	Macrophyllum macrophyllum		[MSW2] See Harrison (1975, Mammalian Species, 62).; [MSW3] See Harrison (1975). See Emmons (1997) for distribution map.; [HMW] Phyllostoma macrophyllum Schinz, 1821 , “In den Waldern von Brasilien .” Restricted by M. P. zu Wied-Neuwied in 1826 as “Flusse Mucuri [= Mucuri River],” Bahia , Brazil . This species is monotypic.; [batnames2022] See Harrison (1975). See Emmons (1997) for distribution map.; [batnames2023] See Harrison (1975). See Emmons (1997) for distribution map.; [batnames2025_1.7] See Harrison (1975). See Emmons (1997) for distribution map.						nieuwiedii.			neuwiedii			macrophyllum 	macrophyllum - neuwiedii, nieuwiedii	macrophyllum, nieuwiedii		macrophyllum 	macrophyllum - neuwiedii, nieuwiedii	macrophyllum, nieuwiedii 	macrophyllum, nieuwiedii, neuwiedii	macrophyllum	macrophyllum - neuwiedii, nieuwiedii	macrophyllum (H. R. Schinz, 1821)|nieuwiedii J. E. Gray, 1838 [nomen novum]|neuwiedii P. Gervais, 1855 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Long-legged bat	S Mexico – N Argentina	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Macrophyllum macrophyllum	Brazil, Bahia, Rio Mucuri.	Schinz	1821	Das Thierreich, 1:163.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Long-legged bat	S Mexico – N Argentina	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Schinz	1821	Das Thierreich, 1:163.	See Harrison (1975, Mammalian Species, 62).	Tabasco (Mexico) to Peru, Bolivia, SE Brazil, and NE Argentina.	Brazil, Bahia, Rio Mucuri.		SCHINZ	1821	Size relatively small (forearm length, 35-38 mm; condylobasal length, 13-15 mm).	Distribution: Same as for genus.	No subspecies.		75	species	M. macrophyllum	SCHINZ	1821	Macrophyllum	genus	Macrophyllum macrophyllum				Size relatively small (forearm length, 35-38 mm; condylobasal length, 13-15 mm).	No subspecies.		1. M. macrophyllum (SCHINZ 1821).	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Phyllostominae		Macrophyllum macrophyllum	Macrophyllum		macrophyllum	Schinz	y	1821		Das Thierreich	1		163		Long-legged Bat	Brazil, Bahia, Río Mucuri.	Tabasco (Mexico) south to Peru, Bolivia, SE Brazil, Paraguay, and NE Argentina.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	neuwiedii Gervais, 1855; nieuwiedii Gray, 1838.	See Harrison (1975). See Emmons (1997) for distribution map.	03A687BCFFA0FFA013B9FDF2F78EF31B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	501	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFA0FFA013B9FDF2F78EF31B.xml	Macrophyllum macrophyllum	Phyllostomidae	Macrophyllum	macrophyllum		1821	Macrophylle pécheur @fr | Langbeinfledermaus @de | Macrofilode patas largas @es	Phyllostoma macrophyllum Schinz, 1821 , “In den Waldern von Brasilien .” Restricted by M. P. zu Wied-Neuwied in 1826 as “Flusse Mucuri [= Mucuri River],” Bahia , Brazil . This species is monotypic.	Extreme S Mexico ( Tabasco and E Chiapas ), S through Central America and South America E of Andes to N & E Bolivia , Paraguay , most of Brazil , and extreme NE Argentina .	Head—body 40-6 cm, tail 38-49 mm, ear 17-20 mm, hindfoot 13-16 mm, forearm 33-40 mm; weight 6-11 g. The Long-legged Bat is small and slender compared with other phyllostomines. Color is uniformly dark brown to blackish brown, with very little contrast between dorsum and venter. Fur is very long and dense. Legs and feet are markedly elongated, reminiscent of those of Greater Bulldog Bat (Noctilio leporinus). Wings are short and broad, and uropatagium is very broad, with distinctive rows of warts resembling an inverted menorah candelabrum. Tail is long and extends to edge of uropatagium, which almost has a straight edge. Face is very distinctive and impossible to confuse with any other bat. Broad, tall, and pointed noseleaf is very prominent, proportionately broadest of any phyllostomid. Lower edge of noseleaf is large and evidently free and unfused with upper lip. Chin has large central wart surrounded by several smaller circular warts. Eyes are brown. Face is hairy, and inside of earsis sparsely covered with hairs. Tragus is long, narrow, and triangular. Dental formulais12/2,C1/1,P2/3,M 3/3 ( x2 ) = 34. Chromosomal complement has 2n = 32 and FN = 56.	Humid tropical forests and tropical deciduous forests always associated with water bodies, usually small streamsin forests. Usually inhabits areas with little or no human disturbance, although one colony in Barro Colorado Island, Panama , roosts in a partially submerged boat some distance from the coast but shaded under forest.	Morphology has long hinted at a diet dominated by water-associated animals, predominantly insects. In Panama , Long-legged Bats feed on gerrid water striders, pyralid moths, and other volant insects associated with water bodies. They might be among the few examples of phyllostomids that are obligate insectivores because no evidence exists that they eat fruit or pollen. They always forage less than 1 m over water at relatively high speeds (15-30 km/h). Like other “fishing” bats, such as the Common Bulldog Bat and the Fish-eating Bat ( Myotis vivesi ), Long-legged Bats skim insects off the water surface with their hindclaws, being the only phyllostomids known to do this. L.onglegged Bats combine gleaning insects from the water surface with aerial pursuit of volant insects. They forage in groups of 3-10 individuals in small coves less than 1-5 m in diameter very close to shore. Foraging flights can last up to one hour. They remain in flight for long periods, resulting in total flight distance of as much of 35-47 km,although they remain relatively close to day roosts. Foraging areas can be as large as 41-5 ha.	Female Long-legged Bats carry one embryo. Seasonality of births likely follows insect availability to optimize survival of young.	Long-legged Bats leave their roost c.20 minutes after sunset and return ¢.30 minutes before sunrise. They remain active most of the night but stop all activity during heavy rain. Most individuals return to day roosts for short periods at night, but one female tended to remain in her foraging areas away from the roost all night until she returned at ¢.05:00 h. They seem to maintain several night roosts and live in caves, culverts under roads, and abandoned buildings.	[Long-legged Bats have relatively large home ranges compared with other phyllostomids. Median home range was 23-9 ha in Panama (range 7-3-150-3 ha). Home ranges tend to be distinctly elongated because individuals remain close to shore and always feed over water. Long-legged Bats form harems in their roosts, butlittle is known about their social organization beyond this. They can share roosts with several other bat species including the Common Vampire Bat ( Desmodus rotundus ), the Common Mustached Bat ( Pteronotus parnellii ), short-tailed bats ( Carollia spp.), and Pallas’s Long-tongued Bat ( Glossophaga soricina ). Colonies have 2-60 individuals.	Classified as Least Concern on The [UCN Red List. Nevertheless, because the Long-legged Bat relies on relatively undisturbed interface between water and well-preserved forest, is rare acrossits distribution, and seems to disappear from disturbed areas, some countries such as Mexico have listed it as threatened.	Gray (1838) | Harrison (1975a) | Meyer etal. (2005) | Weinbeer & Kalko (2007) | Weinbeer, Kalko & Jung (2013) | Weinbeer, Meyer & Kalko (2006) | Wied-Neuwied (1826)	https://zenodo.org/record/6458650/files/figure.png	24. Long-legged Bat Macrophyllum macrophyllum French: Macrophylle pécheur / German: Langbeinfledermaus / Spanish: Macrofilo de patas largas Taxonomy. Phyllostoma macrophyllum Schinz, 1821 , “In den Waldern von Brasilien .” Restricted by M. P. zu Wied-Neuwied in 1826 as “Flusse Mucuri [= Mucuri River],” Bahia , Brazil . This species is monotypic. Distribution. Extreme S Mexico ( Tabasco and E Chiapas ), S through Central America and South America E of Andes to N & E Bolivia , Paraguay , most of Brazil , and extreme NE Argentina . Descriptive notes. Head—body 40-6 cm, tail 38-49 mm, ear 17-20 mm, hindfoot 13-16 mm, forearm 33-40 mm; weight 6-11 g. The Long-legged Bat is small and slender compared with other phyllostomines. Color is uniformly dark brown to blackish brown, with very little contrast between dorsum and venter. Fur is very long and dense. Legs and feet are markedly elongated, reminiscent of those of Greater Bulldog Bat (Noctilio leporinus). Wings are short and broad, and uropatagium is very broad, with distinctive rows of warts resembling an inverted menorah candelabrum. Tail is long and extends to edge of uropatagium, which almost has a straight edge. Face is very distinctive and impossible to confuse with any other bat. Broad, tall, and pointed noseleaf is very prominent, proportionately broadest of any phyllostomid. Lower edge of noseleaf is large and evidently free and unfused with upper lip. Chin has large central wart surrounded by several smaller circular warts. Eyes are brown. Face is hairy, and inside of earsis sparsely covered with hairs. Tragus is long, narrow, and triangular. Dental formulais12/2,C1/1,P2/3,M 3/3 ( x2 ) = 34. Chromosomal complement has 2n = 32 and FN = 56. Habitat. Humid tropical forests and tropical deciduous forests always associated with water bodies, usually small streamsin forests. Usually inhabits areas with little or no human disturbance, although one colony in Barro Colorado Island, Panama , roosts in a partially submerged boat some distance from the coast but shaded under forest. Food and Feeding. Morphology has long hinted at a diet dominated by water-associated animals, predominantly insects. In Panama , Long-legged Bats feed on gerrid water striders, pyralid moths, and other volant insects associated with water bodies. They might be among the few examples of phyllostomids that are obligate insectivores because no evidence exists that they eat fruit or pollen. They always forage less than 1 m over water at relatively high speeds (15-30 km/h). Like other “fishing” bats, such as the Common Bulldog Bat and the Fish-eating Bat ( Myotis vivesi ), Long-legged Bats skim insects off the water surface with their hindclaws, being the only phyllostomids known to do this. L.onglegged Bats combine gleaning insects from the water surface with aerial pursuit of volant insects. They forage in groups of 3-10 individuals in small coves less than 1-5 m in diameter very close to shore. Foraging flights can last up to one hour. They remain in flight for long periods, resulting in total flight distance of as much of 35-47 km,although they remain relatively close to day roosts. Foraging areas can be as large as 41-5 ha. Breeding. Female Long-legged Bats carry one embryo. Seasonality of births likely follows insect availability to optimize survival of young. Activity patterns. Long-legged Bats leave their roost c.20 minutes after sunset and return ¢.30 minutes before sunrise. They remain active most of the night but stop all activity during heavy rain. Most individuals return to day roosts for short periods at night, but one female tended to remain in her foraging areas away from the roost all night until she returned at ¢.05:00 h. They seem to maintain several night roosts and live in caves, culverts under roads, and abandoned buildings. Movements, Home range and Social organization. [Long-legged Bats have relatively large home ranges compared with other phyllostomids. Median home range was 23-9 ha in Panama (range 7-3-150-3 ha). Home ranges tend to be distinctly elongated because individuals remain close to shore and always feed over water. Long-legged Bats form harems in their roosts, butlittle is known about their social organization beyond this. They can share roosts with several other bat species including the Common Vampire Bat ( Desmodus rotundus ), the Common Mustached Bat ( Pteronotus parnellii ), short-tailed bats ( Carollia spp.), and Pallas’s Long-tongued Bat ( Glossophaga soricina ). Colonies have 2-60 individuals. Status and Conservation. Classified as Least Concern on The [UCN Red List. Nevertheless, because the Long-legged Bat relies on relatively undisturbed interface between water and well-preserved forest, is rare acrossits distribution, and seems to disappear from disturbed areas, some countries such as Mexico have listed it as threatened. Bibliography. Gray (1838), Harrison (1975a), Meyer etal. (2005), Weinbeer & Kalko (2007), Weinbeer, Kalko & Jung (2013), Weinbeer, Meyer & Kalko (2006), Wied-Neuwied (1826).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Macrophyllum macrophyllum	Macrophyllum		macrophyllum	Schinz	1821	1	Das Thierreich	0.1549	Long-legged Bat	 neuwiedii Gervais, 1855; nieuwiedii Gray, 1838.	Brazil, Bahia, Río Mucuri.	Tabasco (Mexico) south to Peru, Bolivia, SE Brazil, Paraguay, and NE Argentina.	Not listed.	Least Concern	See Harrison (1975). See Emmons (1997) for distribution map.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Macrophyllum macrophyllum	23	Long-legged Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	PHYLLOSTOMINAE	MACROPHYLLINI	Macrophyllum	NA	macrophyllum	Schinz	1821	1						"In den Wäldern von Brasilien." Restricted by M. P. zu Wied-Neuwied in 1826 as "Flusse Mucuri [= Mucuri River]," Bahia, Brazil.			macrophyllum (Schinz, 1821)|nieuwiedii J. E. Gray, 1838	NA	NA	Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil|Paraguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Macrophyllum_macrophyllum	0	sciname match	Macrophyllum_macrophyllum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	12615	Macrophyllum macrophyllum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Macrophyllum	macrophyllum	(Schinz, 1821)		20000000	Macrophyllum macrophyllum	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	It lives in colonies with up to 70 individuals. This species is water dependent (Meyer et al . 2005) and it feeds on aquatic insects.	Deforestation is a localised threat to this species. There are no major threats to the population overall.	This species is rare in Argentina (Barquez pers. comm.) and Mexico (Arroyo-Cabrales pers. comm.).	Unknown	This species occurs in Tabasco (Mexico) south to Peru, Bolivia, Brazil, Paraguay and northeastern Argentina (Simmons 2005). It is also found in Venezuela, the Guianas and Suriname.	This species is not used.	Terrestrial	It is found in protected areas. In Mexico, this species is listed as threatened under NOM - 059 - SEMARNAT - 2001 (Arroyo-Cabrales pers. comm.).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Macrophyllum		macrophyllum	Schinz	1821	1	Das Thierreich	0.154861	Long-legged Bat	 neuwiedii Gervais, 1855; nieuwiedii Gray, 1838.	Brazil, Bahia, Río Mucuri.	Tabasco (Mexico) south to Peru, Bolivia, SE Brazil, Paraguay, and NE Argentina.	Not listed.	Least Concern	See Harrison (1975). See Emmons (1997) for distribution map.	Macrophyllum macrophyllum	1004966	23	Long-legged Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	PHYLLOSTOMINAE	MACROPHYLLINI	Macrophyllum	NA	macrophyllum	Schinz	1821	1						"In den Wäldern von Brasilien." Restricted by M. P. zu Wied-Neuwied in 1826 as "Flusse Mucuri [= Mucuri River]," Bahia, Brazil.			macrophyllum (Schinz, 1821)|nieuwiedii J. E. Gray, 1838	NA	NA				Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil|Paraguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Macrophyllum_macrophyllum	0	sciname match	Macrophyllum_macrophyllum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Macrophyllum_macrophyllum	1004966	23	Long-legged Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Phyllostominae	Macrophyllini	Macrophyllum	NA	macrophyllum	H. R. Schinz	1	Phyllostoma macrophyllum	Schinz, H.R. 1821. Das Thierreich eingetheilt nach dem Bau der Thiere als Grundlage ihrer Naturgeschichte und der vergleichenden Anatomie. J. G. Cotta'sche Buchhandlung, Stuttgart, 894 pp.	https://www.biodiversitylibrary.org/page/51528920	untraced (number not known)	nonexistent		"In den Wäldern von Brasilien." Restricted by M. P. zu Wied-Neuwied in 1826 as "Flusse Mucuri [= Mucuri River]," Bahia, Brazil.			NA	NA				Mexico|Belize|Guatemala|Honduras|El Salvador|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil|Paraguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Macrophyllum_macrophyllum	0	sciname match	Macrophyllum_macrophyllum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Macrophyllum		macrophyllum	Schinz	1821	1	Das Thierreich	0.154861	Long-legged Bat	neuwiedii Gervais, 1855; nieuwiedii Gray, 1838.	Brazil, Bahia, Río Mucuri.	Tabasco (Mexico) south to Peru, Bolivia, SE Brazil, Paraguay, and NE Argentina.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/12615/22025883/' target='_blank'>Least Concern</a>	See Harrison (1975). See Emmons (1997) for distribution map.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Macrophyllum macrophyllum; Macrophyllum macrophyllum; Macrophyllum macrophyllum; Macrophyllum macrophyllum; Macrophyllum macrophyllum; Macrophyllum macrophyllum; neuwiedii; neuwiedii; nieuwiedii; macrophyllum; nieuwiedii; Macrophylle pécheur; Langbeinfledermaus; Macrofilode patas largas; Long-legged Bat; Long-legged Bat; Long-legged Bat; M. macrophyllum