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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L588	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Tonatia schulzi	Tonatia schulzi	Tonatia schulzi	Tonatia schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi	Lophostoma schulzi		[HMW] Tonatia schulzi Genoways & S.L. Williams, 1980 , “ 3 km SW Rudi Kappelvliegveld, 320 m , Brokopondo , Suriname .” A phylogenetic analysis of genetic data revalidated the genus Lophostoma for species that were previously in Tonatia , except TI: bidens and 1. saurophila , thus reassigning brasiliense , carriken , evotis , schulzi , and silvicola to Lophostoma . Monotypic.; [IUCN] Split from Tonatia .; [MDD2023] may be included under Tonatia based on chromosomal similarities, although further research is needed; [MDD2025_2.0] may be included under Tonatia based on chromosomal similarities, although further research is needed; [MDD2025_2.2] may be included under Tonatia based on chromosomal similarities, although further research is needed														schulzi	Split from Tonatia .			schulzi 	schulzi 			schulzi (Genoways & S. L. Williams, 1980)					Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Tonatia schulzi	Surinam, Brokopondo, 3 km S.W. Rudi Koppelvliegveld.	Genoways and Williams	1980	Ann. Carnegie Mus., 49:205.	Distribution: Guianas and extreme northern Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Guianas, N Brazil; ref. 4.46	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Genoways and Williams	1980	Ann. Carnegie Mus., 49:205.		Guianas, N Brazil.	Surinam, Brokopondo, 3 km SW Rudi Koppelvliegveld.		GENOWAYS & WILLIAMS	1980	Size fairly small (forearm length, 42-44 mm; condylobasal length, 18-20 mm). Ear pinna relatively long. Postorbital constriction and sagittal crest well developed. Numerous warts on dorsal surfaces of wing and leg bones. Underpartes grayish.	Distribution: Guianas and extreme northern Brazil.	No subspecies.		75	species	T. schulzi	GENOWAYS & WILLIAMS	1980	Tonatia	genus	Tonatia schulzi				Size fairly small (forearm length, 42-44 mm; condylobasal length, 18-20 mm). Ear pinna relatively long. Postorbital constriction and sagittal crest well developed. Numerous warts on dorsal surfaces of wing and leg bones. Underpartes grayish.	No subspecies.		3. T. schulzi GENOWAYS & WILLIAMS 1980.	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Phyllostominae		Lophostoma schulzi	Lophostoma		schulzi	Genoways and Williams	y	1980		Ann. Carnegie Mus.	49		205		Schulz's Round-eared Bat	Surinam, Brokopondo, 3 km SW Rudi Koppelvliegveld.	Guianas, N Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable as Tonatia schulzi.			03A687BCFFA5FFA5138DF9A0FAFCFEAA	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	504	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFA5FFA5138DF9A0FAFCFEAA.xml	Lophostoma schulzi	Phyllostomidae	Lophostoma	schulzi		1980	Lophostome de Schulz @fr | Schulz-Rundohrblattnase @de | Lofostoma de Schulz @es	Tonatia schulzi Genoways & S.L. Williams, 1980 , “ 3 km SW Rudi Kappelvliegveld, 320 m , Brokopondo , Suriname .” A phylogenetic analysis of genetic data revalidated the genus Lophostoma for species that were previously in Tonatia , except TI: bidens and 1. saurophila , thus reassigning brasiliense , carriken , evotis , schulzi , and silvicola to Lophostoma . Monotypic.	Endemic to the Guianas and N Brazil (Amazonas, Para, and Amapastates).	Head-body 63-69 mm, tail 12-13 mm, ear 27-29 mm, hindfoot 11-6 14 mm , forearm 42-45 mm; weight 15 g . Greater lengths of skulls are 22-8-24 mm. Schulz’s Round-eared Bat is medium-sized and only larger than the Pygmy Round-eared Bat ( L. brasiliense ). It can be distinguished by small wartlike granulations on dorsal surfaces of forearms, digits, calcars, hindlimbs, ears, and noseleaf (characteristics not found in any other phyllostomid). Dorsal hairs are pale brown, with pale bases, becoming almost white. It has post-auricular patches that are sparsely haired and paler than back. Venteris slightly paler than dorsum and can vary markedly among adults, ranging from medium gray to almost white, and younger adults have darker venters, suggesting that ventral pelage becomes paler with age. Ears are long, with single low band ofskin running from each ear and meeting in middle of forehead;tragus is attenuated but with distinctive prominent projection on inner margin. Calcar is longer than hindfoot. Schulz’s Round-eared Bat resembles other species of Lophostoma by having narrow I. It has been recorded that clitoris of a female Schulz’s Round-eared Bat is remarkably elongated and resembles a penis, unlike congeners that have intermediate stages of elongation.	Tropical lowland and lower montane rainforests ( Guiana highland regions) above elevations of 1500 m . Schulz’s Round-eared Bats are mainly recorded in primary forests, characterized by dense vegetation where understory often provides moderate groundcover, but they also inhabit regenerating secondary forests in Brazil .	Schulz’s Round-eared Bat is a generalist, gleaning insectivore. It might prefer large insects such as beetles and katydids.	A female taken in August in French Guiana had a 28 mm length (crown— rump) fetus.	Schulz’s Round-eared Batis nocturnal. Some behavior is expected to be similar to that of its congeners, so it might roost in large active arboreal hymenopteran (termites or ants) nests that are excavated and maintained.	No information.	Classified as Least Concern on The IUCN Red List.	Farneda et al. (2018) | Genoways & Williams (1980, 1984) | Gribel & Taddei (1989) | Lim & Tavares (2012) | Lim, Engstrom, Genowayset al. (2005) | Marques-Aguiar & Oren (1987) | Simmons & Voss (1998)	https://zenodo.org/record/6458666/files/figure.png	32. Schulz’s Round-eared Bat Lophostoma schulzi French: Lophostome de Schulz / German: Schulz-Rundohrblattnase / Spanish: Lofostoma de Schulz Taxonomy. Tonatia schulzi Genoways & S.L. Williams, 1980 , “ 3 km SW Rudi Kappelvliegveld, 320 m , Brokopondo , Suriname .” A phylogenetic analysis of genetic data revalidated the genus Lophostoma for species that were previously in Tonatia , except TI: bidens and 1. saurophila , thus reassigning brasiliense , carriken , evotis , schulzi , and silvicola to Lophostoma . Monotypic. Distribution. Endemic to the Guianas and N Brazil (Amazonas, Para, and Amapastates). Descriptive notes. Head-body 63-69 mm, tail 12-13 mm, ear 27-29 mm, hindfoot 11-6 14 mm , forearm 42-45 mm; weight 15 g . Greater lengths of skulls are 22-8-24 mm. Schulz’s Round-eared Bat is medium-sized and only larger than the Pygmy Round-eared Bat ( L. brasiliense ). It can be distinguished by small wartlike granulations on dorsal surfaces of forearms, digits, calcars, hindlimbs, ears, and noseleaf (characteristics not found in any other phyllostomid). Dorsal hairs are pale brown, with pale bases, becoming almost white. It has post-auricular patches that are sparsely haired and paler than back. Venteris slightly paler than dorsum and can vary markedly among adults, ranging from medium gray to almost white, and younger adults have darker venters, suggesting that ventral pelage becomes paler with age. Ears are long, with single low band ofskin running from each ear and meeting in middle of forehead;tragus is attenuated but with distinctive prominent projection on inner margin. Calcar is longer than hindfoot. Schulz’s Round-eared Bat resembles other species of Lophostoma by having narrow I. It has been recorded that clitoris of a female Schulz’s Round-eared Bat is remarkably elongated and resembles a penis, unlike congeners that have intermediate stages of elongation. Habitat. Tropical lowland and lower montane rainforests ( Guiana highland regions) above elevations of 1500 m . Schulz’s Round-eared Bats are mainly recorded in primary forests, characterized by dense vegetation where understory often provides moderate groundcover, but they also inhabit regenerating secondary forests in Brazil . Food and Feeding. Schulz’s Round-eared Bat is a generalist, gleaning insectivore. It might prefer large insects such as beetles and katydids. Breeding. A female taken in August in French Guiana had a 28 mm length (crown— rump) fetus. Activity patterns. Schulz’s Round-eared Batis nocturnal. Some behavior is expected to be similar to that of its congeners, so it might roost in large active arboreal hymenopteran (termites or ants) nests that are excavated and maintained. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Farneda et al. (2018), Genoways & Williams (1980, 1984), Gribel & Taddei (1989), Lim & Tavares (2012), Lim, Engstrom, Genowayset al. (2005), Marques-Aguiar & Oren (1987), Simmons & Voss (1998).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Lophostoma schulzi	Lophostoma		schulzi	Genoways & Williams	1980	1	Ann. Carnegie Mus.	52:25:00	Schulz's Round-eared Bat	None.	Surinam, Brokopondo, 3 km SW Rudi Koppelvliegveld.	Guianas, N Brazil.	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lophostoma schulzi	23	Schulz's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	PHYLLOSTOMINAE	PHYLLOSTOMINI	Lophostoma	NA	schulzi	Genoways & S. L. Williams	1980	1	Tonatia_schulzi	Genoways, H. H., & Williams, S. L. (1980). Results of the Alcoa Foundation-Suriname expeditions. I. A new species of bat of the genus Tonatia (Mammalia: Phyllostomatidae). Annals of the Carnegie Museum, 49, 205.	https://www.biodiversitylibrary.org/item/216130#page/237/mode/1up	CM 63687		"3 km SW Rudi Kappelvliegveld, 320 m, Brokopondo, Suriname."			schulzi (Genoways & S. L. Williams, 1980)	NA	NA	Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_schulzi	0	sciname match	Lophostoma_schulzi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	21987	Lophostoma schulzi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lophostoma	schulzi	Genoways &; Williams, 1980	Split from Tonatia .	20000000	Lophostoma schulzi	Least Concern		2016	2016-07-06 00:00:00 UTC	3.1	English	Listed as Least Concern because it is relatively widely distributed, however, is restricted to the Guyana Shield and known from very few records.	Feeding habits and roosting sites of L. schulzi are unknown. A poorly known gleaning insectivorous species associated with multistratal tropical forest. In Paracou, Simmons and Voss (1998) netted this species in well-drained primary forest, swampy primary forest, and creekside primary forest. Other specimens have been mistnetted above a trail in secondary vegetation at a site surrounded by mature tropical rain forest (Williams and Genoways 2008).	Deforestation is a threat to this species.	Very little information is available about the population status of this species although it is known to be rare within a restricted distribution in the Guyana Shield; therefore, their populations are expected to decline in response to habitat disturbance. Simmons and Voss (1998) caught seven individuals at Paracou, French Guiana, which suggests that the species could be not so rare locally.	Unknown	The species is known from the Guianas and northern Brazil (Simmon 2005, Williams and Genoways 2008).		Terrestrial	Retention of forests in Guyana Shield is needed.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Lophostoma		schulzi	Genoways & Williams	1980	1	Ann. Carnegie Mus.	52:25:00	Schulz's Round-eared Bat	None.	Surinam, Brokopondo, 3 km SW Rudi Koppelvliegveld.	Guianas, N Brazil.	Not listed.	Least Concern		Lophostoma schulzi	1004977	23	Schulz's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	PHYLLOSTOMINAE	PHYLLOSTOMINI	Lophostoma	NA	schulzi	Genoways & S. L. Williams	1980	1	Tonatia_schulzi	Genoways, H. H., & Williams, S. L. (1980). Results of the Alcoa Foundation-Suriname expeditions. I. A new species of bat of the genus Tonatia (Mammalia: Phyllostomatidae). Annals of the Carnegie Museum, 49, 205.	https://www.biodiversitylibrary.org/item/216130#page/237/mode/1up	CM 63687		"3 km SW Rudi Kappelvliegveld, 320 m, Brokopondo, Suriname."			schulzi (Genoways & S. L. Williams, 1980)	may be included under Tonatia based on chromosomal similarities, although further research is needed	da Silva, N. K. N., Nagamachi, C. Y., Rodrigues, L. R. R., O’Brien, P. C. M., Yang, F., Ferguson-Smith, M. A., & Pieczarka, J. C. (2022). Chromosome painting and phylogenetic analysis suggest that the genus Lophostoma (Chiroptera, Phyllostomidae) is paraphyletic. Scientific Reports, 12(1), 1-11.				Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_schulzi	0	sciname match	Lophostoma_schulzi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lophostoma_schulzi	1004977	23	Schulz's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Phyllostominae	Phyllostomini	Lophostoma	NA	schulzi	Genoways & S. L. Williams	1	Tonatia schulzi	Genoways, H.H. and Williams, S.L. 1980-07-07. Results of the Alcoa Foundation-Suriname Expeditions. I. A new species of bat of the genus _Tonatia_ (Mammalia: Phyllostomatidae). Annals of Carnegie Museum 49:203-211.	https://www.biodiversitylibrary.org/page/52334417	CM 63687	holotype		"3 km SW Rudi Kappelvliegveld, 320 m, Brokopondo, Suriname."	3.766667	-56.1667	may be included under Tonatia based on chromosomal similarities, although further research is needed	da Silva, N. K. N., Nagamachi, C. Y., Rodrigues, L. R. R., O’Brien, P. C. M., Yang, F., Ferguson-Smith, M. A., & Pieczarka, J. C. (2022). Chromosome painting and phylogenetic analysis suggest that the genus Lophostoma (Chiroptera, Phyllostomidae) is paraphyletic. Scientific Reports, 12(1), 1-11.				Guyana|Suriname|French Guiana|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_schulzi	0	sciname match	Lophostoma_schulzi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Lophostoma		schulzi	Genoways & Williams	1980	1	Ann. Carnegie Mus.	52:25:00	Schulz's Round-eared Bat	None.	Surinam, Brokopondo, 3 km SW Rudi Koppelvliegveld.	Guianas, N Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/21987/22041951/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lophostoma schulzi; Lophostoma schulzi; Lophostoma schulzi; Lophostoma schulzi; Lophostoma schulzi; Lophostoma schulzi; schulzi; Lophostome de Schulz; Schulz-Rundohrblattnase; Lofostoma de Schulz; Schulz's Round-eared Bat; Schulz's Round-eared Bat; Schulz's Round-eared Bat; L. schulzi