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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L583	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Tonatia carrikeri	Tonatia carrikeri	Tonatia carrikeri	Tonatia carrikeri	Tonatia carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri	Lophostoma carrikeri		[MSW3] See McCarthy et al. (1992).; [HMW] Chrotopterus carrikeri J. A. Allen, 1910 , “Rio Mocho,” Bolivar , Venezuela . Lophostoma carriker : was formerly in the genus ZTonatia, but T. E. Lee and collaborators in 2002 showed that Tonatiawas paraphyletic and recommended to restrict it to 1. bidens and 71. saurophila , while reclassifying remaining taxa to Lophostoma . Molecular and morphological analyses showed that L. yasuni , described from eastern Ecuador , was a junior synonym of L. carrikeri . Monotypic.; [batnames2022] Includes  yasuni ; see Camacho et al., 2016. See McCarthy et al. (1992).; [MDD2022] includes the recently described L. yasuni; [IUCN] The new molecular and morphological evidence places Lophostoma yasuni within the clade of L. carrikeri . L. yasuni should therefore be considered as a synonym of L. carrikeri (Camacho et al. 2016).; [batnames2023] Includes  yasuni ; see Camacho et al., 2016. See McCarthy et al. (1992).; [MDD2023] includes the recently described L. yasuni; [MDD2025_2.0] includes the recently described L. yasuni; [batnames2025_1.7] Includes  yasuni; see Camacho et al., 2016. See McCarthy et al. (1992).; [MDD2025_2.2] includes the recently described L. yasuni											yasuni	carrikeri 	carrikeri - yasuni	carrikeri , yasuni	The new molecular and morphological evidence places Lophostoma yasuni within the clade of L. carrikeri . L. yasuni should therefore be considered as a synonym of L. carrikeri (Camacho et al. 2016).	carrikeri 	carrikeri - yasuni	carrikeri, yasuni 	carrikeri, yasuni 	carrikeri	carrikeri - yasuni	carrikeri (J. A. Allen, 1910)|yasuni R. M. Fonseca & C. M. Pinto, 2004		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Venezuela, E Peru, Bolivia, Surinam	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Tonatia carrikeri	Venezuela, Bolivar, Rio Mocho.	J. A. Allen	1910	Bull. Am. Mus. Nat. Hist., 28:147.	Distribution: Confined to tropical South America east of the Andes from Colombia to Surinam and south to Bolivia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela, E Peru, Bolivia, Guianas	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	J. A. Allen	1910	Bull. Am. Mus. Nat. Hist., 28:147.		Colombia, Venezuela, Surinam, N Brazil, Bolivia, Peru.	l^nezuela, Bolivar, Rio Mocho.		J. A. ALLEN	1910	Size medium (forearm length, 43-50 mm; condylobasal length, 18-22 mm). Ear pinna relatively long. Postorbital constriction and sagittal crest well developed. No warts on wing and leg bones. Underparts white.	Distribution: Confined to tropical South America east of the Andes from Colombia to Surinam and south to Bolivia.	No subspecies.		75	species	T. carrikeri	J. A. ALLEN	1910	Tonatia	genus	Tonatia carrikeri				Size medium (forearm length, 43-50 mm; condylobasal length, 18-22 mm). Ear pinna relatively long. Postorbital constriction and sagittal crest well developed. No warts on wing and leg bones. Underparts white.	No subspecies.		4. T. carrikeri (J. A. ALLEN 1910).	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Phyllostominae		Lophostoma carrikeri	Lophostoma		carrikeri	J. A. Allen	y	1910		Bull. Am. Mus. Nat. Hist.	28		147		Carriker's Round-eared Bat	Venezuela, Bolivar, Río Mocho.	Colombia, Venezuela, Guianas, N Brazil, Bolivia, Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable as Tonata carrikeri.		See McCarthy et al. (1992).	03A687BCFFA2FFA516B9F643FB74F791	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	503	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFA2FFA516B9F643FB74F791.xml	Lophostoma carrikeri	Phyllostomidae	Lophostoma	carrikeri		1910	Lophostome a ventre blanc @fr | Carriker Rundohrblattnase @de | Lofostomade vientre blanco @es	Chrotopterus carrikeri J. A. Allen, 1910 , “Rio Mocho,” Bolivar , Venezuela . Lophostoma carriker : was formerly in the genus ZTonatia, but T. E. Lee and collaborators in 2002 showed that Tonatiawas paraphyletic and recommended to restrict it to 1. bidens and 71. saurophila , while reclassifying remaining taxa to Lophostoma . Molecular and morphological analyses showed that L. yasuni , described from eastern Ecuador , was a junior synonym of L. carrikeri . Monotypic.	E Colombia , S Venezuela , the Guianas, NE Ecuador , E Peru , N Brazil , and N Bolivia .	Head—body 66-76 mm, tail 10-14-9 mm, ear 24-25-9 mm, hindfoot 14-5-15 mm, forearm 42:2-47-7 mm; weight 18-25 g. Greatest lengths of skulls are 23-26-6 mm. Carriker’s Round-eared Bat is medium-sized and larger than the Pygmy Round-eared Bat ( L. brasiliense ) but smaller than Davis's Round-eared Bat ( L. evotis ), the Western Round-eared Bat (L. occidentale), and the White-throated Round-eared Bat ( L. silvicola ). Most measurements of Carriker’s Round-eared Bat overlap with those of Kalko’s Round-eared Bat ( IL. kalkoae ) and Schulz’s Round-eared Bats ( L. schulzi ). Carriker’s Round-eared Batis easily identified by plain white ventral fur on its throat, chest, and abdomen, which is bordered along flanks by gray-brown dorsal fur. Gular region is pale to whitish. Dorsal pelageis long and tricolored, with pale to whitish tips. Noseleaf, chin, and base of ears are dark brown. These characteristics are shared with Kalko’s Round-eared Bat, although the latter has dark brown gular fur and white to pale post-auricular patches. Skull is constricted postorbitally and slightly concave in orbital region; sagittal crests can vary from well developed in adult males to moderately developed or absent in females and young males. Females have an elongated clitoris and swollen labia.	Mesic and riparian forests, igapo, varzea, semideciduous savannas, hill forests, gallery forests, primary and secondary terra firma forests, and dry forests in the Amazon Basin. In Brazil , Carriker’s Round-eared Bat has been reported in a transitional locality between semideciduous forest and riparian forest. Generally, it seemsto prefer undisturbed forested areas.	Carriker’s Round-eared Bat is a gleaning insectivore. It prefers large beetles, crickets, cicadas, katydids, and other arthropods.	Reproductive pattern of Carriker’s Round-eared Bat is bimodal, with peaks in April-May and October, and reproductively active males have found in June. In Guyana , reproductively active males were found in March-April and a pregnant female and a lactating female in April.	Carriker’s Round-eared Bat is nocturnal. It prefers hollowed arboreal termite nets as shelters, living in small colonies. Observations of congeners suggest that only single adult males construct roost cavities and that they use particular and strong biting behaviors during excavation.	From observation of congeners,it can be inferred that social system Carriker’s Round-eared Bat is probably harem-based, with resource—defense polygyny where males offer roosts as a resource to females.	Classified as Least Concern on The IUCN Red List. Carriker’s Round-eared Bat seems to be locally rare but widespread.	Allen (1910) | Camacho, Chavez & Burneo (2016) | Camacho, Tirira et al. (2014) | Fonseca & Pinto (2004) | Lee et al. (2002) | Lim, B.K. et al. (1999) | McCarthy et al. (1992) | Zortéa et al. (2009)	https://zenodo.org/record/6458664/files/figure.png	31. Carriker’s Round-eared Bat Lophostoma carrikeri French: Lophostome a ventre blanc / German: CarrikerRundohrblattnase / Spanish: Lofostoma de vientre blanco Taxonomy. Chrotopterus carrikeri J. A. Allen, 1910 , “Rio Mocho,” Bolivar , Venezuela . Lophostoma carriker : was formerly in the genus ZTonatia, but T. E. Lee and collaborators in 2002 showed that Tonatiawas paraphyletic and recommended to restrict it to 1. bidens and 71. saurophila , while reclassifying remaining taxa to Lophostoma . Molecular and morphological analyses showed that L. yasuni , described from eastern Ecuador , was a junior synonym of L. carrikeri . Monotypic. Distribution. E Colombia , S Venezuela , the Guianas, NE Ecuador , E Peru , N Brazil , and N Bolivia . Descriptive notes. Head—body 66-76 mm, tail 10-14-9 mm, ear 24-25-9 mm, hindfoot 14-5-15 mm, forearm 42:2-47-7 mm; weight 18-25 g. Greatest lengths of skulls are 23-26-6 mm. Carriker’s Round-eared Bat is medium-sized and larger than the Pygmy Round-eared Bat ( L. brasiliense ) but smaller than Davis's Round-eared Bat ( L. evotis ), the Western Round-eared Bat (L. occidentale), and the White-throated Round-eared Bat ( L. silvicola ). Most measurements of Carriker’s Round-eared Bat overlap with those of Kalko’s Round-eared Bat ( IL. kalkoae ) and Schulz’s Round-eared Bats ( L. schulzi ). Carriker’s Round-eared Batis easily identified by plain white ventral fur on its throat, chest, and abdomen, which is bordered along flanks by gray-brown dorsal fur. Gular region is pale to whitish. Dorsal pelageis long and tricolored, with pale to whitish tips. Noseleaf, chin, and base of ears are dark brown. These characteristics are shared with Kalko’s Round-eared Bat, although the latter has dark brown gular fur and white to pale post-auricular patches. Skull is constricted postorbitally and slightly concave in orbital region; sagittal crests can vary from well developed in adult males to moderately developed or absent in females and young males. Females have an elongated clitoris and swollen labia. Habitat. Mesic and riparian forests, igapo, varzea, semideciduous savannas, hill forests, gallery forests, primary and secondary terra firma forests, and dry forests in the Amazon Basin. In Brazil , Carriker’s Round-eared Bat has been reported in a transitional locality between semideciduous forest and riparian forest. Generally, it seemsto prefer undisturbed forested areas. Food and Feeding. Carriker’s Round-eared Bat is a gleaning insectivore. It prefers large beetles, crickets, cicadas, katydids, and other arthropods. Breeding. Reproductive pattern of Carriker’s Round-eared Bat is bimodal, with peaks in April-May and October, and reproductively active males have found in June. In Guyana , reproductively active males were found in March-April and a pregnant female and a lactating female in April. Activity patterns. Carriker’s Round-eared Bat is nocturnal. It prefers hollowed arboreal termite nets as shelters, living in small colonies. Observations of congeners suggest that only single adult males construct roost cavities and that they use particular and strong biting behaviors during excavation. Movements, Home range and Social organization. From observation of congeners,it can be inferred that social system Carriker’s Round-eared Bat is probably harem-based, with resource—defense polygyny where males offer roosts as a resource to females. Status and Conservation. Classified as Least Concern on The IUCN Red List. Carriker’s Round-eared Bat seems to be locally rare but widespread. Bibliography. Allen (1910), Camacho, Chavez & Burneo (2016), Camacho, Tirira et al. (2014), Fonseca & Pinto (2004), Lee et al. (2002), Lim, B.K. et al. (1999), McCarthy et al. (1992), Zortéa et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Lophostoma carrikeri	Lophostoma		carrikeri	J. A. Allen	1910	1	Bull. Am. Mus. Nat. Hist.	30:27:00	Carriker's Round-eared Bat	 yasuni Fonseca and Pinto, 2004.	Venezuela, Bolivar, Río Mocho.	Colombia, Venezuela, Guianas, N&C Brazil, Bolivia, Ecuador, Peru.	Not listed.	Least Concern	Includes  yasuni ; see Camacho et al., 2016. See McCarthy et al. (1992).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lophostoma carrikeri	23	Carriker's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	PHYLLOSTOMINAE	PHYLLOSTOMINI	Lophostoma	NA	carrikeri	J. A. Allen	1910	1	Chrotopterus_carrikeri	Allen, J. A. (1910). Mammals from the Caura District of Venezuela, with description of a new species of Chrotopterus. Bulletin of the American Museum of Natural History, 28, 147.	https://digitallibrary.amnh.org/bitstream/handle/2246/1830//v2/dspace/ingest/pdfSource/bul/B028a12.pdf?sequence=1&isAllowed=y	AMNH 30181		"Rio Mocho," Bolívar, Venezuela.			carrikeri (J. A. Allen, 1910)|yasuni Fonseca & Pinto, 2004	includes the recently described L. yasuni	Fonseca, R. M., & Pinto, C. M. (2004). A New Lophostoma (Chiroptera: Phyllostomidae: Phyllostominae) from the Amazonia of Ecuador. Occasional Papers of the Museum of Texas Tech University, 242, 1-9.|Camacho Monge, M. A., & Burneo Nuñez, S. F. (2016). A taxonomic revision of the Yasuni Round-eared bat, Lophostoma yasuni (Chiroptera: Phyllostomidae). Zootaxa, 4114(3), 246-260.	Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_carrikeri	0	sciname match	Lophostoma_carrikeri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	100000000	Lophostoma carrikeri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lophostoma	carrikeri	(J.A. Allen, 1910)	The new molecular and morphological evidence places Lophostoma yasuni within the clade of L. carrikeri . L. yasuni should therefore be considered as a synonym of L. carrikeri (Camacho et al. 2016).	20000000	Lophostoma carrikeri	Least Concern		2016	2016-07-05 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	The species is associated with a variety of forested habitats, ranging from rainforests ;to savannah scrub forests and orchards, also associated with pristine forested habitats near streams, and secondary tropical rainforest with an understory mainly of immature woody and herbaceous vegetation (Williams and Genoways 2008, Camacho et al . 2014). ;It may invade llanos habitat via gallery forests. It uses hollow termite nests as roosting sites with roosting groups ranging from five to 12 individuals ;(Eisenberg and Redford 1999). It apparently feeds mainly on arthropods (McCarthy et al.  1992).	In Bolivia considered near threatened because known only from two localities. Loss of forest habitats although this is not a major threat.	Absent or rare to uncommon and occasionally locally abundant at sites, this is however a widespread species. Because of this rarity, ;the populations of which are expected to decline in response to habitat disturbance.	Decreasing	Thus far this species has been described in southeastern Colombia, eastern Ecuador, the extreme southern part of Amazonas territory in Venezuela, the Guianas extending to north-central Brazil, and adjacent to Peru and Bolivia (Simmons 2005, Williams and Genoways 2008, Camacho et al . 2014).		Terrestrial	Conservation of large patches of pristine forests habitats. The species occurs in a number of protected areas throughout its range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Lophostoma		carrikeri	J. A. Allen	1910	1	Bull. Am. Mus. Nat. Hist.	30:27:00	Carriker's Round-eared Bat	 yasuni Fonseca and Pinto, 2004.	Venezuela, Bolivar, Río Mocho.	Colombia, Venezuela, Guianas, N&C Brazil, Bolivia, Ecuador, Peru.	Not listed.	Least Concern	Includes  yasuni ; see Camacho et al., 2016. See McCarthy et al. (1992).	Lophostoma carrikeri	1004973	23	Carriker's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	PHYLLOSTOMINAE	PHYLLOSTOMINI	Lophostoma	NA	carrikeri	J. A. Allen	1910	1	Chrotopterus_carrikeri	Allen, J. A. (1910). Mammals from the Caura District of Venezuela, with description of a new species of Chrotopterus. Bulletin of the American Museum of Natural History, 28, 147.	https://digitallibrary.amnh.org/bitstream/handle/2246/1830//v2/dspace/ingest/pdfSource/bul/B028a12.pdf?sequence=1&isAllowed=y	AMNH 30181		"Rio Mocho," Bolívar, Venezuela.			carrikeri (J. A. Allen, 1910)|yasuni Fonseca & C. M. Pinto, 2004	includes the recently described L. yasuni	Fonseca, R. M., & Pinto, C. M. (2004). A New Lophostoma (Chiroptera: Phyllostomidae: Phyllostominae) from the Amazonia of Ecuador. Occasional Papers of the Museum of Texas Tech University, 242, 1-9.|Camacho Monge, M. A., & Burneo Nuñez, S. F. (2016). A taxonomic revision of the Yasuni Round-eared bat, Lophostoma yasuni (Chiroptera: Phyllostomidae). Zootaxa, 4114(3), 246-260.				Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_carrikeri	0	sciname match	Lophostoma_carrikeri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lophostoma_carrikeri	1004973	23	Carriker's Round-eared Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Phyllostominae	Phyllostomini	Lophostoma	NA	carrikeri	J. A. Allen	1	Chrotopterus carrikeri	Allen, J.A. 1910-05-27. Mammals from the Caura District of Venezuela, with description of a new species of _Chrotopterus_. Bulletin of the American Museum of Natural History 28(12):145-149.	https://hdl.handle.net/2246/1830	AMNH M-30181	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-30181	"Rio Mocho," Bolívar, Venezuela.			includes the recently described L. yasuni	Fonseca, R. M., & Pinto, C. M. (2004). A New Lophostoma (Chiroptera: Phyllostomidae: Phyllostominae) from the Amazonia of Ecuador. Occasional Papers of the Museum of Texas Tech University, 242, 1-9.|Camacho Monge, M. A., & Burneo Nuñez, S. F. (2016). A taxonomic revision of the Yasuni Round-eared bat, Lophostoma yasuni (Chiroptera: Phyllostomidae). Zootaxa, 4114(3), 246-260.				Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	LC	0	0	0	Lophostoma_carrikeri	0	sciname match	Lophostoma_carrikeri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Lophostoma		carrikeri	J. A. Allen	1910	1	Bull. Am. Mus. Nat. Hist.	30:27:00	Carriker's Round-eared Bat	yasuni Fonseca and Pinto, 2004.	Venezuela, Bolivar, Río Mocho.	Colombia, Venezuela, Guianas, N&C Brazil, Bolivia, Ecuador, Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/99783878/22041541/' target='_blank'>Least Concern</a>	Includes  yasuni; see Camacho et al., 2016. See McCarthy et al. (1992).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lophostoma carrikeri; Lophostoma carrikeri; Lophostoma carrikeri; Lophostoma carrikeri; Lophostoma carrikeri; Lophostoma carrikeri; yasuni; yasuni; carrikeri; yasuni; Lophostome a ventre blanc; Carriker Rundohrblattnase; Lofostomade vientre blanco; Carriker's Round-eared Bat; Carriker's Round-eared Bat; Carriker's Round-eared Bat; L. carrikeri