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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L569	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax	Lonchophylla mordax		[MSW2] Includes concava; see Handley (1966fl:763); but also see Jones and Carter (1976:16), who provisionally recognized it as a species.; [MSW3] Includes concava; see Handley (1966a); but also see Jones and Carter (1976), who provisionally recognized it as a distinct species.; [HMW] Lonchophylla mordax Thomas, 1903 , “Lamarao, NW. of Bahia [ Brazil ]. Alt. 300 m .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining data from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved the genus as paraphyletic and described Hsunycteris to include L. thomasi , L. cadenai , L. pattoni , and an unnamed species. As a result, Lonchophylla (sensu stricto) now comprises 13 Central American and South American species. Nevertheless, L. mordax , which is the type species of Lonchophylla , was not included in those assessments. Based on morphological similarity, C. O. Handley, Jr. in 1966 included L. concava as a subspecies with a disjunct population; it is now considered a distinct species. The current definition and therefore distribution ofthe Brazilian Nectar Bat have been affected by misidentifications with congeners L. dekeyseri and the recently described L. inexpectata . Monotypic.; [batnames2022] Does not include concava ; see Jones and Carter (1976).; [MDD2022] previously included L. concava; [IUCN] Moratelli and Dias (2015) described the species Lonchophylla inexpectata from the Caatinga of north-eastern Brazil, and noted that specimens of L. inexpectata have in the past been misidentified as L. mordax . Lonchophylla mordax occurs along the eastern border of the Caatinga and in the Atlantic Forest–Caatinga ecotone in North-eastern Brazil (Moratelli and Dias 2015).; [batnames2023] Does not include concava ; see Jones and Carter (1976).; [MDD2023] previously included L. concava; [MDD2025_2.0] previously included L. concava; [batnames2025_1.7] Does not include concava; see Jones and Carter (1976).; [MDD2025_2.2] previously included L. concava				concava		concava.	mordax, concava	mordax, concava						mordax	Moratelli and Dias (2015) described the species Lonchophylla inexpectata from the Caatinga of north-eastern Brazil, and noted that specimens of L. inexpectata have in the past been misidentified as L. mordax . Lonchophylla mordax occurs along the eastern border of the Caatinga and in the Atlantic Forest–Caatinga ecotone in North-eastern Brazil (Moratelli and Dias 2015).			mordax 	mordax 			mordax O. Thomas, 1903		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Ecuador, Bolivia, Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Lonchophylla mordax	Brazil, Bahia, Lamarao.	Thomas	1903	Ann. Mag. Nat. Hist., ser. 7, 12:459.	Distribution: Ranging from Costa Rica to Western Ecuador and also in eastern Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Brazilian long-tongued bat	Colombia, Ecuador, Bolivia, Brazil, ? Peru	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1903	Ann. Mag. Nat. Hist., ser. 7, 12:459.	Includes concava; see Handley (1966fl:763); but also see Jones and Carter (1976:16), who provisionally recognized it as a species.	Costa Rica to Ecuador; E Brazil; perhaps Peru and Bolivia.	Brazil, Bahia, Lamarao.		THOMAS	1903	Size fairly small (forearm length, 32-37 mm; condylobasal length, 20-23 mm; maxillary toothrow length, 7.4-8.3 mm). Lingual cusp of last upper premolar poorly developed or absent. Posterior cusp of anterior lower premolar absent or poorly developed. Posterior palatal emargination U-shaped.	Distribution: Ranging from Costa Rica to Western Ecuador and also in eastern Brazil.	Two subspecies are recognized, often treated as separate species:	L. m. mordax (eastern Brazil), L. m. concava (remainder of range).	78	species	L. mordax	THOMAS	1903	Lonchophylla	genus	Lonchophylla mordax				Size fairly small (forearm length, 32-37 mm; condylobasal length, 20-23 mm; maxillary toothrow length, 7.4- 8.3 mm). Lingual cusp of last upper premolar poorly developed or absent. Posterior cusp of anterior lower premolar absent or poorly developed. Posterior palatal emargination U-shaped.	Two subspecies are recognized, often treated as separate species:		3. L. mordax THOMAS 1903.	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Lonchophyllini	Lonchophylla mordax	Lonchophylla		mordax	Thomas		1903		Ann. Mag. Nat. Hist., ser. 7	12		459		Goldman's Nectar Bat	Brazil, Bahia, Lamarao.	Costa Rica south to Ecuador, Peru, and perhaps Bolivia; E Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	concava Goldman, 1914.	Includes concava; see Handley (1966a); but also see Jones and Carter (1976), who provisionally recognized it as a distinct species.	03A687BCFF9DFF9C16FCF885FF2BF156	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	528	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF9DFF9C16FCF885FF2BF156.xml	Lonchophylla mordax	Phyllostomidae	Lonchophylla	mordax	Thomas	1903	Lonchophylle du Brésil @fr | Brasilien-Nektarfledermaus @de | Loncéfilode Brasil @es | Goldman's Nectar Bat @en	Lonchophylla mordax Thomas, 1903 , “Lamarao, NW. of Bahia [ Brazil ]. Alt. 300 m .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining data from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved the genus as paraphyletic and described Hsunycteris to include L. thomasi , L. cadenai , L. pattoni , and an unnamed species. As a result, Lonchophylla (sensu stricto) now comprises 13 Central American and South American species. Nevertheless, L. mordax , which is the type species of Lonchophylla , was not included in those assessments. Based on morphological similarity, C. O. Handley, Jr. in 1966 included L. concava as a subspecies with a disjunct population; it is now considered a distinct species. The current definition and therefore distribution ofthe Brazilian Nectar Bat have been affected by misidentifications with congeners L. dekeyseri and the recently described L. inexpectata . Monotypic.	NE Brazil , at E border of semiarid Caatinga ecoregion (“sertao”) and Caatinga—Atlantic Forest ecotone (“agreste”) of Alagoas , Sergipe , and Bahia states. Previous records outside this area could be misidentifications.	Head-body 50-56 mm, tail 8-12 mm, ear 13-16 mm, hindfoot 8-10 mm, forearm 33-7-37-4 mm; weight 7-5-11 g. The Brazilian Nectar Bat is a medium-sized species of Lonchophylla . Dorsal fur is cinnamon-brown, with bases of hairs (c.80% of length) whitish; venter is pale to light brown, thus contrasting with dorsum. This distinction is useful to differentiate the Brazilian Nectar Bat from other species of Lonchophylla from eastern Brazil . Lower lip is marked by deep medial groove;its sides are bordered by small rounded papillae. Noseleaf is triangular, slightly higher than wide, more sharply toward its point. Forearms are almost naked. Wing membranes are light brown, and uropatagium is wide, completely enclosing short tail in its proximal one-third. Skull is long and narrow, slightly inflated at interorbital region. Lingual cusp is reduced or moderate in P°, and well-developed metastyles are distinct on upper molars. Dental formula for species of Lonchophyllais12/2,C1/1,PM 2/3, M 3/3 ( x2 ) = 34.	Eastern border ofsemiarid caatinga and agreste biomes in eastern Brazil at elevations up to 500 m .	The Brazilian Nectar Bat feeds on nectar, insects, and probably succulent fruits and pollen. Skull and tooth morphology indicate that it is primarily nectarivorous, with similar habits to other species of Lonchophylla .	No information.	Brazilian Nectar Bats probably roost in hollow trees and small caves.	Brazilian Nectar Bats probably roost in small groups.	Classified as Near Threatened on The IUCN Red List. The Brazilian Nectar Bat is restricted to very specific habitats in eastern Brazil , but it can be locally common at some locations.	Albuja & Gardner (2005) | Griffiths & Gardner (2008b) | Handley (1966b) | Moratelli & Dias (2015) | Nogueira, Dias & Peracchi (2007) | Parlos et al. (2014) | Taddei et al. (1983) | Thomas (1903c) | Woodman & Timm (2006)	https://zenodo.org/record/6458786/files/figure.png	88. Brazilian Nectar Bat Lonchophylla mordax French: Lonchophylle du Brésil / German: Brasilien-Nektarfledermaus / Spanish: Loncéfilo de Brasil Other common names: Goldman's Nectar Bat Taxonomy. Lonchophylla mordax Thomas, 1903 , “Lamarao, NW. of Bahia [ Brazil ]. Alt. 300 m .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining data from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved the genus as paraphyletic and described Hsunycteris to include L. thomasi , L. cadenai , L. pattoni , and an unnamed species. As a result, Lonchophylla (sensu stricto) now comprises 13 Central American and South American species. Nevertheless, L. mordax , which is the type species of Lonchophylla , was not included in those assessments. Based on morphological similarity, C. O. Handley, Jr. in 1966 included L. concava as a subspecies with a disjunct population; it is now considered a distinct species. The current definition and therefore distribution ofthe Brazilian Nectar Bat have been affected by misidentifications with congeners L. dekeyseri and the recently described L. inexpectata . Monotypic. Distribution. NE Brazil , at E border of semiarid Caatinga ecoregion (“sertao”) and Caatinga—Atlantic Forest ecotone (“agreste”) of Alagoas , Sergipe , and Bahia states. Previous records outside this area could be misidentifications. Descriptive notes. Head-body 50-56 mm, tail 8-12 mm, ear 13-16 mm, hindfoot 8-10 mm, forearm 33-7-37-4 mm; weight 7-5-11 g. The Brazilian Nectar Bat is a medium-sized species of Lonchophylla . Dorsal fur is cinnamon-brown, with bases of hairs (c.80% of length) whitish; venter is pale to light brown, thus contrasting with dorsum. This distinction is useful to differentiate the Brazilian Nectar Bat from other species of Lonchophylla from eastern Brazil . Lower lip is marked by deep medial groove;its sides are bordered by small rounded papillae. Noseleaf is triangular, slightly higher than wide, more sharply toward its point. Forearms are almost naked. Wing membranes are light brown, and uropatagium is wide, completely enclosing short tail in its proximal one-third. Skull is long and narrow, slightly inflated at interorbital region. Lingual cusp is reduced or moderate in P°, and well-developed metastyles are distinct on upper molars. Dental formula for species of Lonchophyllais12/2,C1/1,PM 2/3, M 3/3 ( x2 ) = 34. Habitat. Eastern border ofsemiarid caatinga and agreste biomes in eastern Brazil at elevations up to 500 m . Food and Feeding. The Brazilian Nectar Bat feeds on nectar, insects, and probably succulent fruits and pollen. Skull and tooth morphology indicate that it is primarily nectarivorous, with similar habits to other species of Lonchophylla . Breeding. No information. Activity patterns. Brazilian Nectar Bats probably roost in hollow trees and small caves. Movements, Home range and Social organization. Brazilian Nectar Bats probably roost in small groups. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Brazilian Nectar Bat is restricted to very specific habitats in eastern Brazil , but it can be locally common at some locations. Bibliography. Albuja & Gardner (2005), Griffiths & Gardner (2008b), Handley (1966b), Moratelli & Dias (2015), Nogueira, Dias & Peracchi (2007), Parlos et al. (2014), Taddei et al. (1983), Thomas (1903c), Woodman & Timm (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Lonchophylla mordax	Lonchophylla		mordax	Thomas	1903	0	Ann. Mag. Nat. Hist.	ser. 7, 12: 459	Mordant Nectar Bat	None.	Brazil, Bahia, Lamarao.	Costa Rica south to Ecuador, Peru, and perhaps Bolivia; E Brazil.		Near Threatened	Does not include concava ; see Jones and Carter (1976).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lonchophylla mordax	23	Brazilian Nectar Bat	Goldman's Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	LONCHOPHYLLINAE	LONCHOPHYLLINI	Lonchophylla	NA	mordax	O. Thomas	1903	0	Lonchophylla_mordax	Thomas, O. (1903). Notes on South-American monkeys, bats, carnivores, and rodents, with descriptions of new species. Annals and Magazine of Natural History, ser. 7, 12, 459.	https://www.biodiversitylibrary.org/item/94944#page/485/mode/1up	BM 1903.9.5.34		"Lamarão, N.W. of Bahia [Brazil]. Alt. 300 m."			mordax O. Thomas, 1903	previously included L. concava	Woodman, N. (2007). A new species of nectar-feeding bat, genus Lonchophylla, from western Colombia and western Ecuador (Mammalia: Chiroptera: Phyllostomidae). Proceedings of the Biological Society of Washington, 120(3), 340-358.	Brazil	South America	Neotropic	NT	0	0	0	Lonchophylla_mordax	0	sciname match	Lonchophylla_mordax	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	12267	Lonchophylla mordax	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lonchophylla	mordax	Thomas, 1903	Moratelli and Dias (2015) described the species Lonchophylla inexpectata from the Caatinga of north-eastern Brazil, and noted that specimens of L. inexpectata have in the past been misidentified as L. mordax . Lonchophylla mordax occurs along the eastern border of the Caatinga and in the Atlantic Forest–Caatinga ecotone in North-eastern Brazil (Moratelli and Dias 2015).	20000000	Lonchophylla mordax	Near Threatened		2016	2016-08-11 00:00:00 UTC	3.1	English	This species is listed as Near Threatened as its distribution has been restricted as a consequence of a taxonomic revision, and although it can be locally common it is also likely to be declining, especially in the eastern part of its range. This means that the extent of occurrence could be near the limit to be considered Vulnerable (criterion B1ab(iii)); however, the population as a whole could be declining at a rate (criterion A2) which would qualify it for inclusion in a threatened category.	Feeds on nectar, insects, and probably succulent fruits and pollen (Griffiths and Gardner, 2008). Probably roosts in small groups in hollow trees and in small caves. Based on the recent restriction of L. mordax (see Moratelli and Dias, 2015), the species occurs in the Atlantic Forest–Caatinga ecotone ;(agreste), and along the eastern border of the Caatinga (sertão). ;It is found in tall, humid forest. In the northeastern part of its range, it occupies very specific habitats - high plateaus that sustain humid forest in generally semi-arid regions.	The species specific habitats in the eastern part of the range (Caatinga region) are severely threatened.	There is limited information on its populations. However, it seems common throughout its range, being locally common.	Unknown	The species is endemic to southeastern Brazil (Simmons 2005), where it is widespread. However, the recent description of L. inexpectata by Moratelli and Dias (2015) restricts the distribution to just three localities from eastern Brazil, at Bahia and Sergipe states.		Terrestrial	The species-specific habitats, especially in the eastern part of the range, need to be protected. Because of the recent split of L. mordax by Moratelli and Dias (2015), a taxonomic assessment could be necessary.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Lonchophylla		mordax	Thomas	1903	0	Ann. Mag. Nat. Hist.	ser. 7, 12: 459	Mordant Nectar Bat	None.	Brazil, Bahia, Lamarao.	Costa Rica south to Ecuador, Peru, and perhaps Bolivia; E Brazil.		Near Threatened	Does not include concava ; see Jones and Carter (1976).	Lonchophylla mordax	1004937	23	Brazilian Nectar Bat	Goldman's Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	LONCHOPHYLLINAE	LONCHOPHYLLINI	Lonchophylla	NA	mordax	O. Thomas	1903	0	Lonchophylla_mordax	Thomas, O. (1903). Notes on South-American monkeys, bats, carnivores, and rodents, with descriptions of new species. Annals and Magazine of Natural History, ser. 7, 12, 459.	https://www.biodiversitylibrary.org/item/94944#page/485/mode/1up	BM 1903.9.5.34		"Lamarão, N.W. of Bahia [Brazil]. Alt. 300 m."			mordax O. Thomas, 1903	previously included L. concava	Woodman, N. (2007). A new species of nectar-feeding bat, genus Lonchophylla, from western Colombia and western Ecuador (Mammalia: Chiroptera: Phyllostomidae). Proceedings of the Biological Society of Washington, 120(3), 340-358.				Brazil	South America	Neotropic	NT	0	0	0	Lonchophylla_mordax	0	sciname match	Lonchophylla_mordax	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lonchophylla_mordax	1004937	23	Brazilian Nectar Bat	Goldman's Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Lonchophyllinae	Lonchophyllini	Lonchophylla	NA	mordax	O. Thomas	0	Lonchophylla mordax	Thomas, O. 1903-10-01. Notes on South-American monkeys, bats, carnivores, and rodents, with descriptions of new species. Annals and Magazine of Natural History (7)12(70):455-464.	https://www.biodiversitylibrary.org/page/29993221	BMNH:Mamm:1903.9.5.34	holotype	https://data.nhm.ac.uk/object/55780f09-61b7-42d1-94c7-1a7ee0c58e1f	"Lamarão, N.W. of Bahia [Brazil]. Alt. 300 m."			previously included L. concava	Woodman, N. (2007). A new species of nectar-feeding bat, genus Lonchophylla, from western Colombia and western Ecuador (Mammalia: Chiroptera: Phyllostomidae). Proceedings of the Biological Society of Washington, 120(3), 340-358.				Brazil	South America	Neotropic	NT	0	0	0	Lonchophylla_mordax	0	sciname match	Lonchophylla_mordax	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Lonchophylla		mordax	Thomas	1903	0	Ann. Mag. Nat. Hist.	ser. 7, 12: 459	Mordant Nectar Bat	None.	Brazil, Bahia, Lamarao.	Costa Rica south to Ecuador, Peru, and perhaps Bolivia; E Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/12267/22038521/' target='_blank'>Near Threatened</a>	Does not include concava; see Jones and Carter (1976).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lonchophylla mordax; Lonchophylla mordax; Lonchophylla mordax; Lonchophylla mordax; Lonchophylla mordax; Lonchophylla mordax; mordax; concava; mordax; Lonchophylle du Brésil; Brasilien-Nektarfledermaus; Loncéfilode Brasil; Goldman's Nectar Bat; Brazilian Nectar Bat; Goldman's Nectar Bat; Goldman's Nectar Bat; Mordant Nectar Bat; L. mordax