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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L561	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni	Lonchophylla bokermanni		[HMW] Lonchophylla bokermanni Sazima, Vizotto & Taddei, 1978 , “Serra do Cipo, municipio de Jaboticatubas, Estado de Minas Gerais , Brasil .” This species is monotypic.														bokermanni				bokermanni 	bokermanni 			bokermanni Sazima, Vizotto, & Taddei, 1978		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S Brazil; ref. 4.67	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Lonchophylla bokermanni	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.	Sazima et al.	1978	Rev. Brasil. Biol., 38:82.	Distribution: Known only from southeastern Brazil.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		SE Brazil	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Sazima et al.	1978	Rev. Brasil. Biol., 38:82.		SE Brazil.	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.		SAZIMA, VIZOTTO & TADDEI	1978	Size fairly large (forearm length, 38-42 mm; condylobasal length, 23-25 mm; maxillary toothrow length, 7.8-8.6 mm). Rostrum relatively narrow (width across upper molars, 5.2-5.7 mm). Lingual cusp of last upper premolar poorly developed. Posterior cusp of anterior lower premolar poorly developed.	Distribution: Known only from southeastern Brazil.	No subspecies.		78	species	L. bokermanni	SAZIMA, VIZOTTO & TADDEI	1978	Lonchophylla	genus	Lonchophylla bokermanni				Size fairly large (forearm length, 38-42 mm; condylobasal length, 23-25 mm; maxillary toothrow length, 7.8-8.6 mm). Rostrum relatively narrow (width across upper molars, 5.2-5.7 mm). Lingual cusp of last upper premolar poorly developed. Posterior cusp of anterior lower premolar poorly developed.	No subspecies.		6. L. bokermanni SAZIMA, VIZOTTO & TADDEI 1978.	6	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Lonchophyllini	Lonchophylla bokermanni	Lonchophylla		bokermanni	Sazima, Vizotto, and Taddei		1978		Rev. Brasil. Biol.	38		82		Bokermann's Nectar Bat	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.	SE Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.			03A687BCFF81FF811388FAA3F5D9F077	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	532	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF81FF811388FAA3F5D9F077.xml	Lonchophylla bokermanni	Phyllostomidae	Lonchophylla	bokermanni	Sazima, Vizotto & Taddei	1978	Lonchophylle de Bokermann @fr | Bokermann-Nektarfledermaus @de | Loncofilo de Bokermann @es	Lonchophylla bokermanni Sazima, Vizotto & Taddei, 1978 , “Serra do Cipo, municipio de Jaboticatubas, Estado de Minas Gerais , Brasil .” This species is monotypic.	NE & SE Brazil , cerrado biome of Bahia and Minas Geraisstates.	Head-body 60-63-5 mm, hindfoot 8-9 mm, forearm 38-7-41-3 mm; weight c. 12 g in the cerrado of Minas Gerais ; head-body 55-57, tail 10-12 mm, forearm 36-2-39-9 mm; weight 10-11 g in the caatinga of Bahia . No other specific measurements are available. Dorsal fur of Bokermann’s Nectar Bat ranges from yellowish brown to grayish brown. Hairs are bicolored, with light brown bases and dark brown tips. Underparts are lighter than dorsum, ranging from yellowish gray to grayish brown. Head is same color as dorsum. Deep medial sulcus occurs on chin, bordered by small warts. Well-developed bristles occur on upper and lower lip. Ears are round and small. Noseleaf is brown, with free flap at base of horseshoe. Wing membranes and uropatagium are dark brown. Uropatagium is long, reaching distal part oftibia. Calcar is smaller than foot but conspicuous. Tail is ¢.33% the length of uropatagium, with tip perforating uropatagium dorsally. Braincase is anteroposteriorly elongated, forming smooth slope between it and rostrum. Parietal and interparietal are slightly inflated. Rostrum is shorter than braincase. Basisphenoid pits are shallow. Zygomatic arches are incomplete. Palate is relatively short, broad, and V-shaped posteriorly. Mandibular condyle is placed above tooth row. I' are spatulated and project forward. Anterior surface of C' is grooved. P* is buccolingually narrow in occlusal view. Lower incisors are relatively broad and slightly trilobed. Chromosomal complement has 2n = 28 and FN = 50, with twelve pairs of metacentric or submetacentric and small pair of acrocentric autosomes. X-chromosome is metacentric, and Y-chromosome is acrocentric.	Rocky, montane fields (“campos rupestres”) at elevations of 720-1129 m. Vegetation is sparse and contains herbs, shrubs, grasses, and small trees.	Bokermann’s Nectar Bat is a nectarivore. In an altered campos rupestres in Minas Gerais , it consumed pollen and nectar of Bauhinia rufa ( Fabaceae ), and stomach contents contained fragments of ants ( Formicidae ). At another site in Minas Gerais , it consumed pollen and nectar of Encholirium glaziovi: ( Bromeliaceae ), a stemless bromeliad with terminal inflorescence 1-5-1-8 m tall. Bokermann’s Nectar Bat approached inflorescences c.30-60 minutes after dusk, circling around them but not visiting flowers. After these initial approaches, individuals begin to visit flowers. These bouts occurred at intervals of 5-40 minutes. When visiting inflorescences of E. glaziovii, individuals hovered in front of individual flowers and lapped nectar. They approached other flowers after short flights in loops. Two to six individuals perched on two small trees (2-3 m) located near flowers of E. glaziovii and remained in these nocturnal roosts for 5-30 minutes before restarting feeding bouts.	A pregnant Bokermann’s Nectar Bat was captured in January in Bahia , and reproductively inactive females were found in September—October in Minas Gerais .	In Bahia , Bokermann’s Nectar Bats left day roosts between 17:40 h and 22:00 h. In Minas Gerais , individuals began visiting flowers of E. glaziovii c.30 minutes after sunset. Bokermann’s Nectar Bat apparently is a cave-roosting species.	In Caetité ( Bahia ), a colony containing at least nine Bokermann’s Nectar Bats was found inside a mine, sharing it with Seba’s Short-tailed Bat ( Carollia perspicillata ) and Pallas’s Long-tongued Bat ( Glossophaga soricina ).	Classified as Endangered on The IUCN Red List. For 40 years, Bokermann’s Nectar Bat was known only from three localities in Minas Gerais . In 2018, it was recorded in the state of Bahia , extending its distribution 840 km to the north. Even with an estimated extent of occurrence of 17,534 km ?it is still poorly known and restricted to a specific montane habitat.	Almeida, B. et al. (2016) | Claudio, Silveira, Farias, Maas et al. (2018) | Dias et al. (2013) | Nascimento et al. (2013) | Sazima, Vizotto & Taddei (1978) | Sazima, Vogel & Sazima (1989) | Teixeira et al. (2015)	https://zenodo.org/record/6458808/files/figure.png	99. Bokermann’s Nectar Bat Lonchophylla bokermanni French: Lonchophylle de Bokermann / German: Bokermann-Nektarfledermaus / Spanish: Loncofilo de Bokermann Taxonomy. Lonchophylla bokermanni Sazima, Vizotto & Taddei, 1978 , “Serra do Cipo, municipio de Jaboticatubas, Estado de Minas Gerais , Brasil .” This species is monotypic. Distribution. NE & SE Brazil , cerrado biome of Bahia and Minas Geraisstates. Descriptive notes. Head-body 60-63-5 mm, hindfoot 8-9 mm, forearm 38-7-41-3 mm; weight c. 12 g in the cerrado of Minas Gerais ; head-body 55-57, tail 10-12 mm, forearm 36-2-39-9 mm; weight 10-11 g in the caatinga of Bahia . No other specific measurements are available. Dorsal fur of Bokermann’s Nectar Bat ranges from yellowish brown to grayish brown. Hairs are bicolored, with light brown bases and dark brown tips. Underparts are lighter than dorsum, ranging from yellowish gray to grayish brown. Head is same color as dorsum. Deep medial sulcus occurs on chin, bordered by small warts. Well-developed bristles occur on upper and lower lip. Ears are round and small. Noseleaf is brown, with free flap at base of horseshoe. Wing membranes and uropatagium are dark brown. Uropatagium is long, reaching distal part oftibia. Calcar is smaller than foot but conspicuous. Tail is ¢.33% the length of uropatagium, with tip perforating uropatagium dorsally. Braincase is anteroposteriorly elongated, forming smooth slope between it and rostrum. Parietal and interparietal are slightly inflated. Rostrum is shorter than braincase. Basisphenoid pits are shallow. Zygomatic arches are incomplete. Palate is relatively short, broad, and V-shaped posteriorly. Mandibular condyle is placed above tooth row. I' are spatulated and project forward. Anterior surface of C' is grooved. P* is buccolingually narrow in occlusal view. Lower incisors are relatively broad and slightly trilobed. Chromosomal complement has 2n = 28 and FN = 50, with twelve pairs of metacentric or submetacentric and small pair of acrocentric autosomes. X-chromosome is metacentric, and Y-chromosome is acrocentric. Habitat. Rocky, montane fields (“campos rupestres”) at elevations of 720-1129 m. Vegetation is sparse and contains herbs, shrubs, grasses, and small trees. Food and Feeding. Bokermann’s Nectar Bat is a nectarivore. In an altered campos rupestres in Minas Gerais , it consumed pollen and nectar of Bauhinia rufa ( Fabaceae ), and stomach contents contained fragments of ants ( Formicidae ). At another site in Minas Gerais , it consumed pollen and nectar of Encholirium glaziovi: ( Bromeliaceae ), a stemless bromeliad with terminal inflorescence 1-5-1-8 m tall. Bokermann’s Nectar Bat approached inflorescences c.30-60 minutes after dusk, circling around them but not visiting flowers. After these initial approaches, individuals begin to visit flowers. These bouts occurred at intervals of 5-40 minutes. When visiting inflorescences of E. glaziovii, individuals hovered in front of individual flowers and lapped nectar. They approached other flowers after short flights in loops. Two to six individuals perched on two small trees (2-3 m) located near flowers of E. glaziovii and remained in these nocturnal roosts for 5-30 minutes before restarting feeding bouts. Breeding. A pregnant Bokermann’s Nectar Bat was captured in January in Bahia , and reproductively inactive females were found in September—October in Minas Gerais . Activity patterns. In Bahia , Bokermann’s Nectar Bats left day roosts between 17:40 h and 22:00 h. In Minas Gerais , individuals began visiting flowers of E. glaziovii c.30 minutes after sunset. Bokermann’s Nectar Bat apparently is a cave-roosting species. Movements, Home range and Social organization. In Caetité ( Bahia ), a colony containing at least nine Bokermann’s Nectar Bats was found inside a mine, sharing it with Seba’s Short-tailed Bat ( Carollia perspicillata ) and Pallas’s Long-tongued Bat ( Glossophaga soricina ). Status and Conservation. Classified as Endangered on The IUCN Red List. For 40 years, Bokermann’s Nectar Bat was known only from three localities in Minas Gerais . In 2018, it was recorded in the state of Bahia , extending its distribution 840 km to the north. Even with an estimated extent of occurrence of 17,534 km ?it is still poorly known and restricted to a specific montane habitat. Bibliography. Almeida, B. et al. (2016), Claudio, Silveira, Farias, Maas et al. (2018), Dias et al. (2013), Nascimento et al. (2013), Sazima, Vizotto & Taddei (1978), Sazima, Vogel & Sazima (1989), Teixeira et al. (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Lonchophylla bokermanni	Lonchophylla		bokermanni	Sazima, Vizotto, & Taddei	1978	0	Rev. Brasil. Biol.	39:22:00	Bokermann's Nectar Bat	None.	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.	SE Brazil.	Not listed.	Endangered		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lonchophylla bokermanni	23	Bokermann's Nectar Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	LONCHOPHYLLINAE	LONCHOPHYLLINI	Lonchophylla	NA	bokermanni	Sazima, Vizotto, & Taddei	1978	0	Lonchophylla_bokermanni	Sazima, I., Vizotto, L. D., & Taddei, V. A. (1978). Uma nova espÃ©cies de Lonchophylla da Serra do CipÃ³, Minas Geraid, Brasil (Mammalia, Chiroptera, Phyllstomidae). Revista Brasileira de Biologia, 38, 82.	https://www.researchgate.net/publication/277776420_Uma_nova_especie_de_Lonchophylla_da_Serra_do_Cipo_Minas_Gerais_Brasil_Mammalia_Chiroptera_Phyllostomidae	DZSJRP 10347		"Serra do CipÃ³, municÃ­pio de Jaboticatubas, Estado de Minas Gerais, Brasil."			bokermanni Sazima, Vizotto, & Taddei, 1978	NA	NA	Brazil	South America	Neotropic	EN	0	0	0	Lonchophylla_bokermanni	0	sciname match	Lonchophylla_bokermanni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	12263	Lonchophylla bokermanni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lonchophylla	bokermanni	Sazima, Vizotto &; Taddei , 1978		20000000	Lonchophylla bokermanni	Endangered	B1ab(iii)	2016	2016-07-01 00:00:00 UTC	3.1	English	This species is listed as Endangered in virtue of a recent taxonomic assessment that restrict this species to a few localities at the Cerrado, a highly threatened region with an extent of occurrence lower than 5,000 kmÂ², and a continuing decline of habitat quality. The species habitat is severely threatened and fragmented.	<p><span lang="EN-US">Lonchophylla bokermanni <span lang="EN-US"> is a medium-sized nectar-feeding bat, and similarly, to its congeners may probably eat fruits and insects to complement its diet. ;The species is a nectarivore, it pollinates typical plants of the Cerrado, such as Bauhinia rufa (Sazima et al. 1978) and Encholirium glaziovii (Sazima et al. 1989). It is probably dependent of shelters associated with karstic regions. ;It roosts in small colonies of two to five individuals (Marinho-Filho pers. comm.).</span></p>	The geographic range and area of occurrence, the Cerrado region, ;are severely fragmented and the species is known to exist at no more than five localities. The fragmentation of habitat is continuous and there is a presumed decline in the extent and quality of L. bokermanni habitat ;in recent years.<p> ; </p>	<span lang="EN-US">Only a few individuals from the three localities, cited in the published papers are computed to the region of the species occurrence, indicating it must be rare in inventories, contributing with few individuals in the known samples to date. ;Since the description of this species in 1978, it has not been recorded in Minas Gerais (cerrado) despite surveys (Aguiar and Marinho-Filho pers. comm).	Unknown	<span lang="EN-US">Lonchophylla bokermanni <span lang="EN-US"> is known from only three localities: Jaboticatubas, ItambÃ© do Mato Dentro, and Diamantina. All these localities are in the Cerrado biome of eastern Brazil (Dias et al. 2013). <span lang="EN-US"><span lang="EN-US">Atlantic Forest populations formerly assigned to this species are now considered as representing a new species: L. peracchii .</span></span>		Terrestrial	<span lang="EN-US">There is an urgent need of an Action Plan for the Species similar to the one done for L. dekeyseri (see Aguiar et al . 2006). Additional surveys are required to confirm the species subpopulation distribution and genetic condition. Urgent education action is also needed in the area of the species occurrence. ;It is important that the species habitat is protected.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Lonchophylla		bokermanni	Sazima, Vizotto, & Taddei	1978	0	Rev. Brasil. Biol.	39:22:00	Bokermann's Nectar Bat	None.	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.	SE Brazil.	Not listed.	Endangered		Lonchophylla bokermanni	1004929	23	Bokermann's Nectar Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	LONCHOPHYLLINAE	LONCHOPHYLLINI	Lonchophylla	NA	bokermanni	Sazima, Vizotto, & Taddei	1978	0	Lonchophylla_bokermanni	Sazima, I., Vizotto, L. D., & Taddei, V. A. (1978). Uma nova espÃ©cies de Lonchophylla da Serra do CipÃ³, Minas Geraid, Brasil (Mammalia, Chiroptera, Phyllstomidae). Revista Brasileira de Biologia, 38, 82.	https://www.researchgate.net/publication/277776420_Uma_nova_especie_de_Lonchophylla_da_Serra_do_Cipo_Minas_Gerais_Brasil_Mammalia_Chiroptera_Phyllostomidae	DZSJRP 10347		"Serra do CipÃ³, municÃ­pio de Jaboticatubas, Estado de Minas Gerais, Brasil."			bokermanni Sazima, Vizotto, & Taddei, 1978	NA	NA				Brazil	South America	Neotropic	EN	0	0	0	Lonchophylla_bokermanni	0	sciname match	Lonchophylla_bokermanni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lonchophylla_bokermanni	1004929	23	Bokermann's Nectar Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Lonchophyllinae	Lonchophyllini	Lonchophylla	NA	bokermanni	Sazima, Vizotto, & Taddei	0	Lonchophylla bokermanni	Sazima, I., Vizotto, L.D. and Taddei, V.A. 1978. Uma nova espÃ©cie de _Lonchophylla_ da Serra do CipÃ³, Minas Gerais, Brasil (Mammalia, Chiroptera, Phyllostomidae). Revista Brasileira de Biologia 38(1):81-89.	https://www.researchgate.net/publication/277776420_Uma_nova_especie_de_Lonchophylla_da_Serra_do_Cipo_Minas_Gerais_Brasil_Mammalia_Chiroptera_Phyllostomidae	DZSJRP 10347	holotype		"Serra do CipÃ³, municÃ­pio de Jaboticatubas, Estado de Minas Gerais, Brasil."	-19.2667	-43.6	NA	NA				Brazil	South America	Neotropic	EN	0	0	0	Lonchophylla_bokermanni	0	sciname match	Lonchophylla_bokermanni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Lonchophylla		bokermanni	Sazima, Vizotto & Taddei	1978	0	Rev. Brasil. Biol.	39:22:00	Bokermann's Nectar Bat	None.	Brazil, Minas Gerais, Jaboticatubas, Serra do Cipo.	SE Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/12263/22038287/' target='_blank'>Endangered</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lonchophylla bokermanni; Lonchophylla bokermanni; Lonchophylla bokermanni; Lonchophylla bokermanni; Lonchophylla bokermanni; Lonchophylla bokermanni; bokermanni; Lonchophylle de Bokermann; Bokermann-Nektarfledermaus; Loncofilo de Bokermann; Bokermann's Nectar Bat; Bokermann's Nectar Bat; Bokermann's Nectar Bat; L. bokermanni
