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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L56	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens	Anoura latidens		[MSW3] Clearly distinct from geoffroyi; see Solari et al. (1999).; [HMW] Anoura latidens Handley, 1984 , “on Pico Avila, 2150 m , 5 km NNE Caracas (= “Hotel Humbolt [sic], 9.4 km N Caracas”), Distrito Federal, Venezuela .” Related A. carishina was recently synonymized with A. latidens . Monotypic.; [batnames2022] Includes carishina ; see Calderón-Acevedo et al. (2021). Clearly distinct from geoffroyi ; see Solari et al. (1999).; [MDD2022] includes the recently described carishina; [batnames2023] Includes carishina ; see Calderón-Acevedo et al. (2021). Clearly distinct from geoffroyi ; see Solari et al. (1999).; [MDD2023] includes the recently described carishina; [MDD2025_2.0] includes the recently described carishina; [batnames2025_1.7] Includes carishina; see Calderón-Acevedo et al. (2021).     Clearly distinct from geoffroyi; see Solari et al. (1999).; [MDD2025_2.2] includes the recently described carishina											carishina	latidens	latidens - carishina 	latidens, carishina		latidens	latidens - carishina 	latidens, carishina	latidens, carishina	latidens	latidens - carishina	latidens Handley, 1984|carishina Mantilla-Meluk & R. J. Baker, 2010						N/A					Distribution: Ranging from northern Venezuela to eastern Peru.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela – Peru; ref. 4.50	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Handley	1984	Proc. Biol. Soc. Washington, 97:503.		Venezuela, Colombia, Peru.	Venezuela, D.F., Pico Avila.		HANDLEY	1984	Size relatively large (forearm length, 40-46 mm; condylobasal length, 23-24 mm). Tail absent. Anterior lower premolar subequal to middle and posterior premolars. Upper canine not enlarged, its anterior face flat. Posterior upper premolar with a median internal cusp enclosed in the broad triangular base of the tooth, remaining premolars thick. Anterior upper molar with an anteroexternal cusp and crest.	Distribution: Ranging from northern Venezuela to eastern Peru.	No subspecies.		82	species	A. latidens	HANDLEY	1984	Anoura	genus	Anoura latidens				Size relatively large (forearm length, 40-46 mm; condylobasal length, 23-24 mm). Tail absent. Anterior lower premolar subequal to middle and posterior premolars. Upper canine not enlarged, its anterior face flat. Posterior upper premolar with a median internal cusp enclosed in the broad triangular base of the tooth, remaining premolars thick. Anterior upper molar with an anteroexternal cusp and crest.	No subspecies.		4. A. latidens HANDLEY 1984.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Glossophagini	Anoura latidens	Anoura		latidens	Handley		1984		Proc. Biol. Soc. Wash.	97		503		Broad-toothed Tailless Bat	Venezuela, Distrito Federal, Pico Avila.	Venezuela, Guyana, Colombia, Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Clearly distinct from geoffroyi; see Solari et al. (1999).	03A687BCFF94FF94134FF6A2F723F3F4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	521	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF94FF94134FF6A2F723F3F4.xml	Anoura latidens	Phyllostomidae	Anoura	latidens	Handley	1984	Anoura a dents larges @fr | Breitzahn-Langnasenfledermaus @de | Anoura de dientes largos @es	Anoura latidens Handley, 1984 , “on Pico Avila, 2150 m , 5 km NNE Caracas (= “Hotel Humbolt [sic], 9.4 km N Caracas”), Distrito Federal, Venezuela .” Related A. carishina was recently synonymized with A. latidens . Monotypic.	N & Andean Colombia , Venezuela , WC Guyana , and Andean Peru .	Head-body 59-77 mm (tailless), ear 11-17 mm, hindfoot 11-13 mm, forearm 40-46 mm; weight 15-16 g. The Broad-toothed Tailless Bat is among the largest species of Anoura , only slightly smaller than Geoffroy’s Tailless Bat ( A. geoffroyi ). Dorsal pelage is grayish brown, with lighter bases to hairs, and ventral pelage is paler, with whitish throat area. Uropatagium and calcars are greatly reduced and hairy on both surfaces. There is no tail. Muzzle is narrow and elongated. Lower incisors are absent, and zygomatic arches are always incomplete. Upper premolars are broad.	Primarily montane humid forest up to elevations of ¢. 2600 m but also tropical dry forests, yards, orchards, and arid thorn scrub of coastal forests.	The Broad-toothed Tailless Bat mainly eats nectar but also pollen. It has a high basal metabolic rate for a nectar-feeding bat, permitting it to maintain constant body temperature at high elevations based on a high carbohydrate diet.	A pregnant Broad-toothed Tailless Bat was collected in August, and nonreproductive females were collected in February and April.	No information.	No information.	Classified as Least Concern on The IUCN Red List. The Broadtoothed Tailless Bat seems locally common in its large distribution.	Calderén-Acevedo et al. (2019) | Griffiths & Gardner (2008a) | Handley (1984) | Lim & Engstrom (2001) | Mantilla-Meluk & Baker (2010) | Solari et al. (1999) | Soriano et al. (2002)	https://zenodo.org/record/6458740/files/figure.png	70. Broad-toothed Tailless Bat Anoura latidens French: Anoura a dents larges / German: Breitzahn-Langnasenfledermaus / Spanish: Anoura de dientes largos Taxonomy. Anoura latidens Handley, 1984 , “on Pico Avila, 2150 m , 5 km NNE Caracas (= “Hotel Humbolt [sic], 9.4 km N Caracas”), Distrito Federal, Venezuela .” Related A. carishina was recently synonymized with A. latidens . Monotypic. Distribution. N & Andean Colombia , Venezuela , WC Guyana , and Andean Peru . Descriptive notes. Head-body 59-77 mm (tailless), ear 11-17 mm, hindfoot 11-13 mm, forearm 40-46 mm; weight 15-16 g. The Broad-toothed Tailless Bat is among the largest species of Anoura , only slightly smaller than Geoffroy’s Tailless Bat ( A. geoffroyi ). Dorsal pelage is grayish brown, with lighter bases to hairs, and ventral pelage is paler, with whitish throat area. Uropatagium and calcars are greatly reduced and hairy on both surfaces. There is no tail. Muzzle is narrow and elongated. Lower incisors are absent, and zygomatic arches are always incomplete. Upper premolars are broad. Habitat. Primarily montane humid forest up to elevations of ¢. 2600 m but also tropical dry forests, yards, orchards, and arid thorn scrub of coastal forests. Food and Feeding. The Broad-toothed Tailless Bat mainly eats nectar but also pollen. It has a high basal metabolic rate for a nectar-feeding bat, permitting it to maintain constant body temperature at high elevations based on a high carbohydrate diet. Breeding. A pregnant Broad-toothed Tailless Bat was collected in August, and nonreproductive females were collected in February and April. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Broadtoothed Tailless Bat seems locally common in its large distribution. Bibliography. Calderén-Acevedo et al. (2019), Griffiths & Gardner (2008a), Handley (1984), Lim & Engstrom (2001), Mantilla-Meluk & Baker (2010), Solari et al. (1999), Soriano et al. (2002).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Anoura latidens	Anoura		latidens	Handley	1984	0	Proc. Biol. Soc. Wash.	97(3): 513	Broad-toothed Tailless Bat	 carishina Mantilla-Melu & Baker, 2010	Venezuela, Distrito Federal, Pico Avila.	Venezuela, Guyana, Colombia, Peru.	Not listed.	Least Concern	Includes carishina ; see Calderón-Acevedo et al. (2021). Clearly distinct from geoffroyi ; see Solari et al. (1999).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Anoura latidens	23	Broad-toothed Tailless Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	GLOSSOPHAGINAE	CHOERONYCTERINI	Anoura	NA	latidens	Handley	1984	0	Anoura_latidens	Handley, C. O., Jr. (1984). New Species of Mammals from Northern South America: A Longtongued Bat, Genus Anoura Gray. Proceedings of the Biological Society of Washington, 97, 513.	https://www.biodiversitylibrary.org/page/34642304#page/49/mode/1up	USNM 370119		"on Pico Avila, 2150 m, 5 km NNE Caracas (= "Hotel Humbolt [sic], 9.4 km N Caracas"), Distrito Federal, Venezuela."			latidens Handley, 1984|carishina Mantilla-Meluk & R. J. Baker, 2010	includes the recently described carishina	Mantilla-Meluk, H., & Baker, R. J. (2006). Systematics of small Anoura (Chiroptera: Phyllostomidae) from Colombia, with description of a new species. Museum of Texas Tech University, 261, 1-18.|Mantilla-Meluk, H., & Baker, R. J. (2010). New species of Anoura (Chiroptera: Phyllostomidae) from Colombia, with systematic remarks and notes on the distribution of the A. geoffroyi complex. Museum of Texas Tech University, 92, 1-19.|Calderón-Acevedo, C. A., Rodríguez-Posada, M. E., & Muchhala, N. (2021). Morphology and genetics concur that Anoura carishina is a synonym of Anoura latidens (Chiroptera, Glossophaginae). Mammalia, 85(5). https://doi.org/10.1515/mammalia-2020-0183	Colombia|Venezuela|Guyana|Peru	South America	Neotropic	LC	0	0	0	Anoura_latidens	0	sciname match	Anoura_latidens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	1568	Anoura latidens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Anoura	latidens	Handley, 1984		20000000	Anoura latidens	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern because of its wide distribution, presumed large population, occurrence in a number of protected areas and as it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	It is strongly associated with tropical evergreen forest (Handley 1976). It feeds on nectar, insects, fruit and pollen. It roosts in small groups in caves or tree hollows and is found in lowland rainforest, deciduous forest, gardens and plantations. It is common only where there are caves or rock crevices, and rare or perhaps absent from lowland Amazonian forests lacking high ground, rocks and caves (Emmons and Feer 1997).	There are no major threats but the habitat is under threat in some areas (Molinari pers. comm.).	In Venezuela and Colombia, it is not common (Linares 1998, Mantilla pers. comm.).	Stable	This species has a discontinuous range. It occurs in Venezuela, Guyana, Colombia and Peru (Simmons 2005). The elevation range is 50 to 2,100 m asl in Venezuela (Linares 1998).	This species is not used.	Terrestrial	It occurs in national parks throughout its range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Anoura		latidens	Handley	1984	0	Proc. Biol. Soc. Wash.	97(3): 513	Broad-toothed Tailless Bat	 carishina Mantilla-Melu & Baker, 2010	Venezuela, Distrito Federal, Pico Avila.	Venezuela, Guyana, Colombia, Peru.	Not listed.	Least Concern	Includes carishina ; see Calderón-Acevedo et al. (2021). Clearly distinct from geoffroyi ; see Solari et al. (1999).	Anoura latidens	1004896	23	Broad-toothed Tailless Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	GLOSSOPHAGINAE	CHOERONYCTERINI	Anoura	NA	latidens	Handley	1984	0	Anoura_latidens	Handley, C. O., Jr. (1984). New Species of Mammals from Northern South America: A Longtongued Bat, Genus Anoura Gray. Proceedings of the Biological Society of Washington, 97, 513.	https://www.biodiversitylibrary.org/page/34642304#page/49/mode/1up	USNM 370119		"on Pico Avila, 2150 m, 5 km NNE Caracas (= "Hotel Humbolt [sic], 9.4 km N Caracas"), Distrito Federal, Venezuela."			latidens Handley, 1984|carishina Mantilla-Meluk & R. J. Baker, 2010	includes the recently described carishina	Mantilla-Meluk, H., & Baker, R. J. (2006). Systematics of small Anoura (Chiroptera: Phyllostomidae) from Colombia, with description of a new species. Museum of Texas Tech University, 261, 1-18.|Mantilla-Meluk, H., & Baker, R. J. (2010). New species of Anoura (Chiroptera: Phyllostomidae) from Colombia, with systematic remarks and notes on the distribution of the A. geoffroyi complex. Museum of Texas Tech University, 92, 1-19.|Calderón-Acevedo, C. A., Rodríguez-Posada, M. E., & Muchhala, N. (2021). Morphology and genetics concur that Anoura carishina is a synonym of Anoura latidens (Chiroptera, Glossophaginae). Mammalia, 85(5). https://doi.org/10.1515/mammalia-2020-0183				Colombia|Venezuela|Guyana|Peru	South America	Neotropic	LC	0	0	0	Anoura_latidens	0	sciname match	Anoura_latidens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Anoura_latidens	1004896	23	Broad-toothed Tailless Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Glossophaginae	Choeronycterini	Anoura	Anoura	latidens	Handley	0	Anoura latidens	Handley, C.O., Jr. 1984-08-17. New species of mammals from northern South America: a long-tongued bat, genus _Anoura_ Gray. Proceedings of the Biological Society of Washington 97(3):513-521.	https://www.biodiversitylibrary.org/page/34642304	USNM:MAMM:370119	holotype	http://n2t.net/ark:/65665/3b4df6096-a876-4d3f-b5b9-75936fd47e27	"on Pico Avila, 2150 m, 5 km NNE Caracas (= "Hotel Humbolt [sic], 9.4 km N Caracas"), Distrito Federal, Venezuela."			includes the recently described carishina	Mantilla-Meluk, H., & Baker, R. J. (2006). Systematics of small Anoura (Chiroptera: Phyllostomidae) from Colombia, with description of a new species. Museum of Texas Tech University, 261, 1-18.|Mantilla-Meluk, H., & Baker, R. J. (2010). New species of Anoura (Chiroptera: Phyllostomidae) from Colombia, with systematic remarks and notes on the distribution of the A. geoffroyi complex. Museum of Texas Tech University, 92, 1-19.|Calderón-Acevedo, C. A., Rodríguez-Posada, M. E., & Muchhala, N. (2021). Morphology and genetics concur that Anoura carishina is a synonym of Anoura latidens (Chiroptera, Glossophaginae). Mammalia, 85(5). https://doi.org/10.1515/mammalia-2020-0183				Colombia|Venezuela|Guyana|Peru|Bolivia	South America	Neotropic	LC	0	0	0	Anoura_latidens	0	sciname match	Anoura_latidens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Anoura		latidens	Handley	1984	0	Proc. Biol. Soc. Wash.	97(3): 513	Broad-toothed Tailless Bat	carishina Mantilla-Melu & Baker, 2010	Venezuela, Distrito Federal, Pico Avila.	Venezuela, Guyana, Colombia, Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/1568/22106814/' target='_blank'>Least Concern</a>	Includes carishina; see Calderón-Acevedo et al. (2021).     Clearly distinct from geoffroyi; see Solari et al. (1999).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Anourina	Anoura latidens; Anoura latidens; Anoura latidens; carishina; latidens; carishina; Anoura a dents larges; Breitzahn-Langnasenfledermaus; Anoura de dientes largos; Broad-toothed Tailless Bat; Broad-toothed Tailless Bat; Broad-toothed Tailless Bat; Anoura carishina; Anoura latidens carishina; A. latidens