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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L549	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Lasiurus borealis [synonym of]	Lasiurus borealis varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius	Lasiurus varius		[MSW3] Subgenus Lasiurus, borealis species group. Often listed as synonym of borealis or blossevillii, but apparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).; [HMW] Nycticeius varius Poeppig, 1835 , “Antuco,” Chile . See L. egregius . Lasiurus varius was previously considered a subspecies of L. borealis or L. blossevilii . Additional studies based on morphological evidence treated it as a full species. Moreover, recent molecular analyses based on mtDNA and nDNA also recovered L. varius as distinct from other congeners and placed it in a basal position relative to other Red Bats. Monotypic.; [batnames2022] Subgenus Lasiurus , borealis species group. Often listed as synonym of borealis or blossevillii , butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).; [IUCN] Often listed as a synonym of borealis or blossevillii .; [batnames2023] Subgenus Lasiurus , borealis species group. Often listed as synonym of borealis or blossevillii , butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).; [batnames2025_1.7] Subgenus Lasiurus, borealis species group. Often listed as synonym of borealis or blossevillii, butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).									poeppigii			varius 	varius - poeppigii 	varius, poepingii	Often listed as a synonym of borealis or blossevillii .	varius 	varius - poeppigii 	varius, poepingii 	varius, poepingii, poeppigii	varius	varius - poeppigii	varius (Poeppig, 1835)|poepingii (Lesson, 1836)|poeppigii Simmons, 2005 [incorrect subsequent spelling | not used as valid]						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus varius	Lasiurus	Lasiurus	varius	Poeppig		1835		Reis. Chilie, Peru, und Amaz.	1		451		Cinnamon Red Bat	Chile, Antuco.	S Argentina, Chile.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).	poeppigii Lesson, 1836.	Subgenus Lasiurus, borealis species group. Often listed as synonym of borealis or blossevillii, but apparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).	4C3D87E8FF826A3DFF829AE519D7B3D3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	879	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF826A3DFF829AE519D7B3D3.xml	Lasiurus varius	Vespertilionidae	Lasiurus	varius		1835	Lasiure du Chili @fr | Zimtrote Haarschwanzfledermaus @de | Lasiurode Chile @es | Chilean Red Bat @en	Nycticeius varius Poeppig, 1835 , “Antuco,” Chile . See L. egregius . Lasiurus varius was previously considered a subspecies of L. borealis or L. blossevilii . Additional studies based on morphological evidence treated it as a full species. Moreover, recent molecular analyses based on mtDNA and nDNA also recovered L. varius as distinct from other congeners and placed it in a basal position relative to other Red Bats. Monotypic.	From C Chile and WC Argentina S to Tierra del Fuego .	Head-body 55-61 mm , tail 44-58 mm , ear 9-13- 9 mm , hindfoot 6-10 mm , forearm 3642- 1 mm ; weight 7-13 g . Dorsal hairs are tricolored, with black bases, yellowish middles, and bright red cinnamon tips. Ventral hairs are similar to dorsal hairs, but central band is much reduced. There are yellow patches on shoulders, and white wash on hairs is absent. Ears are small, rounded, black, and fringed with hairs on outer basal part. Wing membranes are black. Uropatagium is densely furred, with hairs extending beyond its edge. Skull is robust; rostrum is short and wide. Braincase is high and rounded; caudal spine of palate is well developed; sagittal crest is usually reduced, but it can be evident in anterior part of braincase; and lambdoid crests are slightly developed. I? is triangular and in contact with cingulum of C', which is long and pointed; P* is reduced and not visible in lateral view; M' and M? are well developed, and M® has only two commissures; I is tricuspid, and I,is bicuspid, filling gap between canines; and P, is small. Dental formulais1 1/3, C1/1,P2/2.	Temperate rainforests, xeric shrublands, Patagonian forests, deciduous forests, evergreen forests, pasturelands, pine and eucalypt plantations, and urban areas, usually associated with watercourses, at elevations up to ¢. 1000 m .	Cinnamon Red Bats usually forage alone, preying mainly on aquatic insects. They can forage above canopies but also beneath them and on trails. Diets mainly contain Homoptera, Coleoptera , Hymenoptera , and Diptera .	Pregnant Cinnamon Red Bats were captured in December. Females usually bear two embryos, but there are records of only one embryo.	The Cinnamon Red Bat roosts mainly in trees but was found on rocks along the coast of Chile . Echolocation calls are divided in three phases: search, approach, and final buzz. Search-phase calls are long (c.7 milliseconds) and include single-harmonic calls, with narrow downward FM component, followed by QCF component. They sweep from c¢.52 kHz to ¢.33 kHz, with interpulse intervals of ¢.216 milliseconds. Approach-phase calls are similar to search-phase calls but with shorter and broader bandwidth, higher slope, and shorter interpulse intervals. Terminal buzz consists of FM calls with no QCF elements, high pulse repetition rate, and lower call duration and interpulse intervals. Echolocation behavioris affected by degree of structural clutter of foraging areas.	Population sizes of Cinnamon Red Bats vary seasonally. It might be migratory at least in southern parts ofits distribution. It is mainly solitary, but clusters of four individuals have been observed.	Classified as Least Concern on The IUCN Red List. The Cinnamon Red Bat is widely distributed, presumably with large populations, and occurs in protected areas.	Baird et al. (2015) | Barquez et al. (1999) | Diaz et al. (2017) | Gardner & Handley (2008) | Mann (1978) | Mares et al. (1995) | Novaes, Garbino et al. (2018) | Rodriguez-San Pedro & Simonetti (2014) | Rodriguez-San Pedro etal. (2016) | Simmons (2005) | Solari (2018h)		260. Cinnamon Red Bat Lasiurus varius French: Lasiure du Chili / German: Zimtrote Haarschwanzfledermaus / Spanish: Lasiuro de Chile Other common names: Chilean Red Bat Taxonomy. Nycticeius varius Poeppig, 1835 , “Antuco,” Chile . See L. egregius . Lasiurus varius was previously considered a subspecies of L. borealis or L. blossevilii . Additional studies based on morphological evidence treated it as a full species. Moreover, recent molecular analyses based on mtDNA and nDNA also recovered L. varius as distinct from other congeners and placed it in a basal position relative to other Red Bats. Monotypic. Distribution. From C Chile and WC Argentina S to Tierra del Fuego . Descriptive notes. Head-body 55-61 mm , tail 44-58 mm , ear 9-13- 9 mm , hindfoot 6-10 mm , forearm 3642- 1 mm ; weight 7-13 g . Dorsal hairs are tricolored, with black bases, yellowish middles, and bright red cinnamon tips. Ventral hairs are similar to dorsal hairs, but central band is much reduced. There are yellow patches on shoulders, and white wash on hairs is absent. Ears are small, rounded, black, and fringed with hairs on outer basal part. Wing membranes are black. Uropatagium is densely furred, with hairs extending beyond its edge. Skull is robust; rostrum is short and wide. Braincase is high and rounded; caudal spine of palate is well developed; sagittal crest is usually reduced, but it can be evident in anterior part of braincase; and lambdoid crests are slightly developed. I? is triangular and in contact with cingulum of C', which is long and pointed; P* is reduced and not visible in lateral view; M' and M? are well developed, and M® has only two commissures; I is tricuspid, and I,is bicuspid, filling gap between canines; and P, is small. Dental formulais1 1/3, C1/1,P2/2.M 3/3 (2) = 32, Habitat. Temperate rainforests, xeric shrublands, Patagonian forests, deciduous forests, evergreen forests, pasturelands, pine and eucalypt plantations, and urban areas, usually associated with watercourses, at elevations up to ¢. 1000 m . Food and Feeding. Cinnamon Red Bats usually forage alone, preying mainly on aquatic insects. They can forage above canopies but also beneath them and on trails. Diets mainly contain Homoptera, Coleoptera , Hymenoptera , and Diptera . Breeding. Pregnant Cinnamon Red Bats were captured in December. Females usually bear two embryos, but there are records of only one embryo. Activity patterns. The Cinnamon Red Bat roosts mainly in trees but was found on rocks along the coast of Chile . Echolocation calls are divided in three phases: search, approach, and final buzz. Search-phase calls are long (c.7 milliseconds) and include single-harmonic calls, with narrow downward FM component, followed by QCF component. They sweep from c¢.52 kHz to ¢.33 kHz, with interpulse intervals of ¢.216 milliseconds. Approach-phase calls are similar to search-phase calls but with shorter and broader bandwidth, higher slope, and shorter interpulse intervals. Terminal buzz consists of FM calls with no QCF elements, high pulse repetition rate, and lower call duration and interpulse intervals. Echolocation behavioris affected by degree of structural clutter of foraging areas. Movements, Home range and Social organization. Population sizes of Cinnamon Red Bats vary seasonally. It might be migratory at least in southern parts ofits distribution. It is mainly solitary, but clusters of four individuals have been observed. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Cinnamon Red Bat is widely distributed, presumably with large populations, and occurs in protected areas. Bibliography. Baird et al. (2015), Barquez et al. (1999), Diaz et al. (2017), Gardner & Handley (2008), Mann (1978), Mares et al. (1995), Novaes, Garbino et al. (2018), Rodriguez-San Pedro & Simonetti (2014), Rodriguez-San Pedro etal. (2016), Simmons (2005), Solari (2018h).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Lasiurus varius	Lasiurus	Lasiurus	varius	Poeppig	1835	0	Reis. Chilie, Peru, und Amaz.	0.3549	Cinnamon Red Bat	 poeppigii Lesson, 1836.	Chile, Antuco.	S Argentina, Chile.	Not listed.	Least Concern	Subgenus Lasiurus , borealis species group. Often listed as synonym of borealis or blossevillii , butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lasiurus varius	23	Cinnamon Red Bat	Chilean Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	LASIURINI	Lasiurus	NA	varius	Poeppig	1835	1						"Antuco," Chile.			varius (Poeppig, 1835)|poepingii (Lesson, 1836)	NA	NA	Chile|Argentina	South America	Neotropic	LC	0	0	0	Lasiurus_varius	0	sciname match	Lasiurus_varius	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136690	Lasiurus varius	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Lasiurus	varius	Poeppig, 1835	Often listed as a synonym of borealis or blossevillii .	20000000	Lasiurus varius	Least Concern		2018	2018-03-05 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population, plus the fact that it occurs on protected areas, and no specific threats have been identified so far, making improbable to become threatened at the near future.	It is solitary and insectivore, as others vespertilionid bats (Rodríguez-San Pedro and Simonetti 2014). Usually associated to temperate rainforests of central and southern Chile and Argentina, but also found in plantations of pine or eucalyptus (G. Ossa pers. comm.). Also present in agricultural or modified landscapes, even urban areas, but depending on the presence of clear water courses and aquatic insects, which are the main part of its diet (Ossa 2010). It feed on insects, mostly Homoptera, Coleoptera, Hymenoptera and Diptera (Mann 1978).	No specific threat is known. However, the development of eolic turbines for production of energy could represent a significant threat in the future, as more of these are being set in southern South America.	Almost nothing is known about the populations of this species. Recent echolocation studies suggest it shows a solitary behavior while foraging (Rodríguez-San Pedro and Simonetti 2014). It could support good numbers on certain regions and at certain seasons (G. Ossa pers. comm.).	Unknown	This species is known from west central Argentina to Tierra del Fuego, and Central and southern Chile (Simmons 2005, Gardner and Handley 2008).		Terrestrial	Lost of forest cover on certain parts of its range should be closely monitored. More research is needed to confirm is this is a migratory species, also to know more about its ecology, reproduction, and threats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Lasiurus	Lasiurus	varius	Poeppig	1835	0	Reis. Chilie, Peru, und Amaz.	0.354861	Cinnamon Red Bat	 poeppigii Lesson, 1836.	Chile, Antuco.	S Argentina, Chile.	Not listed.	Least Concern	Subgenus Lasiurus , borealis species group. Often listed as synonym of borealis or blossevillii , butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).	Lasiurus varius	1005589	23	Cinnamon Red Bat	Chilean Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	LASIURINI	Lasiurus	Lasiurus	varius	Poeppig	1835	1						"Antuco," Chile.			varius (Poeppig, 1835)|poepingii (Lesson, 1836)	NA	NA				Chile|Argentina	South America	Neotropic	LC	0	0	0	Lasiurus_varius	0	sciname match	Lasiurus_varius	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lasiurus_varius	1005589	23	Cinnamon Red Bat	Chilean Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus	Lasiurus	varius	Poeppig	1	Nycticeius varius	Poeppig, E.F. 1835. Reise in Chile, Peru und auf dem Amazonenstrome, während der Jahre 1827–1832. Erster Band. Friedrich Fleischer, Leipzig, 466 pp.	https://www.digitale-sammlungen.de/de/view/bsb10366733?page=1				"Antuco," Chile.			NA	NA				Chile|Argentina	South America	Neotropic	LC	0	0	0	Lasiurus_varius	0	sciname match	Lasiurus_varius	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Lasiurus	Lasiurus	varius	Poeppig	1835	1	Reis. Chilie, Peru, und Amaz.	1: 451, footnote	Cinnamon Red Bat	poeppigii Lesson, 1836.	Chile, Antuco.	S Argentina, Chile.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136690/22040066/' target='_blank'>Least Concern</a>	Subgenus Lasiurus, borealis species group. Often listed as synonym of borealis or blossevillii, butapparently distinct; see Barquez (1987), Barquez et al. (1993), and Mares et al. (1995).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lasiurus varius; Lasiurus varius; Lasiurus varius; Lasiurus varius; Lasiurus varius; Lasiurus varius; poeppigii; poeppigii; varius; poepingii; Lasiure du Chili; Zimtrote Haarschwanzfledermaus; Lasiurode Chile; Chilean Red Bat; Cinnamon Red Bat; Chilean Red Bat; Cinnamon Red Bat; Cinnamon Red Bat; L. varius