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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L539	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus	Lasiurus ebenus		[MSW3] Subgenus Lasiurus, borealis species group. Known only from the holotype.; [HMW] Lasiurus ebenus Fazzolari-Corréa, 1994 , “ Brazil : Sao Paulo : Parque Estadual da ITha do Cardoso ( 25°05'S , 47°59'W ).” See L. egregius . Until recently, L. ebenus was known only from the holotype captured in 1991. Its taxonomic status has been questioned because distinction between L. ebenus and sympatric congener L. blossevilii was based mainly on color, which is highly variable. A second specimen captured in 2017 allowed qualitative and morphometric analyses and confirmed L. ebenus as a valid species. Lasiurus ebenus is distinguished from congeners by black membranes and pelage, bone-brown hair on interfemoral and ventral wing membranes, and presence of two upper premolars on each side, in which anterior is minute and medially displaced. Lasiurus ebenus is not yet phylogenetically positioned, but morphological traits indicate that it belongs in the Red Bat group. Monotypic.; [batnames2022] Subgenus Lasiurus , borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).; [batnames2023] Subgenus Lasiurus , borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).; [batnames2025_1.7] Subgenus Lasiurus, borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).														ebenus				ebenus 	ebenus 			ebenus Fazzolari-CorrÃªa, 1994						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus ebenus	Lasiurus	Lasiurus	ebenus	Fazzolari-Corrêa		1994		Mammalia	58		119		Blackish Red Bat	Brazil, São Paulo, Parque Estadual da Ilha do Cardoso, 25°05'S, 47°59'W.	SE Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Lasiurus, borealis species group. Known only from the holotype.	4C3D87E8FF816A3EFF4E9DFB19D0BD2E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	880	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF816A3EFF4E9DFB19D0BD2E.xml	Lasiurus ebenus	Vespertilionidae	Lasiurus	ebenus	Fazzolari-Correa	1994	Lasiure du Brésil @fr | Sao-Paulo-Haarschwanzfledermaus @de | Lasiurode Brasil @es | Blackish Red Bat @en	Lasiurus ebenus Fazzolari-Corréa, 1994 , “ Brazil : Sao Paulo : Parque Estadual da ITha do Cardoso ( 25°05'S , 47°59'W ).” See L. egregius . Until recently, L. ebenus was known only from the holotype captured in 1991. Its taxonomic status has been questioned because distinction between L. ebenus and sympatric congener L. blossevilii was based mainly on color, which is highly variable. A second specimen captured in 2017 allowed qualitative and morphometric analyses and confirmed L. ebenus as a valid species. Lasiurus ebenus is distinguished from congeners by black membranes and pelage, bone-brown hair on interfemoral and ventral wing membranes, and presence of two upper premolars on each side, in which anterior is minute and medially displaced. Lasiurus ebenus is not yet phylogenetically positioned, but morphological traits indicate that it belongs in the Red Bat group. Monotypic.	Known only from two localities in SE Atlantic Forest of Sao Paulo State , SE Brazil .	Head-body 57-462- 1 mm , tail 50-1-58- 1 mm , ear 12-4-15- 3 mm , hindfoot c. 11 mm , forearm 45-6— 45- 7 mm ; weight 12- 5-14 g . Dorsal hairs are tricolored, with black bases, followed by dark brown to bone-brown band, and black distal band. Ventral hairs are bicolored, with large bone-brown bases and black tips. Bone-brown to dark brown fur on uropatagium reaches one-third to one-halfof its length; ventral surface of humeri and forearms are also furred. Ears are broad and rounded, with triangular tragus measuring c.50% the ear length. Muzzle, borders of ears, and lips are black. Calcar is almost twice as long as hindfoot. Rostrum is short; braincase is broad; and sagittal crest is absent. P? is present, and P* is double rooted. Dental formulais11/3,C1/1,P 2/2, M 3/3 (x2) = 32.	Ombrophilous Forest and Premontane Forest areas in Atlantic Forest, associated to watercourses, at elevations of 40-630 m .	The Hairy-tailed Bat is insectivorous. Both specimens were captured over streams c. 1 m over the water surface.	No information.	One Hairy-tailed Bat was captured c.1 hour after sunset.	No information.	Classified as Data Deficient on The IUCNRed List. Main threat to the Hairy-tailed Bat is intensive habitat destruction in south-eastern region of Sao Paulo State . Both specimens were collected in protected areas.	Baird et al. (2015) | Claudio, Barbosa et al. (2018) | Fazzolari-Corréa (1994, 1995) | Gardner & Handley (2008) | Moratelli (2016) | Sampaio et al. (2016f)	https://zenodo.org/record/6398368/files/figure.png	263. Hairy-tailed Bat Lasiurus ebenus French: Lasiure du Brésil / German: Sao-Paulo-Haarschwanzfledermaus / Spanish: Lasiuro de Brasil Other common names: Blackish Red Bat Taxonomy. Lasiurus ebenus Fazzolari-Corréa, 1994 , “ Brazil : Sao Paulo : Parque Estadual da ITha do Cardoso ( 25°05'S , 47°59'W ).” See L. egregius . Until recently, L. ebenus was known only from the holotype captured in 1991. Its taxonomic status has been questioned because distinction between L. ebenus and sympatric congener L. blossevilii was based mainly on color, which is highly variable. A second specimen captured in 2017 allowed qualitative and morphometric analyses and confirmed L. ebenus as a valid species. Lasiurus ebenus is distinguished from congeners by black membranes and pelage, bone-brown hair on interfemoral and ventral wing membranes, and presence of two upper premolars on each side, in which anterior is minute and medially displaced. Lasiurus ebenus is not yet phylogenetically positioned, but morphological traits indicate that it belongs in the Red Bat group. Monotypic. Distribution. Known only from two localities in SE Atlantic Forest of Sao Paulo State , SE Brazil . Descriptive notes. Head-body 57-462- 1 mm , tail 50-1-58- 1 mm , ear 12-4-15- 3 mm , hindfoot c. 11 mm , forearm 45-6— 45- 7 mm ; weight 12- 5-14 g . Dorsal hairs are tricolored, with black bases, followed by dark brown to bone-brown band, and black distal band. Ventral hairs are bicolored, with large bone-brown bases and black tips. Bone-brown to dark brown fur on uropatagium reaches one-third to one-halfof its length; ventral surface of humeri and forearms are also furred. Ears are broad and rounded, with triangular tragus measuring c.50% the ear length. Muzzle, borders of ears, and lips are black. Calcar is almost twice as long as hindfoot. Rostrum is short; braincase is broad; and sagittal crest is absent. P? is present, and P* is double rooted. Dental formulais11/3,C1/1,P 2/2, M 3/3 (x2) = 32. Habitat. Ombrophilous Forest and Premontane Forest areas in Atlantic Forest, associated to watercourses, at elevations of 40-630 m . Food and Feeding. The Hairy-tailed Bat is insectivorous. Both specimens were captured over streams c. 1 m over the water surface. Breeding. No information. Activity patterns. One Hairy-tailed Bat was captured c.1 hour after sunset. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCNRed List. Main threat to the Hairy-tailed Bat is intensive habitat destruction in south-eastern region of Sao Paulo State . Both specimens were collected in protected areas. Bibliography. Baird et al. (2015), Claudio, Barbosa et al. (2018), Fazzolari-Corréa (1994, 1995), Gardner & Handley (2008), Moratelli (2016), Sampaio et al. (2016f).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Lasiurus ebenus	Lasiurus	Lasiurus	ebenus	Fazzolari-Corr&ecaron;a	1994	0	Mammalia	59:59:00	Blackish Red Bat	None.	Brazil, S&atilde;oPaulo, Parque Estadual da Ilha do Cardoso, 25Â°05'S, 47Â°59'W.	SE Brazil.	Not listed.	Data Deficient	Subgenus Lasiurus , borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lasiurus ebenus	23	Hairy-tailed Bat	Blackish Red Bat|Black Hairy-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	LASIURINI	Lasiurus	NA	ebenus	Fazzolari-CorrÃªa	1994	0	Lasiurus_ebenus	Fazzolari-CorrÃªa, S. (1994) Lasiurus ebenus, a new vespertilionid bat from southeastern Brazil. Mammalia, 58, 119.	https://www.degruyter.com/document/doi/10.1515/mamm.1994.58.1.119/html	MZUSP 28125		"Brazil: SÃ£o Paulo: Parque Estadual da Ilha do Cardoso (25Â°05'S, 47Â°59'W)."	-25.08	-47.98	ebenus Fazzolari-CorrÃªa, 1994	NA	NA	Brazil	South America	Neotropic	DD	0	0	0	Lasiurus_ebenus	0	sciname match	Lasiurus_ebenus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	11349	Lasiurus ebenus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Lasiurus	ebenus	Fazzolari-CorrÃªa, 1994		20000000	Lasiurus ebenus	Data Deficient		2016	2016-06-28 00:00:00 UTC	3.1	English	This species is listed as Data Deficient as it is known from a single specimen found in a protected area but within a region that is undergoing intense development.	This bat is poorly known. Probably its natural history is similar to other species of Lasiurus . As an aerial insectivore of uncluttered space, sampling methodology needs to be appropriate to record its activities.	The region of Sao Paulo is undergoing intensive human habitat destruction.	Nowak (1999) reports that Lasiurus ebenus is known only from a single specimen from the vicinity of Sao Paulo in southeastern Brazil, in a region that is undergoing intensive human habitat destruction. This species is extremely rare.	Unknown	Lasiurus ebenus is known only from type locality: Brazil, eastern Sao Paulo, Parque Estadual da Ihla do Cardoso (Eisenberg and Redford 1999). This locality belongs to the Atlantic forest of southeastern Brazil.		Terrestrial	The type locality is in Parque Estadual da Ilha do Cardoso. Further surveys are needed in the surrounding area, using appropriate methodology, to improve understanding of the species' distribution and ecology and to clarify its relationship to other species of Lasiurus .	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Lasiurus	Lasiurus	ebenus	Fazzolari-Corr&ecaron;a	1994	0	Mammalia	59:59:00	Blackish Red Bat	None.	Brazil, S&atilde;oPaulo, Parque Estadual da Ilha do Cardoso, 25Â°05'S, 47Â°59'W.	SE Brazil.	Not listed.	Data Deficient	Subgenus Lasiurus , borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).	Lasiurus ebenus	1005579	23	Hairy-tailed Bat	Blackish Red Bat|Black Hairy-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	LASIURINI	Lasiurus	Lasiurus	ebenus	Fazzolari-CorrÃªa	1994	0	Lasiurus_ebenus	Fazzolari-CorrÃªa, S. (1994) Lasiurus ebenus, a new vespertilionid bat from southeastern Brazil. Mammalia, 58, 119.	https://www.degruyter.com/document/doi/10.1515/mamm.1994.58.1.119/html	MZUSP 28125		"Brazil: SÃ£o Paulo: Parque Estadual da Ilha do Cardoso (25Â°05'S, 47Â°59'W)."	-25.0833	-47.9833	ebenus Fazzolari-CorrÃªa, 1994	NA	NA				Brazil	South America	Neotropic	DD	0	0	0	Lasiurus_ebenus	0	sciname match	Lasiurus_ebenus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lasiurus_ebenus	1005579	23	Hairy-tailed Bat	Blackish Red Bat|Black Hairy-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus	Lasiurus	ebenus	Fazzolari-CorrÃªa	0	Lasiurus ebenus	Fazzolari-CorrÃªa, S. 1994. _Lasiurus ebenus_, a new vespertilionid bat from southeastern Brasil. Mammalia 58(1):119-124.	https://doi.org/10.1515/mamm.1994.58.1.119	MZUSP 28125	holotype		"Brazil: SÃ£o Paulo: Parque Estadual da Ilha do Cardoso (25Â°05'S, 47Â°59'W)."	-25.0833	-47.9833	NA	NA				Brazil	South America	Neotropic	DD	0	0	0	Lasiurus_ebenus	0	sciname match	Lasiurus_ebenus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Lasiurus	Lasiurus	ebenus	Fazzolari-Corr&ecaron;a	1994	0	Mammalia	59:59:00	Blackish Red Bat	None.	Brazil, S&atilde;oPaulo, Parque Estadual da Ilha do Cardoso, 25Â°05'S, 47Â°59'W.	SE Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/11349/22119537/' target='_blank'>Data Deficient</a>	Subgenus Lasiurus, borealis species group. Known only from the holotype until 2018, when a second specimesn was collected; see Cl&aacute;udio et al. (2018).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lasiurus ebenus; Lasiurus ebenus; Lasiurus ebenus; Lasiurus ebenus; Lasiurus ebenus; Lasiurus ebenus; ebenus; Lasiure du Brésil; Sao-Paulo-Haarschwanzfledermaus; Lasiurode Brasil; Blackish Red Bat; Hairy-tailed Bat; Blackish Red Bat; Black Hairy-tailed Bat; Blackish Red Bat; Blackish Red Bat; L. ebenus
