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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L536	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus	Lasiurus castaneus		[MSW2] Subgenus Lasiurus.; [MSW3] Subgenus Lasiurus, borealis species group.; [HMW] Lasiurus castaneus Handley, 1960 , “Tacarcuna Village, 3,200 ft. [= 975 m ], Rio Pucro, Darién , Panama .” See L. egregius . Lasiurus castaneus and L. atratus might be conspecific. Lasiurus castaneus is in the Red Bats group. Monotypic.; [batnames2022] Subgenus Lasiurus , borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).; [batnames2023] Subgenus Lasiurus , borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).; [batnames2025_1.7] Subgenus Lasiurus, borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).														castaneus				castaneus 	castaneus 			castaneus Handley, 1960		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Panama; ref. 4.105	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Lasiurus castaneus	Panama, Darien, Rio Pucro, Tacarcuna Village, 3200 ft. (975 m).	Handley	1960	Proc. U.S. Nat. Mus., 112:468.	Distribution: From Costa Rica to Colombia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Tacarcuna bat	Panama	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Handley	1960	Proc. U.S. Natl. Mus., 112:468.	Subgenus Lasiurus.	Panama, Costa Rica.	Panama, Darien, Rio Pucro, Tacarcuna Village, 3,200 ft. (975 m).		HANDLEY	1960	Size medium (forearm length, 44-45 mm). Basicranial plane tilted upward from palate. Lacrimal tubercle absent. Small anterior upper premolar present.	Distribution: From Costa Rica to Colombia.	No subspecies.		129	species	L. castaneus	HANDLEY	1960	Lasiurus	subgenus	Lasiurus castaneus				Size medium (forearm length, 44-45 mm). Basicranial plane tilted upward from palate. Lacrimal tubercle absent. Small anterior upper premolar present.	No subspecies.		2. L. castaneus HANDLEY 1960.	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus castaneus	Lasiurus	Lasiurus	castaneus	Handley		1960		Proc. U.S. Natl. Mus.	112		468		Tacarcunan Bat	Panama, Darien, Río Pucro, Tacarcuna Village, 3,200 ft. (1,066 m).	Panama, Costa Rica. A record from French Guiana was subsequently reidentified as atratus (Handley, 1996).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Lasiurus, borealis species group.	4C3D87E8FF816A3EFF4E92B91D22B06B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	880	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF816A3EFF4E92B91D22B06B.xml	Lasiurus castaneus	Vespertilionidae	Lasiurus	castaneus	Handley	1960	Lasiure de Tacarcuna @fr | Panama-Haarschwanzfledermaus @de | Lasiuro de Tacarcuna @es | Red-faced Red Bat @en | Tacarcuna Bat @en	Lasiurus castaneus Handley, 1960 , “Tacarcuna Village, 3,200 ft. [= 975 m ], Rio Pucro, Darién , Panama .” See L. egregius . Lasiurus castaneus and L. atratus might be conspecific. Lasiurus castaneus is in the Red Bats group. Monotypic.	Known from scattered localifies in NW Costa Rica , E Panama , and N Brazil (Amazonas State).	Head-body 52-755- 6 mm , tail 47-50 mm , ear 11-417- 7 mm , hindfoot 7-5—- 11 mm , forearm 41-3-45- 5 mm ; weight 11- 5-17 g . Females are larger than males. Dorsum is deep chestnut, and rump and feet are mahogany. Dorsal hairs are tricolored, with black bases, amber middles, and chestnut tips. Ventral hairs are blackish brown, with scattered buff tips on collar. Ears are wide, short, and rounded, with basal two-thirds of outer part furred. Distal end of tragusis triangular. Auricle and antitragus are relatively large. Ears, membranes, and lips are blackish. Face and muzzle vary from black to reddish. Uropatagium is densely furred with mahogany hair that reaches three-quarters ofits length. Skull is short and broad; rostrum is broad and deep. Braincase is narrow, deep, and inclined upward away from plane of palate; lacrimal ridge is not developed; and palatal emargination is wider than depth. P* is double rooted; hypocone on M' and M? are much reduced; and M? is much reduced, with second commissure shorter than first.	Elfin forests, tall lowland evergreen forests, cloud forests, mixed secondary forests, pastures, and over streams and ponds in primary forests from sea level up to elevations of ¢. 1500 m .	The Tacarcunan Bat probably forages over watercourses and streams, capturing insects in flight. Specimens were captured c. 1 m over the water surface.	No information.	Tacarcunan Bats were captured in the first hour after sunset. Echolocation calls are c.6-3 milliseconds, with narrowband and single harmonic, and sweep from ¢.53 kHz to ¢.30 kHz.	No information.	Classified as Data Deficient on The IUCN Red List. The Tacarcunan Bat is rarely captured and occurs in protected areas in Panama .	Baird et al. (2015) | Collen (2012) | Dinerstein (1985) | Gardner & Handley (2008) | Handley (1960) | Morales-Martinez & Ramirez-Chaves (2015) | Novaes, Garbino et al. (2018) | Pineda et al. (2016) | Sampaio et al. (2003) | Simmons (2005) | Villalobos-Chaves & Dick (2014)	https://zenodo.org/record/6398366/files/figure.png	262. Tacarcunan Bat Lasiurus castaneus French: Lasiure de Tacarcuna / German: Panama-Haarschwanzfledermaus / Spanish: Lasiuro de Tacarcuna Other common names: Red-faced Red Bat , Tacarcuna Bat Taxonomy. Lasiurus castaneus Handley, 1960 , “Tacarcuna Village, 3,200 ft. [= 975 m ], Rio Pucro, Darién , Panama .” See L. egregius . Lasiurus castaneus and L. atratus might be conspecific. Lasiurus castaneus is in the Red Bats group. Monotypic. Distribution. Known from scattered localifies in NW Costa Rica , E Panama , and N Brazil (Amazonas State). Descriptive notes. Head-body 52-755- 6 mm , tail 47-50 mm , ear 11-417- 7 mm , hindfoot 7-5—- 11 mm , forearm 41-3-45- 5 mm ; weight 11- 5-17 g . Females are larger than males. Dorsum is deep chestnut, and rump and feet are mahogany. Dorsal hairs are tricolored, with black bases, amber middles, and chestnut tips. Ventral hairs are blackish brown, with scattered buff tips on collar. Ears are wide, short, and rounded, with basal two-thirds of outer part furred. Distal end of tragusis triangular. Auricle and antitragus are relatively large. Ears, membranes, and lips are blackish. Face and muzzle vary from black to reddish. Uropatagium is densely furred with mahogany hair that reaches three-quarters ofits length. Skull is short and broad; rostrum is broad and deep. Braincase is narrow, deep, and inclined upward away from plane of palate; lacrimal ridge is not developed; and palatal emargination is wider than depth. P* is double rooted; hypocone on M' and M? are much reduced; and M? is much reduced, with second commissure shorter than first. Dental formulas 11/3, C1/1,P 2/2, M 3/3 (x2) = 32, Habitat. Elfin forests, tall lowland evergreen forests, cloud forests, mixed secondary forests, pastures, and over streams and ponds in primary forests from sea level up to elevations of ¢. 1500 m . Food and Feeding. The Tacarcunan Bat probably forages over watercourses and streams, capturing insects in flight. Specimens were captured c. 1 m over the water surface. Breeding. No information. Activity patterns. Tacarcunan Bats were captured in the first hour after sunset. Echolocation calls are c.6-3 milliseconds, with narrowband and single harmonic, and sweep from ¢.53 kHz to ¢.30 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Tacarcunan Bat is rarely captured and occurs in protected areas in Panama . Bibliography. Baird et al. (2015), Collen (2012), Dinerstein (1985), Gardner & Handley (2008), Handley (1960), Morales-Martinez & Ramirez-Chaves (2015), Novaes, Garbino et al. (2018), Pineda et al. (2016), Sampaio et al. (2003), Simmons (2005), Villalobos-Chaves & Dick (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Lasiurus castaneus	Lasiurus	Lasiurus	castaneus	Handley	1960	0	Proc. U.S. Natl. Mus.	119:48:00	Tacarcunan Bat	None.	Panama, Darien, Río Pucro, Tacarcuna Village, 3,200 ft. (1,066 m).	Panama, Costa Rica, Brazil. A record from French Guiana was subsequently reidentified as atratus (Handley, 1996).	Not listed.	Data Deficient	Subgenus Lasiurus , borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Lasiurus castaneus	23	Tacarcunan Bat	Red-faced Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	LASIURINI	Lasiurus	NA	castaneus	Handley	1960	0	Lasiurus_castaneus	Handley, C. O., Jr. (1960). Decriptions of New Bats from Panama. Proceedings of the United States National Museum, 112, 468.	https://repository.si.edu/bitstream/handle/10088/16682/USNMP-112_3442_1960.pdf?sequence=1&isAllowed=y	USNM 310263		"Tacarcuna Village, 3,200 ft. [= 975 m], Río Pucro, Darién, Panama."			castaneus Handley, 1960	NA	NA	Costa Rica|Panama|Brazil	North America|South America	Nearctic|Neotropic	DD	0	0	0	Lasiurus_castaneus	0	sciname match	Lasiurus_castaneus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	11348	Lasiurus castaneus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Lasiurus	castaneus	Handley, 1960		20000000	Lasiurus castaneus	Data Deficient		2016	2016-06-30 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of recent information on its extent of occurrence, threats, status and ecological requirements.	The species is poorly known. It has been caught in mist nets set over streams through tall, lowland evergreen forests in Panama and in nets across elfin forests on an exposed ridge of the Continental Divide in Costa Rica (Reid 2009).	Threats for this species are not known.	There is no information available on the population status of this species.	Unknown	This species is restricted to Panama and Costa Rica (Dinerstein 1985, ;Reid 2009), with one record from the ;Central Amazonia of Brazil (Sampaio ;et al. ;2003). ;A record from French Guiana was subsequently reidentified as L. atratus (Handley 1996), and those from Colombia are not verified (Solari et al . 2013).		Terrestrial	The species is found in protected areas in Panama. Research actions are needed into the distribution, habitat, ecology, and threats to this species.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Lasiurus	Lasiurus	castaneus	Handley	1960	0	Proc. U.S. Natl. Mus.	119:48:00	Tacarcunan Bat	None.	Panama, Darien, Río Pucro, Tacarcuna Village, 3,200 ft. (1,066 m).	Panama, Costa Rica, Brazil. A record from French Guiana was subsequently reidentified as atratus (Handley, 1996).	Not listed.	Data Deficient	Subgenus Lasiurus , borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).	Lasiurus castaneus	1005576	23	Tacarcunan Bat	Red-faced Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	LASIURINI	Lasiurus	Lasiurus	castaneus	Handley	1960	0	Lasiurus_castaneus	Handley, C. O., Jr. (1960). Decriptions of New Bats from Panama. Proceedings of the United States National Museum, 112, 468.	https://repository.si.edu/bitstream/handle/10088/16682/USNMP-112_3442_1960.pdf?sequence=1&isAllowed=y	USNM 310263		"Tacarcuna Village, 3,200 ft. [= 975 m], Río Pucro, Darién, Panama."			castaneus Handley, 1960	NA	NA				Costa Rica|Panama|Brazil	North America|South America	Nearctic|Neotropic	DD	0	0	0	Lasiurus_castaneus	0	sciname match	Lasiurus_castaneus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Lasiurus_castaneus	1005576	23	Tacarcunan Bat	Red-faced Red Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Lasiurini	Lasiurus	Lasiurus	castaneus	Handley	0	Lasiurus castaneus	Handley, C.O., Jr. 1960-10-06. Descriptions of new bats from Panama. Proceedings of the United States National Museum 112:459-479.	https://www.biodiversitylibrary.org/page/7717340	USNM:MAMM:310263	holotype	http://n2t.net/ark:/65665/3fe12b6a6-6e8b-44c8-8f54-95ccfc2747fd	"Tacarcuna Village, 3,200 ft. [= 975 m], Río Pucro, Darién, Panama."			NA	NA				Costa Rica|Panama|Brazil|Bolivia	North America|South America	Neotropic	DD	0	0	0	Lasiurus_castaneus	0	sciname match	Lasiurus_castaneus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Lasiurus	Lasiurus	castaneus	Handley	1960	0	Proc. U.S. Natl. Mus.	119:48:00	Tacarcunan Bat	None.	Panama, Darien, Río Pucro, Tacarcuna Village, 3,200 ft. (1,066 m).	Panama, Costa Rica, Brazil. A record from French Guiana was subsequently reidentified as atratus (Handley, 1996).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/11348/22120924/' target='_blank'>Data Deficient</a>	Subgenus Lasiurus, borealis species group. The occurrence of this species in Brazil was first recorded by Sampaio et al.(2003), but they expressed some doubt about their identification of these specimens as belonging to L. castaneus. Their identifications were subsequently confirmed by Gardner and Handley (2008). Most likely occurs in Nicaragua, but specimens or published literature accounts appear lacking (H. York, pers. comm. 2/27/2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lasiurus castaneus; Lasiurus castaneus; Lasiurus castaneus; Lasiurus castaneus; Lasiurus castaneus; Lasiurus castaneus; castaneus; Lasiure de Tacarcuna; Panama-Haarschwanzfledermaus; Lasiuro de Tacarcuna; Red-faced Red Bat; Tacarcuna Bat; Tacarcunan Bat; Red-faced Red Bat; Tacarcunan Bat; Tacarcunan Bat; L. castaneus