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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L522	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Neoromicia somalica malagasyensis	Neoromicia malagasyensis	Laephotis malagasyensis	Laephotis malagasyensis	Neoromicia malagasyensis	Laephotis malagasyensis	Laephotis malagasyensis	Laephotis malagasyensis	Laephotis malagasyensis	Laephotis malagasyensis		[HMW] Eptesicus somalicus malagasyensis Peterson, Eger & Mitchell, 1995 , 220 km NE Tuléar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar ( 22°41' S , 45°13' E ). Originally described as a subspecies of N. somalica ; it is treated as a separate species based on cranial traits, and perhaps also bacular morphology. Its distinction from N. matroka is still open to question. Monotypic.; [batnames2022] Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus ; see Goodman and Ranivo (2004).; [MDD2022] split from Neoromicia somalica; moved from Neoromicia to Laephotis; [IUCN] The Madagascar populations of this species were included within Eptesicus (=Neoromicia ) somalicus malagasyensis (Petersen et al. 1995). However, this taxon was elevated to distinct species status (Goodman and Ranvio 2004), which is supported by the most recent taxonomic assessment (Bates et al . 2006, Goodman et al. 2015).; [batnames2023] Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus ; see Goodman and Ranivo (2004).; [MDD2023] split from Neoromicia somalica; moved from Neoromicia to Laephotis; [MDD2025_2.0] split from Neoromicia somalica; moved from Neoromicia to Laephotis; [batnames2025_1.7] Previously included in Neoromicia; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus; see Goodman and Ranivo (2004).; [MDD2025_2.2] split from Neoromicia somalica; moved from Neoromicia to Laephotis														malagasyensis	The Madagascar populations of this species were included within Eptesicus (=Neoromicia ) somalicus malagasyensis (Petersen et al. 1995). However, this taxon was elevated to distinct species status (Goodman and Ranvio 2004), which is supported by the most recent taxonomic assessment (Bates et al . 2006, Goodman et al. 2015).			malagasyensis	malagasyensis			malagasyensis (R. L. Peterson, Eger, & L. Mitchell, 1995)						N/A																																								NA																											4C3D87E8FFC66A7AFA859F131FB4B85D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	819	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFC66A7AFA859F131FB4B85D.xml	Neoromicia malagasyensis	Vespertilionidae	Neoromicia	malagasyensis		1995	Vespere de |'lsalo @fr | German @en | salo-Zwergfledermaus @en | Neoromicia de @es | salo @en	Eptesicus somalicus malagasyensis Peterson, Eger & Mitchell, 1995 , 220 km NE Tuléar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar ( 22°41' S , 45°13' E ). Originally described as a subspecies of N. somalica ; it is treated as a separate species based on cranial traits, and perhaps also bacular morphology. Its distinction from N. matroka is still open to question. Monotypic.	Madagascar , known only from four localities around the Isalo Massif.	Tail 30-4 mm, ear 9-8 mm, hindfoot 5: 3 mm (one specimen), forearm 30-1-32 mm. Dorsal pelage of the Isalo Serotine is dark brown, with mid-dorsal hairs c¢. 7 mm long; ventral pelage is sharply bicolored, with a mixture of dark buff and gray, becoming paler toward tail. Ears are mid-brown and translucent. Single available baculum is 2-2 mm long, has distal end flattened and deflected ventrally, as in the Malagasy Serotine ( N. matroka ), but has a smaller surface area, and lateral flanges and vertical projection are less developed; dental morphology of the two is similar. On following pages: 121. Roberts's Serotine ( Neoromicia robertsi ); 122. Stanley's Serotine ( Neoromicia stanleyi); 123. Zulu Serotine ( Neoromicia zuluensis ): 124. Tiny Serotine ( Neoromicia guineensis ); 125. Somali Serotine ( Neoromicia somalica ); 126. Banana Serotine ( Neoromicia nanus ); 127. Rendall’'s Serotine ( Neoromicia rendall): 128. Dark-brown Serotine ( Neoromicia brunnea ); 129. Rosevear's Serotine ( Neoromicia rosevear): 130. Isabelline Serotine ( Neoromicia isabella); 131. Grandidier’s Serotine ( Neoromicia grandidieri ); 132. White-winged Serotine ( Neoromicia tenuipinnis ); 133. Heller's Serotine ( Neoromicia helios ); 134. De Winton's Long-eared Bat ( Laephotis wintoni ); 135. Angolan Long-eared Bat ( Laephotis angolensis ); 136. Botswanan Long-eared Bat ( Laephotis botswanae ); 137. Namib Long-eared Bat ( Laephotis namibensis ): 138. Sind Bat ( Rhyneptesicus nasutus ); 139. Collared Sprite ( Thainycteris aureocollaris ); 140. Necklace Sprite ( Thainycteris torquatus ); 141. Bronze Sprite ( Arielulus circumdatus ); 142. Social Sprite ( Arielulus societatis ); 143. Coppery Sprite ( Arielulus cuprosus ).	The Isalo Serotine has been netted foraging over water in transitional gallery tropical dry forest in canyons, at elevations of 450-700 m.	No information.	In Isalo National Park, a male captured in December had slightly enlarged testes, and a female had large mammae, indicating recently ceased lactation.	Based on calls ofsix individuals, FM/QCF components are produced at low-duty cycle, with maximum energy in fundamental harmonic at ¢.45-8 kHz.	No information.	Classified as Vulnerable on The IUCN Red List. The Isalo Serotine is only known from a few specimens.	Bates et al. (2006) | Goodman & Ranivo (2004) | Kofoky et al. (2009) | Ramasindrazana et al. (2016) | Simmons (2005)	https://zenodo.org/record/6398025/files/figure.png	120. Isalo Serotine Neoromicia malagasyensis French: Vespere de |'lsalo / German: Isalo-Zwergfledermaus / Spanish: Neoromicia de Isalo Taxonomy. Eptesicus somalicus malagasyensis Peterson, Eger & Mitchell, 1995 , 220 km NE Tuléar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar ( 22°41' S , 45°13' E ). Originally described as a subspecies of N. somalica ; it is treated as a separate species based on cranial traits, and perhaps also bacular morphology. Its distinction from N. matroka is still open to question. Monotypic. Distribution. Madagascar , known only from four localities around the Isalo Massif. Descriptive notes. Tail 30-4 mm, ear 9-8 mm, hindfoot 5: 3 mm (one specimen), forearm 30-1-32 mm. Dorsal pelage of the Isalo Serotine is dark brown, with mid-dorsal hairs c¢. 7 mm long; ventral pelage is sharply bicolored, with a mixture of dark buff and gray, becoming paler toward tail. Ears are mid-brown and translucent. Single available baculum is 2-2 mm long, has distal end flattened and deflected ventrally, as in the Malagasy Serotine ( N. matroka ), but has a smaller surface area, and lateral flanges and vertical projection are less developed; dental morphology of the two is similar. On following pages: 121. Roberts's Serotine ( Neoromicia robertsi ); 122. Stanley's Serotine ( Neoromicia stanleyi); 123. Zulu Serotine ( Neoromicia zuluensis ): 124. Tiny Serotine ( Neoromicia guineensis ); 125. Somali Serotine ( Neoromicia somalica ); 126. Banana Serotine ( Neoromicia nanus ); 127. Rendall’'s Serotine ( Neoromicia rendall): 128. Dark-brown Serotine ( Neoromicia brunnea ); 129. Rosevear's Serotine ( Neoromicia rosevear): 130. Isabelline Serotine ( Neoromicia isabella); 131. Grandidier’s Serotine ( Neoromicia grandidieri ); 132. White-winged Serotine ( Neoromicia tenuipinnis ); 133. Heller's Serotine ( Neoromicia helios ); 134. De Winton's Long-eared Bat ( Laephotis wintoni ); 135. Angolan Long-eared Bat ( Laephotis angolensis ); 136. Botswanan Long-eared Bat ( Laephotis botswanae ); 137. Namib Long-eared Bat ( Laephotis namibensis ): 138. Sind Bat ( Rhyneptesicus nasutus ); 139. Collared Sprite ( Thainycteris aureocollaris ); 140. Necklace Sprite ( Thainycteris torquatus ); 141. Bronze Sprite ( Arielulus circumdatus ); 142. Social Sprite ( Arielulus societatis ); 143. Coppery Sprite ( Arielulus cuprosus ). Habitat. The Isalo Serotine has been netted foraging over water in transitional gallery tropical dry forest in canyons, at elevations of 450-700 m. Food and Feeding. No information. Breeding. In Isalo National Park, a male captured in December had slightly enlarged testes, and a female had large mammae, indicating recently ceased lactation. Activity patterns. Based on calls ofsix individuals, FM/QCF components are produced at low-duty cycle, with maximum energy in fundamental harmonic at ¢.45-8 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCN Red List. The Isalo Serotine is only known from a few specimens. It occurs in Isalo National Park, occupying a restricted area of 1905 km ² Bibliography. Bates et al. (2006), Goodman & Ranivo (2004), Kofoky et al. (2009), Ramasindrazana et al. (2016), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Laephotis malagasyensis	Laephotis		malagasyensis	Peterson, Eger & Mitchell	1995	1	Faune de Madagascar	85:40:00	Isalo Serotine	None.		C Madagascar.	Not listed.	Vulnerable as Neoromicia malagasyensis 	Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus ; see Goodman and Ranivo (2004).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Laephotis malagasyensis	23	Isalo Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Laephotis	NA	malagasyensis	R. L. Peterson, Eger, & L. Mitchell	1995	1	Eptesicus_somalicus_malagasyensis	Peterson, R. L., Eger, J. L. & Mitchell, L. (1995) Faune de Madagascar. 84. ChiroptÃ¨res. MusÃ©um national d'Histoire naturelle, Paris, 100.		ROM 42713		220 km NE TuleÌar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar (22Â°41' S, 45Â°13' E).	-22.68	45.217	malagasyensis (R. L. Peterson, Eger, & L. Mitchell, 1995)	split from Neoromicia somalica; moved from Neoromicia to Laephotis	Goodman, S. M., & Ranivo, J. (2004). The taxonomic status of Neoromicia somalicus malagasyensis. Mammalian Biology, 69(6), 434-438.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.	Madagascar	Africa	Afrotropic	VU	0	0	0	Neoromicia_malagasyensis	1	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136820	Neoromicia malagasyensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Neoromicia	malagasyensis	(Peterson, Eger &; Mitchell, 1995)	The Madagascar populations of this species were included within Eptesicus (=Neoromicia ) somalicus malagasyensis (Petersen et al. 1995). However, this taxon was elevated to distinct species status (Goodman and Ranvio 2004), which is supported by the most recent taxonomic assessment (Bates et al . 2006, Goodman et al. 2015).	20000000	Neoromicia malagasyensis	Vulnerable	B1ab(iii)+2ab(iii)	2019	2016-04-25 00:00:00 UTC	3.1	English	This species is listed as Vulnerable because it has an extent of occurrence (EOO) of only 7,301 kmÂ², and is currently known only from the area around the Isalo Massif (likely having a maximum area of occupancy (AOO) of &lt;2,000 kmÂ²), where it is presumed to be at risk of habitat loss.	This species has been found in transitional, gallery tropical dry forest in canyons where it has been netted foraging over water (Goodman and Ranivo 2004, Bates et al. 2006). Its roosting requirements have yet to be described.	The threats to this species are not known, but its habitat, particularly outside the Parc National dâ€™Isalo, is presumably undergoing some decline. It has yet to be recorded from synanthropic settings and it probably relies on natural roost sites in trees and other vegetation.	There is no information on the population status of this species.	Unknown	This species is endemic to Madagascar, where it is restricted to an area of 1,905 kmÂ² around the Isalo Massif (Bates et al. 2006, Goodman and Ramasindrazana 2013). It has been recorded between 450 and 700 m above sea level (Goodman and Ranivo 2004) and is currently known from four localities (Goodman and Ramasindrazana 2013, Goodman et al. 2015).		Terrestrial	This species is present in Parc National dâ€™Isalo. Future conservation measures should focus on roosting sites, but further research is needed to investigate the roosting and foraging habitats of this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Laephotis		malagasyensis	Peterson, Eger & Mitchell	1995	1	Faune de Madagascar	85:40:00	Isalo Serotine	None.		C Madagascar.	Not listed.	Vulnerable as Neoromicia malagasyensis 	Previously included in Neoromicia ; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus ; see Goodman and Ranivo (2004).	Laephotis malagasyensis	1005733	23	Isalo Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Laephotis	NA	malagasyensis	R. L. Peterson, Eger, & L. Mitchell	1995	1	Eptesicus_somalicus_malagasyensis	Peterson, R. L., Eger, J. L. & Mitchell, L. (1995) Faune de Madagascar. 84. ChiroptÃ¨res. MusÃ©um national d'Histoire naturelle, Paris, 100.		ROM 42713		220 km NE TuleÌar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar (22Â°41' S, 45Â°13' E).	-22.6833	45.21667	malagasyensis (R. L. Peterson, Eger, & L. Mitchell, 1995)	split from Neoromicia somalica; moved from Neoromicia to Laephotis	Goodman, S. M., & Ranivo, J. (2004). The taxonomic status of Neoromicia somalicus malagasyensis. Mammalian Biology, 69(6), 434-438.|Monadjem, A., Demos, T. C., Dalton, D. L., Webala, P. W., Musila, S., Kerbis Peterhans, J. C., & Patterson, B. D. (2020). A revision of the pipistrelle-like bats (Mammali: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society, zlaa087.				Madagascar	Africa	Afrotropic	VU	0	0	0	Neoromicia_malagasyensis	1	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Laephotis_malagasyensis	1005733	23	Isalo Serotine		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Laephotis	NA	malagasyensis	R. L. Peterson, Eger, & L. Mitchell	1	Eptesicus somalicus malagasyensis	Peterson, R.L., Eger, J.L. and Mitchell, L. 1995. ChiroptÃ¨res. Faune de Madagascar 84:1-204.		ROM:MAM:42713	holotype	http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-42713	220 km NE TuleÌar (= Toleara), Ilakaka, East of Sakaraha sous-prefecture, Madagascar (22Â°41' S, 45Â°13' E).	-22.7833	44.8	split from Neoromicia somalica; moved from Neoromicia to Laephotis	Goodman, S. M., & Ranivo, J. (2004). The taxonomic status of Neoromicia somalicus malagasyensis. Mammalian Biology, 69(6), 434-438.|Monadjem, A., T. C. Demos, D. L. Dalton, P. W. Webala, S. Musila, J. C. K. Peterhans and B. D. Patterson (2021). A revision of pipistrelle-like bats (Mammalia: Chiroptera: Vespertilionidae) in East Africa with the description of new genera and species. Zoological Journal of the Linnean Society 191(4): 1114â€“1146.				Madagascar	Africa	Afrotropic	VU (as Neoromicia malagasyensis)	0	0	0	Neoromicia_malagasyensis	1	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Laephotis		malagasyensis	Peterson, Eger & Mitchell	1995	1	Faune de Madagascar	85:40:00	Isalo Serotine	None.		C Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136820/22044073/' target='_blank'>Vulnerable as Neoromicia malagasyensis</a>	Previously included in Neoromicia; see Hoofer and Van Den Bussche (2003); Roehers et al. (2010) and Amador et al. (2016). Distinct from somalicus; see Goodman and Ranivo (2004).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Neoromicia malagasyensis; Laephotis malagasyensis; Laephotis malagasyensis; Neoromicia malagasyensis; Laephotis malagasyensis; malagasyensis; Vespere de; lsalo; German; salo-Zwergfledermaus; Neoromicia de; salo; Isalo Serotine; Isalo Serotine; L. malagasyensis
