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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L516	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi	Kerivoula whiteheadi		[MSW2] Subgenus Kerivoula.; [MSW3] Reviewed by Hill (1965) and Corbet and Hill (1992).; [HMW] Kerwvoula whiteheadi Thomas, 1894 , “ Isabella , N.E. Luzon,” Philippines . Phylogenetic position of K. whitehead: is uncertain because it has not been included in any genetic studies. Three subspecies recognized.; [batnames2022] Does not include pusilla or bicolor , which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).; [MDD2022] previously included K. pusilla (along with bicolor); [batnames2023] Does not include pusilla or bicolor , which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).; [MDD2023] previously included K. pusilla (along with bicolor); [MDD2025_2.0] previously included K. pusilla (along with bicolor); [batnames2025_1.7] Does not include pusilla or bicolor, which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).; [MDD2025_2.2] previously included K. pusilla (along with bicolor)				pusilia, bicolor		bicolor, pusilla (see Hill, 1965:532-533).	bicolor, pusilla, whiteheadi	whiteheadi, bicolor, pusilla		whiteheadi, bicolor, pusilla				whiteheadi				whiteheadi	whiteheadi			whiteheadi O. Thomas, 1894		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S Thailand, Borneo, Philippines	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula whiteheadi	Philippines, Luzon, Isabella.	Thomas	1894	Ann. Mag. Nat. Hist., ser. 6, 14:460.	Distribution: Known only from southern Thailand, Borneo, and the Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Thailand, Malaya, Borneo, Philippines	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1894	Ann. Mag. Nat. Hist., ser. 6, 14:460.	Subgenus Kerivoula.	Philippines, Borneo, S Thailand, W Malaysia.	Philippines, Luzon, Isabella.		THOMAS	1894	Rostrum relatively long, slender, and narrowed anteriorly. Anterior and middle upper and lower premolars narrow and elongate. Lingual margins of anterior and middle upper molars rounded with narrow shelves. Size relatively small (forearm length, 27-33 mm; condylobasal length, 11 -12 mm). Inner upper incisor bicuspid. Fringe of hair on posterior margin of uropatagium poorly developed.	Distribution: Known only from southern Thailand, Borneo, and the Philippines.	Three subspecies are currently recognized:	K. w. bicolor (southern Thailand), K. w. pusilla (Borneo, southern Philippines), K. w. whiteheadi (northern Philippines).	97	species	K. whiteheadi	THOMAS	1894	Kerivoula	subgenus	Kerivoula whiteheadi				Rostrum relatively long, slender, and narrowed anteriorly. An- terior and middle upper and lower premolars narrow and elongate. Lingual margins of anterior and middle upper molars rounded with narrow shelves. Size relatively small (forearm length, 27-33 mm; condylobasal length, 11 -12 mm). In- ner upper incisor bicuspid. Fringe of hair on posterior margin of uropatagium poorly developed.	Three subspecies are currently recognized:		1. K. whiteheadi THOMAS 1894.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula whiteheadi	Kerivoula		whiteheadi	Thomas		1894		Ann. Mag. Nat. Hist., ser. 6	14		460		Whitehead's Woolly Bat	Philippines, Luzon, Isabella, Molino.	Philippines, Borneo, S Thailand, W Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	bicolor Thomas, 1904; pusilla Thomas, 1894.	Reviewed by Hill (1965) and Corbet and Hill (1992).	4C3D87E8FF766AC9FF7394C61C07B3C9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	901	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF766AC9FF7394C61C07B3C9.xml	Kerivoula whiteheadi	Vespertilionidae	Kerivoula	whiteheadi	Thomas	1894	Kérivoule de Whitehead @fr | Whitehead-Wollfledermaus @de | Querivoula de Whitehead @es	Kerwvoula whiteheadi Thomas, 1894 , “ Isabella , N.E. Luzon,” Philippines . Phylogenetic position of K. whitehead: is uncertain because it has not been included in any genetic studies. Three subspecies recognized.	K.w. whitehead :Thomas,1894—Philippines(Luzon,Mindoro,Palawan,Panay,Cebu,Bohol,andMindanaoIs). K.w .bicolorThomas,1904—extremeSThailandandCPeninsularMalaysia. K. w. pusilla Thomas, 1894 — N Borneo.	Head-body c. 39-44 mm , tail 32-42 mm , ear 13-18 mm , hindfoot 7-9 mm , forearm 28-33 mm ; weight 3-5-5- 5 g . Fur is dense and woolly. Dorsal pelage is generally dark brown but can be reddish brown (typical on Mindanao); venteris paler (hairs with dark gray bases dorsally and ventrally). Wings of mainland individuals apparently have white tips; membranes and ears are dark. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavity just below tips on posterior borders; tragus is narrow and tall, with virtually straight anterior margin except for very slight convexity neartip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and calcaris long. Upper and lower premolars are elongated and oval in cross section, with sharp cusps, and I? is tall and bicuspid.	Secondary and disturbed forests, agricultural areas, various grasslands (including cogon grassland on Palawan Island), and montane forests ( Philippines ) from sea level up to elevations of ¢. 1490 m .	Whitehead’s Woolly Bats forage low to the ground and are slow fliers.	No information.	Whitehead’s Woolly Bats roost in foliage, including under large dead leaves.	Whitehead’s Woolly Bats roost in groups of 20-30 individuals in Sabah , Borneo.	Classified as Least Concern on The IUCN Red List. Whitehead’s Woolly Bat is found in various disturbed habitats, butit is fairly difficult to encounter. It does not seem to face any major threats.	Alviola (2000) | Esselstyn, Widmann & Heaney (2004) | Francis (2008a) | Francis, Rosell-Ambal & Tabaranza (2008) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Ingle et al. (1999) | Khan et al. (2008) | Kingston et al. (2006)	https://zenodo.org/record/6398472/files/figure.png	312. Whitehead’s Woolly Bat Kerivoula whiteheadi French: Kérivoule de Whitehead / German: Whitehead-Wollfledermaus / Spanish: Querivoula de Whitehead Taxonomy. Kerwvoula whiteheadi Thomas, 1894 , “ Isabella , N.E. Luzon,” Philippines . Phylogenetic position of K. whitehead: is uncertain because it has not been included in any genetic studies. Three subspecies recognized. Subspecies and Distribution. K.w.whitehead:Thomas,1894—Philippines(Luzon,Mindoro,Palawan,Panay,Cebu,Bohol,andMindanaoIs). K.w.bicolorThomas,1904—extremeSThailandandCPeninsularMalaysia. K. w. pusilla Thomas, 1894 — N Borneo. Descriptive notes. Head-body c. 39-44 mm , tail 32-42 mm , ear 13-18 mm , hindfoot 7-9 mm , forearm 28-33 mm ; weight 3-5-5- 5 g . Fur is dense and woolly. Dorsal pelage is generally dark brown but can be reddish brown (typical on Mindanao); venteris paler (hairs with dark gray bases dorsally and ventrally). Wings of mainland individuals apparently have white tips; membranes and ears are dark. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavity just below tips on posterior borders; tragus is narrow and tall, with virtually straight anterior margin except for very slight convexity neartip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and calcaris long. Upper and lower premolars are elongated and oval in cross section, with sharp cusps, and I? is tall and bicuspid. Habitat. Secondary and disturbed forests, agricultural areas, various grasslands (including cogon grassland on Palawan Island), and montane forests ( Philippines ) from sea level up to elevations of ¢. 1490 m . Food and Feeding. Whitehead’s Woolly Bats forage low to the ground and are slow fliers. Breeding. No information. Activity patterns. Whitehead’s Woolly Bats roost in foliage, including under large dead leaves. Movements, Home range and Social organization. Whitehead’s Woolly Bats roost in groups of 20-30 individuals in Sabah , Borneo. Status and Conservation. Classified as Least Concern on The IUCN Red List. Whitehead’s Woolly Bat is found in various disturbed habitats, butit is fairly difficult to encounter. It does not seem to face any major threats. Bibliography. Alviola (2000), Esselstyn, Widmann & Heaney (2004), Francis (2008a), Francis, Rosell-Ambal & Tabaranza (2008), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Ingle et al. (1999), Khan et al. (2008), Kingston et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula whiteheadi	Kerivoula		whiteheadi	Thomas	1894	0	Ann. Mag. Nat. Hist.	ser. 6, 14: 460	Whitehead's Woolly Bat	None.	Philippines, Luzon, Isabella, Molino.	Philippines, Borneo, S Thailand, W Malaysia.	Not listed.	Least Concern	Does not include pusilla or bicolor , which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula whiteheadi	23	Whitehead's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	whiteheadi	O. Thomas	1894	0	Kerivoula_Whiteheadi	Thomas, O. (1894). Descriptions of two new bats of the genus Kerivoula. Annals and Magazine of Natural History, ser. 6, 14, 460.	https://www.biodiversitylibrary.org/item/78508#page/494/mode/1up	BM 1894.10.9.2		"Isabella, N.E. Luzon," Philippines.			whiteheadi O. Thomas, 1894	previously included K. pusilla (along with bicolor)	Sedlock, J. L., Heaney, L. R., Balete, D. S., & Ruedi, M. (2020). Philippine bats of the genus Kerivoula (Chiroptera: Vespertilionidae): Overview and assessment of variation in K. pellucida and K. whiteheadi. Zootaxa, 4755(3), 454-490.	Philippines	Asia	Indomalaya	LC	0	0	0	Kerivoula_whiteheadi	0	sciname match	Kerivoula_whiteheadi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10987	Kerivoula whiteheadi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	whiteheadi	Thomas, 1894		20000000	Kerivoula whiteheadi	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern as this species is widespread and probably more common than first thought. There are no major threats at present, it occurs in a few protected areas and its populations are unlikely to be declining fast enough to qualify for listing in a more threatened category.	In the Philippines, the species has been found in disturbed forest and agricultural areas (Sanborn 1952). On Palawan island, it has been reported from cogon grassland at around 60 m asl (Esselstyn et al . 2004) and in secondary forest beside a creek at 450 m asl on Mt. Makiling (Laguna, Luzon) (Heaney et al . 1998, Sedlock 2001). Surveys found the species to be common in old growth forests near streams (P. Alviola pers comm. 2018).	There are no major threats to this species throughout its range. In the Philippines, the species is locally threatened at higher elevations in montane forest on ultramafic soils through activities associated with mining (D. Balete pers. comm. 2006).	Originally this species was considered to be extremely rare but this was probably due to inappropriate survey methodology and an inadequate number of surveys. Surveys using harp traps in montane forest on mount Isarog (Camarines Sur Province, Luzon) have found the species to be common at high elevations (1,400 m asl) (J. Sedlock pers. comm. 2006). It is known only from isolated records in Malaysia, and it is not certain whether this reflects genuine rarity for this species or if it is a sign of the sampling difficulties associated with it.	Unknown	This species occurs in southern Thailand, Borneo, and the Philippines. In Malaysian Borneo, the species is known only from two localities in Sarawak (Mulu and Usun Apau) and from Sabah (Kinabatangan). In the Philippines, records are from Bohol (FMNH), Cebu (FMNH), Luzon (Aurora (FMNH), Bulacan (Duya), Cagayan (FMNH), Camarines Norte (KU), Camarines Sur (FMNH), Isabela (Duya, BMNH), Laguna (FMNH), Nueva Vizcaya (FMNH), Quezon (FMNH), and Rizal (FMNH) provinces), Mindanao (Davao del Norte, Davao del Sur, Davao Oriental, Surigao del Sur (FMNH), and Zamboanga del Norte Provinces), Mindoro (Duya, KU), Palawan, Panay (Heaney et al. 1998, Taylor 1934) Polillo (Alviola 1999), and Sibuyan (Alvarez).		Terrestrial	It occurs in protected areas throughout its range. Taxonomic research is needed to clarify the status of some populations.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		whiteheadi	Thomas	1894	0	Ann. Mag. Nat. Hist.	ser. 6, 14: 460	Whitehead's Woolly Bat	None.	Philippines, Luzon, Isabella, Molino.	Philippines, Borneo, S Thailand, W Malaysia.	Not listed.	Least Concern	Does not include pusilla or bicolor , which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).	Kerivoula whiteheadi	1005309	23	Whitehead's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	whiteheadi	O. Thomas	1894	0	Kerivoula_Whiteheadi	Thomas, O. (1894). Descriptions of two new bats of the genus Kerivoula. Annals and Magazine of Natural History, ser. 6, 14, 460.	https://www.biodiversitylibrary.org/item/78508#page/494/mode/1up	BM 1894.10.9.2		"Isabella, N.E. Luzon," Philippines.			whiteheadi O. Thomas, 1894	previously included K. pusilla (along with bicolor)	Sedlock, J. L., Heaney, L. R., Balete, D. S., & Ruedi, M. (2020). Philippine bats of the genus Kerivoula (Chiroptera: Vespertilionidae): Overview and assessment of variation in K. pellucida and K. whiteheadi. Zootaxa, 4755(3), 454-490.				Philippines	Asia	Indomalaya	LC	0	0	0	Kerivoula_whiteheadi	0	sciname match	Kerivoula_whiteheadi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_whiteheadi	1005309	23	Whitehead's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	whiteheadi	O. Thomas	0	Kerivoula Whiteheadi	Thomas, O. 1894-12-01. Descriptions of two new bats of the genus _Kerivoula_. Annals and Magazine of Natural History (6)14(84):460-462.	https://www.biodiversitylibrary.org/page/15626287	BMNH:Mamm:1894.10.9.2	holotype	https://data.nhm.ac.uk/object/2015fdc9-0e58-484d-b431-1e2b86ab5bea	"Isabella, N.E. Luzon," Philippines.			previously included K. pusilla (along with bicolor)	Sedlock, J. L., Heaney, L. R., Balete, D. S., & Ruedi, M. (2020). Philippine bats of the genus Kerivoula (Chiroptera: Vespertilionidae): Overview and assessment of variation in K. pellucida and K. whiteheadi. Zootaxa, 4755(3), 454-490.				Philippines	Asia	Indomalaya	LC	0	0	0	Kerivoula_whiteheadi	0	sciname match	Kerivoula_whiteheadi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		whiteheadi	Thomas	1894	0	Ann. Mag. Nat. Hist.	ser. 6, 14: 460	Whitehead's Woolly Bat	None.	Philippines, Luzon, Isabella, Molino.	Philippines, Borneo, S Thailand, W Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10987/22023276/' target='_blank'>Least Concern</a>	Does not include pusilla or bicolor, which we recognize as a subspecies of pusilla following Sedlock et al. (2020). Reviewed by Hill (1965) and Corbet and Hill (1992). Phillippines whiteheadi reviewed by Sedlock et al. (2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula whiteheadi; Kerivoula whiteheadi; Kerivoula whiteheadi; Kerivoula whiteheadi; Kerivoula whiteheadi; Kerivoula whiteheadi; whiteheadi; bicolor; pusilla; whiteheadi; bicolor; pusilla; whiteheadi; Kérivoule de Whitehead; Whitehead-Wollfledermaus; Querivoula de Whitehead; Whitehead's Woolly Bat; Whitehead's Woolly Bat; Whitehead's Woolly Bat; K. whiteheadi