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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L515	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania	Kerivoula titania		[HMW] Kerivoula titania Bates et al., 2007 , “Seima Biodiversity Conservation Area, Mondul Kiri Province , Cambodia , 12°10.49°N , 106°58.55'E , 290. m asl.” Kerivoula titania appears to be sister to K. minuta , and this clade is sister to the rest of the hardwickii species group, although it might also be related to K. intermedia . Specimens previously attributed to K. fitania from Hainan and Taiwan islands now represent K. furva . Monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														titania				titania	titania			titania P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, C. M. Francis, & Csorba, 2007						N/A																																								NA																											4C3D87E8FF716ACFFA519FCD1DFEBFB9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	898	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF716ACFFA519FCD1DFEBFB9.xml	Kerivoula titania	Vespertilionidae	Kerivoula	titania	Bates	2007	Kérivoule titania @fr | Titania-Wollfledermaus @de | Querivoulatitania @es	Kerivoula titania Bates et al., 2007 , “Seima Biodiversity Conservation Area, Mondul Kiri Province , Cambodia , 12°10.49°N , 106°58.55'E , 290. m asl.” Kerivoula titania appears to be sister to K. minuta , and this clade is sister to the rest of the hardwickii species group, although it might also be related to K. intermedia . Specimens previously attributed to K. fitania from Hainan and Taiwan islands now represent K. furva . Monotypic.	E Myanmar , N Thailand , Laos , Vietnam , and E Cambodia .	Head—body 38- 4—41 mm , tail 45-8-53- 5 mm , ear 12- 5-15 mm , hindfoot 7-1-8- 3 mm , forearm 32-4-35- 9 mm ; weight 4-7- 9 g . Dorsal pelage is brownish gray (hairs with black bases, light gray middles, and dark gray tips); venteris a little lighter (hairs black with whitish tips tinged with gray-brown). Face and lips are hairy, except nostrils. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are semi-translucent and gray, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes. Fleshy lobe at base of thumb is oval and smooth in middle. Skull has broad, flattened braincase; rostrum has large narial pit and well developed rostral sulcus; and basioccipital pits are shallow. I? is unicuspid, with cingulum on its internal posterior border; I is equal in crown area to I?, with well-defined internally positioned cusp; canines are smooth, with deep longitudinal groove on outer sides; upper premolars are compressed in tooth row; M? lacks metastyle; I and I, are tricuspid, and I, is smaller with only well-defined central cusp and no lateral cusps; lower premolars are compressed in tooth row; and talonid of M,is reduced to ¢.70% of trigonid size.	Lowland and montane primary and secondary evergreen and deciduous forests at elevations of 260-1600 m ( Vietnam ).	No information.	Female Titania’s Woolly Bats in late pregnancy were caughtin early May in Ba Be National Park, northern Vietnam , and late April in Laos . Two pregnant females were captured in mid-May and two lactating females in early June in Laos , suggesting births occur in late May and possibly earlyJune.	In Thailand , call shape is steep FM sweep, with start frequencies of 217-243 kHz (mean 227 kHz), end frequencies of 74-8-90 kHz (82-6 kHz), and peak frequencies of 137-7-168-7 kHz (157 kHz).	No information.	Classified as Least Concern on The IUCN Red List. Titania's Woolly Bat does not seem to currently face any major threats, but becauseit is forest dependent, logging and agricultural development might be future threats.	Bates, Struebig et al. (2007) | Chanson (2008) | Douangboubpha et al. (2016) | Kruskop (2013a) | Kuo Haochih et al. (2017) | Tu Vuong Tan et al. (2018) | Wu Yi, Li Yuchun et al. (2012)	https://zenodo.org/record/6398458/files/figure.png	305. Titania’s Woolly Bat Kerivoula titania French: Kérivoule titania / German: Titania-Wollfledermaus / Spanish: Querivoula titania Taxonomy. Kerivoula titania Bates et al., 2007 , “Seima Biodiversity Conservation Area, Mondul Kiri Province , Cambodia , 12°10.49°N , 106°58.55'E , 290. m asl.” Kerivoula titania appears to be sister to K. minuta , and this clade is sister to the rest of the hardwickii species group, although it might also be related to K. intermedia . Specimens previously attributed to K. fitania from Hainan and Taiwan islands now represent K. furva . Monotypic. Distribution. E Myanmar , N Thailand , Laos , Vietnam , and E Cambodia . Descriptive notes. Head—body 38- 4—41 mm , tail 45-8-53- 5 mm , ear 12- 5-15 mm , hindfoot 7-1-8- 3 mm , forearm 32-4-35- 9 mm ; weight 4-7- 9 g . Dorsal pelage is brownish gray (hairs with black bases, light gray middles, and dark gray tips); venteris a little lighter (hairs black with whitish tips tinged with gray-brown). Face and lips are hairy, except nostrils. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are semi-translucent and gray, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes. Fleshy lobe at base of thumb is oval and smooth in middle. Skull has broad, flattened braincase; rostrum has large narial pit and well developed rostral sulcus; and basioccipital pits are shallow. I? is unicuspid, with cingulum on its internal posterior border; I is equal in crown area to I?, with well-defined internally positioned cusp; canines are smooth, with deep longitudinal groove on outer sides; upper premolars are compressed in tooth row; M? lacks metastyle; I and I, are tricuspid, and I, is smaller with only well-defined central cusp and no lateral cusps; lower premolars are compressed in tooth row; and talonid of M,is reduced to ¢.70% of trigonid size. Habitat. Lowland and montane primary and secondary evergreen and deciduous forests at elevations of 260-1600 m ( Vietnam ). Food and Feeding. No information. Breeding. Female Titania’s Woolly Bats in late pregnancy were caughtin early May in Ba Be National Park, northern Vietnam , and late April in Laos . Two pregnant females were captured in mid-May and two lactating females in early June in Laos , suggesting births occur in late May and possibly earlyJune. Activity patterns. In Thailand , call shape is steep FM sweep, with start frequencies of 217-243 kHz (mean 227 kHz), end frequencies of 74-8-90 kHz (82-6 kHz), and peak frequencies of 137-7-168-7 kHz (157 kHz). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Titania's Woolly Bat does not seem to currently face any major threats, but becauseit is forest dependent, logging and agricultural development might be future threats. Bibliography. Bates, Struebig et al. (2007), Chanson (2008), Douangboubpha et al. (2016), Kruskop (2013a), Kuo Haochih et al. (2017), Tu Vuong Tan et al. (2018), Wu Yi, Li Yuchun et al. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula titania	Kerivoula		titania	Bates, Struebig, Hayes, Furey, Mya, Thong, Tien, Son, Harrison, Francis, & Csorba	2007	0	Acta Chiropterol.	9(2): 326	Titania&apos;s Woolly Bat	None.	Cambodia, Mondul Kiri Province Seima Biodiversity Conservation Area	Yunnan (China) and Myanmar, Thailand, Laos, Cambodia, Vietnam	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula titania	23	Titania's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	titania	P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, Francis, & Csorba	2007	0	Kerivoula_titania	Bates, P. J., Struebig, M. J., Hayes, B. D., Furey, N. M., Mya, K. M., Thong, V. D., Tien, P. D., Son, N. T., Harrison, D. L, Francis, C. M., & Csorba, G. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica, 9(2), 326.	https://bioone.org/journals/acta-chiropterologica/volume-9/issue-2/1733-5329(2007)9[323:ANSOKC]2.0.CO;2/A-new-species-of-Kerivoula-Chiroptera--Vespertilionidae-from-Southeast/10.3161/1733-5329(2007)9[323:ANSOKC]2.0.CO;2.full	HNHM 2005.81.58		"Seima Biodiversity Conservation Area, Mondul Kiri Province, Cambodia, 12°10.49'N, 106°58.55'E, 290 m a.s.l."	12.175	106.98	titania P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, Francis, & Csorba, 2007	recently described	Bates, P. J., Struebig, M. J., Hayes, B. D., Furey, N. M., Mya, K. M., Thong, V. D., ... & Csorba, G. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica, 9(2), 323-337.	Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Kerivoula_titania	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136817	Kerivoula titania	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	titania	Bates, Struebig, Hayes, Furey, Mya Mya, Thong, Tien, Son, Harrison, Francis &; Csorba, 2007		20000000	Kerivoula titania	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern, although only recently described, it is known from a large area, it has a presumed large population, and is unlikely to be declining fast enough for listing in a more threatened category.	It has been collected in lowland and montane, primary and secondary evergreen and deciduous forest, often associated with areas of limestone. It is most likely a forest dependent species. Nothing is known of the roosting behaviour of this species.	There are no major threats to this species throughout its range. Being a forest dependent species, logging and agricultural development is an ongoing threat to some populations.	There is no information available on the abundance or population trends of this species. It has only been recently described, but in the past, specimens of this species had been assigned to other Kerivoula species. Hence its recent description is not an indication or rarity but rather taxonomic revision.	Unknown	This species is currently known from across Southeast Asia (Myanmar, Thailand, Cambodia, Lao PDR and Viet Nam), Guangdong Province and Hainan Island (China) and Taiwan. It has been collected up to 1,600 m asl in Viet Nam, ca. 923 m asl in Hainan Island, and ca. 350-920 m asl in Taiwan.		Terrestrial	This species has been recorded from many protected areas within its range.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		titania	Bates, Struebig, Hayes, Furey, Mya, Thong, Tien, Son, Harrison, Francis, & Csorba	2007	0	Acta Chiropterol.	9(2): 326	Titania&apos;s Woolly Bat	None.	Cambodia, Mondul Kiri Province Seima Biodiversity Conservation Area	Yunnan (China) and Myanmar, Thailand, Laos, Cambodia, Vietnam	Not listed.	Least Concern		Kerivoula titania	1005308	23	Titania's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	titania	P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, Francis, & Csorba	2007	0	Kerivoula_titania	Bates, P. J., Struebig, M. J., Hayes, B. D., Furey, N. M., Mya, K. M., Thong, V. D., Tien, P. D., Son, N. T., Harrison, D. L, Francis, C. M., & Csorba, G. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica, 9(2), 326.	https://bioone.org/journals/acta-chiropterologica/volume-9/issue-2/1733-5329(2007)9[323:ANSOKC]2.0.CO;2/A-new-species-of-Kerivoula-Chiroptera--Vespertilionidae-from-Southeast/10.3161/1733-5329(2007)9[323:ANSOKC]2.0.CO;2.full	HNHM 2005.81.58		"Seima Biodiversity Conservation Area, Mondul Kiri Province, Cambodia, 12°10.49'N, 106°58.55'E, 290 m a.s.l."	12.17483	106.9758	titania P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, Francis, & Csorba, 2007	recently described	Bates, P. J., Struebig, M. J., Hayes, B. D., Furey, N. M., Mya, K. M., Thong, V. D., ... & Csorba, G. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica, 9(2), 323-337.				Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Kerivoula_titania	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_titania	1005308	23	Titania's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	titania	P. J. J. Bates, Struebig, Hayes, Furey, Khin Mya Mya, Vu Dinh Thong, Pham Duc Tien, Nguyen Truong Son, D. L. Harrison, C. M. Francis, & Csorba	0	Kerivoula titania	Bates, P.J.J., Struebig, M.J., Hayes, B.D., Furey, N.M., Khin Mya Mya, Thong, V.D., Tien, P.D., Son, N.T., Harrison, D.L., Francis, C.M. and Csorba, G. 2007-12. A new species of _Kerivoula_ (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica 9(2):323-337.	https://doi.org/10.3161/1733-5329(2007)9[323:ANSOKC]2.0.CO;2	HNHM 2005.81.58	holotype		"Seima Biodiversity Conservation Area, Mondul Kiri Province, Cambodia, 12°10.49'N, 106°58.55'E, 290 m a.s.l."	12.17483	106.9758	recently described	Bates, P. J., Struebig, M. J., Hayes, B. D., Furey, N. M., Mya, K. M., Thong, V. D., ... & Csorba, G. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from Southeast Asia. Acta Chiropterologica, 9(2), 323-337.				Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Kerivoula_titania	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		titania	Bates, Struebig, Hayes, Furey, Mya, Thong, Tien, Son, Harrison, Francis & Csorba	2007	0	Acta Chiropterol.	9(2): 326	Titania&apos;s Woolly Bat	None.	Cambodia, Mondul Kiri Province Seima Biodiversity Conservation Area	Yunnan (China) and Myanmar, Thailand, Laos, Cambodia, Vietnam	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136817/22044302/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula titania; Kerivoula titania; Kerivoula titania; Kerivoula titania; Kerivoula titania; titania; Kérivoule titania; Titania-Wollfledermaus; Querivoulatitania; Titania's Woolly Bat; Titania&apos;s Woolly Bat; K. titania