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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L514	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula smithi	Kerivoula smithi	Kerivoula smithii	Kerivoula smithi	Kerivoula smithi	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii	Kerivoula smithii		[MSW2] Subgenus Kerivoula.; [MSW3] Sometimes mispelled smithi but the original spelling is smithii.; [HMW] Kerivoula smithii Thomas, 1880 , “Old Calabar,” Nigeria . This species is monotypic.; [batnames2022] Sometimes mispelled smithi but the original spelling is smithii .; [batnames2023] Sometimes mispelled smithi but the original spelling is smithii .; [batnames2025_1.7] Sometimes mispelled smithi but the original spelling is smithii.														smithii				smithii	smithii, smithi			smithii O. Thomas, 1880|smithi O. Thomas, 1901 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Smith's woolly bat	Nigeria – E Zaire, Kenya	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula smithi	Nigeria, Calabar.	Thomas	1880	Ann. Mag. Nat. Hist., ser. 5, 6:166.	Distribution: Known only from forested regions of Nigeria, Cameroon, northern Zaire, and Kenya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Smith's woolly bat	Nigeria – E Zaire, Kenya	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1880	Ann. Mag. Nat. Hist., ser. 5, 6:166.	Subgenus Kerivoula.	Nigeria, Cameroon, N and E Zaire, Kenya, Ivory Coast, Liberia.	Nigeria, Calabar.		THOMAS	1880	Rostrum fairly short and not narrowed anteriorly. Anterior and middle upper premolars more or less circular in section but anterior and middle lowers elongate. Lingual margins of anterior and middle upper molars more or less rounded, but with broad shelves. Size medium (forearm length, 32-35 mm; condylobasal length, 12-14 mm). Inner upper incisor bicuspid and relatively long. Fringe of hair on posterior margin of uropatagium poorly developed. Outer lower incisors unicuspid.	Distribution: Known only from forested regions of Nigeria, Cameroon, northern Zaire, and Kenya.	No subspecies.		98	species	K. smithi	THOMAS	1880	Kerivoula	subgenus	Kerivoula smithi				Rostrum fairly short and not narrowed anteriorly. Anterior and middle upper premolars more or less circular in section but anterior and middle lowers elongate. Lingual margins of anterior and middle upper molars more or less rounded, but with broad shelves. Size medium (forearm length, 32-35 mm; condylobasal length, 12-14 mm). Inner upper incisor bicuspid and relatively long. Fringe of hair on posterior margin of uropatagium poorly developed. Outer lower incisors unicuspid.	No subspecies.		4. K. smithi THOMAS 1880.	4	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula smithii	Kerivoula		smithii	Thomas		1880		Ann. Mag. Nat. Hist., ser. 5	6		166		Smith's Woolly Bat	Nigeria, Calabar.	Nigeria, Cameroon, N and E Dem. Rep. Congo, Kenya. Previous records from Côte d’Ivoire and Liberia are apparently erroneous (J. Fahr, pers. comm.)	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Sometimes mispelled smithi but the original spelling is smithii.	4C3D87E8FF746ACBFF7A920C1D56B047	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	903	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF746ACBFF7A920C1D56B047.xml	Kerivoula smithii	Vespertilionidae	Kerivoula	smithii	Thomas	1880	Kérivoule de Smith @fr | Smith-Wollfledermaus @de | Querivoula de Smith @es	Kerivoula smithii Thomas, 1880 , “Old Calabar,” Nigeria . This species is monotypic.	Known from a few scattered records from Nigeria and Cameroon E through DR Congo to Uganda and Kenya .	Head-body ¢.39 44 mm , tail 40-49 mm , ear 11-15 mm , hindfoot 5-7-8- 2 mm , forearm 32-36 mm ; weight 5-9- 7 g . Sexes are similar. Pelage is dense, woolly, and frizzled, with many hairs having hooked tips. Dorsal pelage is grayish brown to dark brown, with silvery sheen and silvery or straw-colored frosting. Dorsal hairs are blackish or dark brown, often with silvery or straw-colored tips. Mid-dorsal hairs are ¢. 9 mm . Ventral pelage is paler brown. Ventral hairs are grayish brown or blackish brown, usually with pale brown or fawn tips, but some have whitish silvery tips. Wings are blackish or dark brown, with scattered hairs extending to dorsal surfaces of forearm, third digit, thumb, tibia, and tail. Uropatagium is dark brown, with posterior margin and sparse hairs but no comb-like fringe of hooked bristle-like hairs. Ears are widely separated, funnel-shaped, short for a vespertilionid, dark brown or blackish, and deeply emarginated below tips. Tragusis long, tapering, and curved outward, and base has small triangular lobule over shallow emargination. Eyes are minute. Head is high-domed. Muzzle is long, pointed, partly hidden byfacial pelage. Tail is ¢.92-131% of head-body length. Skull is small (greatest skull lengths 13-1-14- 1 mm ); braincase is high-domed; rostrum is narrow; and frontal region is sharply angular to plane as in otherspecies of Kerivoula . I? is bicuspid andtaller than I’; I? is unicuspid or weakly bicuspid; P? is slightly higher than P?, and much smaller than P%I and I, are tricuspid; I, is unicuspid; and lower premolars are subequal in size.	Mainly lowland rainforests but also mangroves (southern Nigeria ), Congolese swamp forests DR Congo ), montane forests ( Cameroon highlands, Uganda , and Kenya ), and riverine forests in Somalia-Masai bushland, at elevations of 900-2800 m .	No information.	In Kenya , a pregnant Smith’s Woolly Bat was captured in September from Bura, near Garissa , and another in December from Warwick. In Uganda , two females were pregnant in late November. In north-eastern DR Congo , two females each with an attached young were found in early October. Litter size is one.	Fourteen calls from the Aberdare Range in Kenya had mean peak frequency of95-9 (81-5-104-9 kHz), maximum frequency of 126-3 (115-6-137-2 kHz), minimum frequency of 65-3 (44-9-91-8 kHz), duration of 61 milliseconds (25-1-85-5 milliseconds), and interpulse interval of 0-2 milliseconds (0-1-0-6 milliseconds).	Ectoparasites include the mite Rodhainyssus myotis (Acari, Gastronyssidae ).	Classified as Least Concern on The IUCN Red List.	ACR (2017) | Eisenring et al. (2016) | Koopman (1994)	https://zenodo.org/record/6398492/files/figure.png	320. Smith’s Woolly Bat Kerivoula smithii French: Kérivoule de Smith / German: Smith-Wollfledermaus / Spanish: Querivoula de Smith Taxonomy. Kerivoula smithii Thomas, 1880 , “Old Calabar,” Nigeria . This species is monotypic. Distribution. Known from a few scattered records from Nigeria and Cameroon E through DR Congo to Uganda and Kenya . Descriptive notes. Head-body ¢.39 44 mm , tail 40-49 mm , ear 11-15 mm , hindfoot 5-7-8- 2 mm , forearm 32-36 mm ; weight 5-9- 7 g . Sexes are similar. Pelage is dense, woolly, and frizzled, with many hairs having hooked tips. Dorsal pelage is grayish brown to dark brown, with silvery sheen and silvery or straw-colored frosting. Dorsal hairs are blackish or dark brown, often with silvery or straw-colored tips. Mid-dorsal hairs are ¢. 9 mm . Ventral pelage is paler brown. Ventral hairs are grayish brown or blackish brown, usually with pale brown or fawn tips, but some have whitish silvery tips. Wings are blackish or dark brown, with scattered hairs extending to dorsal surfaces of forearm, third digit, thumb, tibia, and tail. Uropatagium is dark brown, with posterior margin and sparse hairs but no comb-like fringe of hooked bristle-like hairs. Ears are widely separated, funnel-shaped, short for a vespertilionid, dark brown or blackish, and deeply emarginated below tips. Tragusis long, tapering, and curved outward, and base has small triangular lobule over shallow emargination. Eyes are minute. Head is high-domed. Muzzle is long, pointed, partly hidden byfacial pelage. Tail is ¢.92-131% of head-body length. Skull is small (greatest skull lengths 13-1-14- 1 mm ); braincase is high-domed; rostrum is narrow; and frontal region is sharply angular to plane as in otherspecies of Kerivoula . I? is bicuspid andtaller than I’; I? is unicuspid or weakly bicuspid; P? is slightly higher than P?, and much smaller than P%I and I, are tricuspid; I, is unicuspid; and lower premolars are subequal in size. Habitat. Mainly lowland rainforests but also mangroves (southern Nigeria ), Congolese swamp forests DR Congo ), montane forests ( Cameroon highlands, Uganda , and Kenya ), and riverine forests in Somalia-Masai bushland, at elevations of 900-2800 m . Food and Feeding. No information. Breeding. In Kenya , a pregnant Smith’s Woolly Bat was captured in September from Bura, near Garissa , and another in December from Warwick. In Uganda , two females were pregnant in late November. In north-eastern DR Congo , two females each with an attached young were found in early October. Litter size is one. Activity patterns. Fourteen calls from the Aberdare Range in Kenya had mean peak frequency of95-9 (81-5-104-9 kHz), maximum frequency of 126-3 (115-6-137-2 kHz), minimum frequency of 65-3 (44-9-91-8 kHz), duration of 61 milliseconds (25-1-85-5 milliseconds), and interpulse interval of 0-2 milliseconds (0-1-0-6 milliseconds). Movements, Home range and Social organization. Ectoparasites include the mite Rodhainyssus myotis (Acari, Gastronyssidae ). Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. ACR (2017), Eisenring et al. (2016), Koopman (1994).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula smithii	Kerivoula		smithii	Thomas	1880	0	Ann. Mag. Nat. Hist.	ser. 5, 6: 166	Smith's Woolly Bat	None.	Nigeria, Calabar.	Nigeria, Cameroon, N and E Dem. Rep. Congo, Kenya. Previous records from Côte d’Ivoire and Liberia are apparently erroneous (J. Fahr, pers. comm.)	Not listed.	Least Concern	Sometimes mispelled smithi but the original spelling is smithii .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula smithii	23	Smith's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	smithii	O. Thomas	1880	0	Kerivoula_smithii	Thomas, O. (1880). On bats from Old Calabar. Annals and Magazine of Natural History, ser. 5, 6, 166.	https://www.biodiversitylibrary.org/item/81219#page/180/mode/1up	BM 1880.7.21.9		"Old Calabar," Nigeria.			smithii O. Thomas, 1880	NA	NA	Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya	Africa	Afrotropic	LC	0	0	0	Kerivoula_smithii	0	sciname match	Kerivoula_smithii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10986	Kerivoula smithii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	smithii	Thomas, 1880		20000000	Kerivoula smithii	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species has been recorded from several forest types including lowland and montane tropical moist forest, tropical dry forest and swamp forest. Happold (1987) reported that in Nigeria, the species is found in 'the rainforest zone east of the Niger River'. It may use the nests of weaver birds and sunbirds, and tree cavities as roosting sites (Happold 1987). It is not known if the species is adaptable to habitat disturbance.	There appear to be no major threats to this little known species.	This species has been very rarely recorded (Happold 1987).	Unknown	This African bat is known from scattered records in a distribution band ranging from Nigeria and Cameroon in the west, through the Democratic Republic of the Congo, to Uganda and Kenya in the east. Earlier records from Côte d'Ivoire and Liberia (Happold 1987; Koopman et al. 1995) appear to be in error (J. Fahr pers. comm. in Simmons 2005). It has been recorded at elevations of 900 to 2,800 m Asl.		Terrestrial	It is not known if the species is present within any protected areas. Further studies are needed into the distribution, abundance, and general ecology of this apparently rare species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		smithii	Thomas	1880	0	Ann. Mag. Nat. Hist.	ser. 5, 6: 166	Smith's Woolly Bat	None.	Nigeria, Calabar.	Nigeria, Cameroon, N and E Dem. Rep. Congo, Kenya. Previous records from Côte d’Ivoire and Liberia are apparently erroneous (J. Fahr, pers. comm.)	Not listed.	Least Concern	Sometimes mispelled smithi but the original spelling is smithii .	Kerivoula smithii	1005307	23	Smith's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	smithii	O. Thomas	1880	0	Kerivoula_smithii	Thomas, O. (1880). On bats from Old Calabar. Annals and Magazine of Natural History, ser. 5, 6, 166.	https://www.biodiversitylibrary.org/item/81219#page/180/mode/1up	BM 1880.7.21.9		"Old Calabar," Nigeria.			smithii O. Thomas, 1880	NA	NA				Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya	Africa	Afrotropic	LC	0	0	0	Kerivoula_smithii	0	sciname match	Kerivoula_smithii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_smithii	1005307	23	Smith's Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	smithii	O. Thomas	0	Kerivoula smithii	Thomas, O. 1880-08-01. On bats from Old Calabar. Annals and Magazine of Natural History (5)6(32):164-167.	https://www.biodiversitylibrary.org/page/25170391	BMNH:Mamm:1880.7.21.9	holotype	https://data.nhm.ac.uk/object/e1dfff7e-210e-4e5e-ac0f-3338673d63fb	"Old Calabar," Nigeria.			NA	NA				Nigeria|Cameroon|Democratic Republic of the Congo|Uganda|Kenya	Africa	Afrotropic	LC	0	0	0	Kerivoula_smithii	0	sciname match	Kerivoula_smithii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		smithii	Thomas	1880	0	Ann. Mag. Nat. Hist.	ser. 5, 6: 166	Smith's Woolly Bat	None.	Nigeria, Calabar.	Nigeria, Cameroon, N and E Dem. Rep. Congo, Kenya. Previous records from Côte d’Ivoire and Liberia are apparently erroneous (J. Fahr, pers. comm.)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10986/22023189/' target='_blank'>Least Concern</a>	Sometimes mispelled smithi but the original spelling is smithii.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula smithii; Kerivoula smithii; Kerivoula smithii; Kerivoula smithii; Kerivoula smithii; Kerivoula smithii; smithii; Kérivoule de Smith; Smith-Wollfledermaus; Querivoula de Smith; Smith's Woolly Bat; Smith's Woolly Bat; Smith's Woolly Bat; K. smithii