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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L509	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida	Kerivoula pellucida		[MSW2] Subgenus Kerivoula. Includes bombifrons-, see Hill (1965:539).; [MSW3] Includes bombifrons; see Hill (1965).; [HMW] Vespertilio pellucidus Waterhouse, 1845 , Philippines . Phylogenetic position of K. pellucida is uncertain. It might be related to the African radiation of Kerivoula or to K. picta rather than to other Asiatic Kerivoula . Monotypic.; [batnames2022] Includes bombifrons ; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).; [IUCN] This species requires taxonomic review (L. Heaney pers. comm. 2006).; [batnames2023] Includes bombifrons ; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).; [batnames2025_1.7] Includes bombifrons; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).				bonibifrons		bombifrons,			bombifrons			pellucida	pellucida - bombifrons	pellucida, bombifrons	This species requires taxonomic review (L. Heaney pers. comm. 2006).	pellucida	pellucida - bombifrons	pellucida, bombifrons	pellucida, bombifrons	pellucida	pellucida - bombifrons	pellucida (G. R. Waterhouse, 1845)|bombifrons Lyon, 1911		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Clear-winged bat	Philippines, Malaya, Borneo, ? Sumatra, Java	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula pellucida	Philippines.	Waterhouse	1845	Proc. Zool. Soc. Lond., 1845:6.	Distribution: Known from Malaya, Sumatra, Java, Borneo, and the southern Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Clear-winged bat	Philippines, Malaya, Borneo, ? Sumatra, Java	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Waterhouse	1845	Proc. Zool. Soc. Lend., 1845:6.	Subgenus Kerivoula. Includes bombifrons-, see Hill (1965:539).	Borneo, Philippines, Java and Sumatra, W Malaysia.	Philippines.		WATERHOUSE	1845	Rostrum of medium proportions, slightly narrowed anteriorly. Anterior upper and lower premolars and middle upper premolar nearly circular in section but middle lower slightly ovate. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size fairly small (forearm length, 28-35 mm; condylobasal length, 12-14 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	Distribution: Known from Malaya, Sumatra, Java, Borneo, and the southern Philippines.	No subspecies.		99	species	K. pellucida	WATERHOUSE	1845	Kerivoula	subgenus	Kerivoula pellucida				Rostrum of medium proportions, slightly narrowed anteriorly. Anterior upper and lower premolars and middle upper premolar nearly circular in section but middle lower slightly ovate. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size fairly small (forearm length, 28-35 mm; condylobasal length, 12-14 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	No subspecies.		13. K. pellucida (WATERHOUSE 1845).	13	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula pellucida	Kerivoula		pellucida	Waterhouse	y	1845		Proc. Zool. Soc. Lond.	1845		6		Clear-winged Woolly Bat	Philippines.	Borneo, Philippines, Java and Sumatra, W Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	bombifrons Lyon, 1911.	Includes bombifrons; see Hill (1965).	4C3D87E8FF776AC8FF529F0D16C5B863	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	900	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF776AC8FF529F0D16C5B863.xml	Kerivoula pellucida	Vespertilionidae	Kerivoula	pellucida		1845	Kérivoule diaphane @fr | Transparentfligel-Wollfledermaus @de | Querivoulade alasclaras @es	Vespertilio pellucidus Waterhouse, 1845 , Philippines . Phylogenetic position of K. pellucida is uncertain. It might be related to the African radiation of Kerivoula or to K. picta rather than to other Asiatic Kerivoula . Monotypic.	Thai-Malay Peninsula, Sumatra, Borneo except EC & SE, Java, and Philippines ( Palawan , Cebu , Bohol , Mindanao, and Jolo Is).	Head-body 35-4 41- 4 mm , tail 40-51- 8 mm , ear 13- 2-17 mm , hindfoot 5-4-7- 8 mm , forearm 29- 32.1 mm; weight 3-5-5- 5 g . Dorsal pelage is pale orange-brown (hairs with pale grayish white bases); venter is grayish white. Face is pale orangish to pinkish. Membranes and ears are pale orange-brown and nearly completely translucent. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity neartip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and calcar is long. Compared with Hardwicke’s Woolly Bat ( K. hardwickii ), skull has narrower interorbital width, with generally more elongated palatal length and narrower cranial length (both with some overlap in measurements), and shape of anterior region to rostrum is more flattened.	Primarily understories of primary and secondary lowland forests from sea level up to elevations of ¢. 1200 m .	No information.	Female Clear-winged Woolly Bats have one young per pregnancy.	The Clear-winged Woolly Bat roosts in dead leaves of small shrubs in the Philippines and banana leaves in Indochina. In Peninsular Malaysia , calls are steep FM sweep, with start frequencies of 159-2-188-8 kHz (mean 178-5 kHz), end frequencies of 30-4-78-2 kHz (58-5 kHz), peak frequencies of 57-6-138-4 kHz (96-4 kHz), and durations of 1-4-4 milliseconds (2-3 milliseconds).	The Clear-winged Woolly Bat roosts in small groups.	Classified as Near Threatened on The IUCN Red List. The Clear-winged Woolly Bat is widespread and fairly common, but lowland primary forests are being deforested across its distribution.	Bumrungsri et al. (2006) | Douangboubpha et al. (2016) | Esselstyn, Widmann & Heaney (2004) | Francis (2008a) | Francis, Rosell-Ambal, Kingston & Nusalawo (2008) | Hasan & Abdullah (2011) | Heaney et al. (1998) | Hill (1965b) | Khan (2008) | Khan et al. (2010) | Kingston et al. (1999)	https://zenodo.org/record/6398468/files/figure.png	310. Clear-winged Woolly Bat Kerivoula pellucida French: Kérivoule diaphane / German: Transparentfligel-Wollfledermaus / Spanish: Querivoula de alasclaras Taxonomy. Vespertilio pellucidus Waterhouse, 1845 , Philippines . Phylogenetic position of K. pellucida is uncertain. It might be related to the African radiation of Kerivoula or to K. picta rather than to other Asiatic Kerivoula . Monotypic. Distribution. Thai-Malay Peninsula, Sumatra, Borneo except EC & SE, Java, and Philippines ( Palawan , Cebu , Bohol , Mindanao, and Jolo Is). Descriptive notes. Head-body 35-4 41- 4 mm , tail 40-51- 8 mm , ear 13- 2-17 mm , hindfoot 5-4-7- 8 mm , forearm 29- 32.1 mm; weight 3-5-5- 5 g . Dorsal pelage is pale orange-brown (hairs with pale grayish white bases); venter is grayish white. Face is pale orangish to pinkish. Membranes and ears are pale orange-brown and nearly completely translucent. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragusis narrow and tall, with virtually straight anterior margin except for very slight convexity neartip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and calcar is long. Compared with Hardwicke’s Woolly Bat ( K. hardwickii ), skull has narrower interorbital width, with generally more elongated palatal length and narrower cranial length (both with some overlap in measurements), and shape of anterior region to rostrum is more flattened. Habitat. Primarily understories of primary and secondary lowland forests from sea level up to elevations of ¢. 1200 m . Food and Feeding. No information. Breeding. Female Clear-winged Woolly Bats have one young per pregnancy. Activity patterns. The Clear-winged Woolly Bat roosts in dead leaves of small shrubs in the Philippines and banana leaves in Indochina. In Peninsular Malaysia , calls are steep FM sweep, with start frequencies of 159-2-188-8 kHz (mean 178-5 kHz), end frequencies of 30-4-78-2 kHz (58-5 kHz), peak frequencies of 57-6-138-4 kHz (96-4 kHz), and durations of 1-4-4 milliseconds (2-3 milliseconds). Movements, Home range and Social organization. The Clear-winged Woolly Bat roosts in small groups. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Clear-winged Woolly Bat is widespread and fairly common, but lowland primary forests are being deforested across its distribution. Bibliography. Bumrungsri et al. (2006), Douangboubpha et al. (2016), Esselstyn, Widmann & Heaney (2004), Francis (2008a), Francis, Rosell-Ambal, Kingston & Nusalawo (2008), Hasan & Abdullah (2011), Heaney et al. (1998), Hill (1965b), Khan (2008), Khan et al. (2010), Kingston et al. (1999).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula pellucida	Kerivoula		pellucida	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	######	Clear-winged Woolly Bat	 bombifrons Lyon, 1911.	Philippines.	Borneo, Philippines, Java and Sumatra, W Malaysia.	Not listed.	Near Threatened	Includes bombifrons ; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula pellucida	23	Clear-winged Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	pellucida	Waterhouse	1845	1						Philippines.			pellucida (Waterhouse, 1845)|bombifrons Lyon, 1911	NA	NA	Thailand|Indonesia|Brunei|Malaysia|Philippines	Asia	Indomalaya	NT	0	0	0	Kerivoula_pellucida	0	sciname match	Kerivoula_pellucida	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10983	Kerivoula pellucida	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	pellucida	(Waterhouse, 1845)	This species requires taxonomic review (L. Heaney pers. comm. 2006).	20000000	Kerivoula pellucida	Near Threatened	A3c	2020	2019-07-17 00:00:00 UTC	3.1	English	Kerivoula pellucida is ;listed as ;Near Threatened (nearly meets criterion A3c) as its global population is suspected to decline by ;an estimated 25-30% ;over the next 15 years (three generations; generation length = 5 years; Pacifici et al . 2013). The species is restricted to primary forests and its extent of occurrence (EOO), area of occupancy (AOO), and habitat is expected to continue to decline ;at a rapid rate from deforestation.	In Malaysia the species is generally dependent on primary forest (T. Kingston pers. comm. 2006). There are records of two groups roosting in dead leaves on small shrubs in lowland forest in the Philippines (Taylor 1934, Heaney et al . 1998). On Palawan Island, there are records from secondary lowland forest at 80 m asl (Esselstyn et al . 2004). Recorded at limestone forest areas of ca 500 m asl (Mohd-Azlan et al. 2005) and also reported in swamp and heath forest in Central Kalimantan (Struebig et al. 2006)	This species is dependent on primary forest which is being lost due to logging, plantations, conversion to agriculture and forest fires.	This is a common species in primary forests on Sumatra (M. Nusalawo pers. comm. 2006), and in Peninsular Malaysia it is fairly common in the understorey of primary forest. Harp and tunnel traps are required to survey this species. It is suspected that the population in Malaysia has undergone a decline of at least 30% in the past 15 years (T. Kingston pers. comm. 2006). Across its entire global range, deforestation, conversion of habitat to agriculture, and forest fires are expected to cause the population to decline by around 25-30%.	Decreasing	The species occurs on Borneo, Java, Peninsular Malaysia, extremem southern Thailand, Sumatra, and the Philippines. In the Philippines, there are records from Jolo (Taylor 1934), Mindanao (Davao del Norte Province) and Palawan (Heaney et al. 1998). In Sabah this species was recorded in Imbak Canyon, and Telupid.  This species is also reported as a common species in Western and Northeastern Sarawak by Shazali et al. (2018).		Terrestrial	The species occurs in a number of protected areas throughout its range.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		pellucida	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	########	Clear-winged Woolly Bat	 bombifrons Lyon, 1911.	Philippines.	Borneo, Philippines, Java and Sumatra, W Malaysia.	Not listed.	Near Threatened	Includes bombifrons ; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).	Kerivoula pellucida	1005303	23	Clear-winged Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	pellucida	Waterhouse	1845	1						Philippines.			pellucida (Waterhouse, 1845)|bombifrons Lyon, 1911	NA	NA				Thailand|Indonesia|Brunei|Malaysia|Philippines	Asia	Indomalaya	NT	0	0	0	Kerivoula_pellucida	0	sciname match	Kerivoula_pellucida	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_pellucida	1005303	23	Clear-winged Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	pellucida	G. R. Waterhouse	1	Vespertilio pellucidus	Waterhouse, G.R. 1845-04. Descriptions of species of Bats collectd in the Philippine Islands, and presented to the Society by H. Cuming, Esq. Proceedings of the Zoological Society of London 1845:3-10.	https://www.biodiversitylibrary.org/page/12862502	BMNH:Mamm:1855.12.26.264	holotype	https://data.nhm.ac.uk/object/ddb45b8e-6482-4b49-afb6-83c8f08c78a8	Philippines.			NA	NA				Thailand|Indonesia|Brunei|Malaysia|Philippines	Asia	Indomalaya	NT	0	0	0	Kerivoula_pellucida	0	sciname match	Kerivoula_pellucida	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		pellucida	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	########	Clear-winged Woolly Bat	bombifrons Lyon, 1911.	Philippines.	Borneo, Philippines, Java and Sumatra, W Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10983/22021330/' target='_blank'>Near Threatened</a>	Includes bombifrons; see Hill (1965). Phillippines pellucida reviewed by Sedlock et al. (2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula pellucida; Kerivoula pellucida; Kerivoula pellucida; Kerivoula pellucida; Kerivoula pellucida; Kerivoula pellucida; bombifrons; bombifrons; pellucida; bombifrons; Kérivoule diaphane; Transparentfligel-Wollfledermaus; Querivoulade alasclaras; Clear-winged Woolly Bat; Clear-winged Woolly Bat; Clear-winged Woolly Bat; K. pellucida
