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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L508	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa	Kerivoula papillosa		[MSW2] Subgenus Kerivoula.; [MSW3] Does not include lenis; see Vanitharani et al. (2003). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis, which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).; [HMW] Vespertilio papillosus Temminck, 1840 , “L’ile de Java , dans le district sauvage de Bantam . Se trouve aussi a Sumatra dans le district de Padang [= Theisland ofJava, in the wild district of Bantam . It is also found in Sumatra in the district of Padang].” Restricted by G. H. H. Tate in 1940 to “ Java .” As currently defined, K. papillosa represents a species complex that is paraphyletic with K. lenas, which togetherare sisterto the hardwickspecies group. There appearto be several sympatrically distributed forms (at least three), and manyspecimens currently attributed to K. papillosa likely represent K. lenis . Genetic and morphological data from across the distribution are needed to determine taxonomy of this complex. Kerivoula lenis was previously considered a subspecies of K. papillosa , but genetic and morphological data support its status as a separate species, especially because the two are broadly sympatric. Identity of Philippines specimens needs to be addressed further. Two subspecies recognized.; [batnames2022] Does not include lenis ; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis , which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).; [IUCN] This species does not include Kerivoula lenis (Simmons 2005), although some specimens referred to this species may represent lenis , which appears to be broadly sympatric with papillosa (Vanitharani et al . 2003).; [batnames2023] Does not include lenis ; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis , which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).; [batnames2025_1.7] Does not include lenis; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis, which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).						lenis, malayana.	lenis, papillosa, malayana	papillosa, malayana		papillosa, malayana		papillosa, malayana		papillosa, malayana	This species does not include Kerivoula lenis (Simmons 2005), although some specimens referred to this species may represent lenis , which appears to be broadly sympatric with papillosa (Vanitharani et al . 2003).	papillosa, malayana		papillosa, malayana	papillosa, malayana	malayana, papillosa		papillosa (Temminck, 1840)|malayana Chasen, 1940		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Papillose bat	NE India – Java, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula papillosa	Indonesia, Java, Bantam.	Temminck	1840	Monogr. Mamm., 2:220.	Distribution: Ranging from northeastern India to Vietnam, then south and east to Java and Celebes.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Papillose bat	NE India – Java, Borneo, Sulawesi; refs. 4.12, 143	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1840	Monogr. Mamm., 2:220.	Subgenus Kerivoula.	NE India, Vietnam, W Malaysia, Sumatra, Java, Sulawesi, Borneo.	Indonesia, Java, Bantam.		TEMMINCK	1840	Rostrum relatively short and broad, widened anteriorly. Anterior and middle premolars slightly elongate. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size relatively large (forearm length, 38-44 mm; condylobasal length, 14-17 mm). Inner upper incisor bicuspid, but secondary cusp small. No fringe of hair on posterior margin of uropatagium. Upper canine enlarged.	Distribution: Ranging from northeastern India to Vietnam, then south and east to Java and Celebes.	Three subspecies are recognized:	K. p. lenis (northeastern India, Burma), K. p. papillosa (Java), K. p. malayana (remainder of range).	99	species	K. papillosa	TEMMINCK	1840	Kerivoula	subgenus	Kerivoula papillosa				Rostrum relatively short and broad, widened anteriorly. Anterior and middle premolars slightly elongate. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size relatively large (forearm length, 38-44 mm; condylobasal length, 14-17 mm). Inner upper incisor bicuspid, but secondary cusp small. No fringe of hair on posterior margin of uropatagium. Upper canine enlarged.	Three subspecies are recognized:		16. K. papillosa (TEMMINCK 1840).	16	_K. p. malayana_ Chasen, 1940; _K. p. papillosa_ (Temminck, 1840)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula papillosa	Kerivoula		papillosa	Temminck	y	1840		Monogr. Mamm.	2		220		Papillose Woolly Bat	Indonesia, Java, Bantam (restricted by Tate, 1940).	Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Java, Sulawesi, Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	malayana Chasen, 1940.	Does not include lenis; see Vanitharani et al. (2003). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis, which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).	4C3D87E8FF706AC8FA7F90EA1CBDB800	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	899	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF706AC8FA7F90EA1CBDB800.xml	Kerivoula papillosa	Vespertilionidae	Kerivoula	papillosa		1840	Kérivoule mamelonnée @fr | Warzen-Wollfledermaus @de | Querivoulapiloso de Java @es | Papillose Bat @en	Vespertilio papillosus Temminck, 1840 , “L’ile de Java , dans le district sauvage de Bantam . Se trouve aussi a Sumatra dans le district de Padang [= Theisland ofJava, in the wild district of Bantam . It is also found in Sumatra in the district of Padang].” Restricted by G. H. H. Tate in 1940 to “ Java .” As currently defined, K. papillosa represents a species complex that is paraphyletic with K. lenas, which togetherare sisterto the hardwickspecies group. There appearto be several sympatrically distributed forms (at least three), and manyspecimens currently attributed to K. papillosa likely represent K. lenis . Genetic and morphological data from across the distribution are needed to determine taxonomy of this complex. Kerivoula lenis was previously considered a subspecies of K. papillosa , but genetic and morphological data support its status as a separate species, especially because the two are broadly sympatric. Identity of Philippines specimens needs to be addressed further. Two subspecies recognized.	K.p.papillosaTemminck,1840—Sumatra,N&WBorneo,Java,N&CSulawesi,includ-ingnearbyKabaenaandButonIs,andPhilippines(LuzonandBoholIs). K. p. malayana Chasen, 1940 — WC & S Thailand , NE Laos , Vietnam , C Cambodia , and Peninsular Malaysia .	Head-body 40-3-52- 2 mm , tail 47-56- 4 mm , ear 14-17 mm , hindfoot 7-9- 10 mm , forearm 39-49 mm ; weight 6-13 g . Furis dense and woolly. Dorsal pelage is brown to buffy brown; venter is paler. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragus is narrow and tall, with virtuallystraight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are light brown, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes, and calcar is long. Skull has high braincase and domed forehead, with more robust rostrum than in the Lenis Woolly Bat ( K. lenis ). Compared to the Lenis Woolly Bat, dentition is more robust overall, upper premolars are wider, and lower premolars are larger, especially P. Chromosomal complement has 2n = 38 and FN = 54 ( Thailand ).	Lowland mixed deciduous dipterocarp forests and secondary lowland forests (Luzon) from sea level up to elevations of ¢. 1050 m .	No information.	No information.	The Papillose Woolly Bat roosts in small tree hollows. Call shape is steep FM sweep. Calls of two (B and C group) ofthe three genetic lineages in Thailand were recorded: B group, start frequencies of 180-196 kHz (mean 190-3 kHz), end frequencies of 60-69 kHz (65-5 kHz), and peak frequencies of 109-9-157-6 kHz (131-6 kHz) and C group,start frequencies of 169-207-7 kHz (189-8 kHz), end frequencies of 63-5- 89-8 kHz (75-7 kHz), and peak frequencies of 112-1-161-4 kHz (140-8 kHz).	The Papillose Woolly Bat reportedly roosts in groups of 1-14 individuals.	Classified as Least Concern on The IUCN Red List. The Papillose Woolly Bat appears to be common in much of its wide distribution. Nevertheless, its taxonomy is unresolved, and it likely represents several species that would have smaller distributions. Further taxonomic studies are needed before a proper assessment can be made. Deforestation might be a threat in some regions.	Douangboubpha et al. (2016) | Duya et al. (2007) | Francis (2008a) | Hasan & Abdullah (2011) | Hutson & Kingston (2008e) | Khan et al. (2010) | Kruskop (2013a) | Kuo Haochih et al. (2017) | Struebig, Horsburgh et al. (2008) | Tate (1940) | Vanitharani et al. (2003)	https://zenodo.org/record/6398464/files/figure.png	308. Papillose Woolly Bat Kerivoula papillosa French: Kérivoule mamelonnée / German: Warzen-Wollfledermaus / Spanish: Querivoula piloso de Java Other common names: Papillose Bat Taxonomy. Vespertilio papillosus Temminck, 1840 , “L’ile de Java , dans le district sauvage de Bantam . Se trouve aussi a Sumatra dans le district de Padang [= Theisland ofJava, in the wild district of Bantam . It is also found in Sumatra in the district of Padang].” Restricted by G. H. H. Tate in 1940 to “ Java .” As currently defined, K. papillosa represents a species complex that is paraphyletic with K. lenas, which togetherare sisterto the hardwickspecies group. There appearto be several sympatrically distributed forms (at least three), and manyspecimens currently attributed to K. papillosa likely represent K. lenis . Genetic and morphological data from across the distribution are needed to determine taxonomy of this complex. Kerivoula lenis was previously considered a subspecies of K. papillosa , but genetic and morphological data support its status as a separate species, especially because the two are broadly sympatric. Identity of Philippines specimens needs to be addressed further. Two subspecies recognized. Subspecies and Distribution. K.p.papillosaTemminck,1840—Sumatra,N&WBorneo,Java,N&CSulawesi,includ-ingnearbyKabaenaandButonIs,andPhilippines(LuzonandBoholIs). K. p. malayana Chasen, 1940 — WC & S Thailand , NE Laos , Vietnam , C Cambodia , and Peninsular Malaysia . Descriptive notes. Head-body 40-3-52- 2 mm , tail 47-56- 4 mm , ear 14-17 mm , hindfoot 7-9- 10 mm , forearm 39-49 mm ; weight 6-13 g . Furis dense and woolly. Dorsal pelage is brown to buffy brown; venter is paler. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavityjust below tips on posterior borders; tragus is narrow and tall, with virtuallystraight anterior margin except for very slight convexity near tip and concave posterior margin with small hooked basal lobe. Membranes are light brown, with hairs scattered on posterior margin of uropatagium and no definite fringe; wings are attached at base of outer toes, and calcar is long. Skull has high braincase and domed forehead, with more robust rostrum than in the Lenis Woolly Bat ( K. lenis ). Compared to the Lenis Woolly Bat, dentition is more robust overall, upper premolars are wider, and lower premolars are larger, especially P. Chromosomal complement has 2n = 38 and FN = 54 ( Thailand ). Habitat. Lowland mixed deciduous dipterocarp forests and secondary lowland forests (Luzon) from sea level up to elevations of ¢. 1050 m . Food and Feeding. No information. Breeding. No information. Activity patterns. The Papillose Woolly Bat roosts in small tree hollows. Call shape is steep FM sweep. Calls of two (B and C group) ofthe three genetic lineages in Thailand were recorded: B group, start frequencies of 180-196 kHz (mean 190-3 kHz), end frequencies of 60-69 kHz (65-5 kHz), and peak frequencies of 109-9-157-6 kHz (131-6 kHz) and C group,start frequencies of 169-207-7 kHz (189-8 kHz), end frequencies of 63-5- 89-8 kHz (75-7 kHz), and peak frequencies of 112-1-161-4 kHz (140-8 kHz). Movements, Home range and Social organization. The Papillose Woolly Bat reportedly roosts in groups of 1-14 individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Papillose Woolly Bat appears to be common in much of its wide distribution. Nevertheless, its taxonomy is unresolved, and it likely represents several species that would have smaller distributions. Further taxonomic studies are needed before a proper assessment can be made. Deforestation might be a threat in some regions. Bibliography. Douangboubpha et al. (2016), Duya et al. (2007), Francis (2008a), Hasan & Abdullah (2011), Hutson & Kingston (2008e), Khan et al. (2010), Kruskop (2013a), Kuo Haochih et al. (2017), Struebig, Horsburgh et al. (2008), Tate (1940), Vanitharani et al. (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula papillosa	Kerivoula		papillosa	Temminck	1840	1	Monogr. Mamm.	0.2361	Papillose Woolly Bat	<b> malayana </b>Chasen, 1940.	Indonesia, Java, Bantam (restricted by Tate, 1940).	Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Java, Sulawesi, Borneo.	Not listed.	Least Concern	Does not include lenis ; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis , which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula papillosa	23	Papillose Woolly Bat	Papillose Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	papillosa	Temminck	1840	1						"L'ïle de Java, dans le district sauvage de Bantam. Se trouve aussi à Sumatra dans le district de Padang [= The island of Java, in the wild district of Bantam. It is also found in Sumatra in the district of Padang]." Restricted by G. H. H. Tate in 1940 to "Java."			papillosa (Temminck, 1840)|malayana Chasen, 1940	NA	NA	Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Kerivoula_papillosa	0	sciname match	Kerivoula_papillosa	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10981	Kerivoula papillosa	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	papillosa	(Temminck, 1840)	This species does not include Kerivoula lenis (Simmons 2005), although some specimens referred to this species may represent lenis , which appears to be broadly sympatric with papillosa (Vanitharani et al . 2003).	20000000	Kerivoula papillosa	Least Concern		2021	2020-11-28 00:00:00 UTC	3.1	English	Kerivoula papillosa is assessed as Least Concern as the species is widespread, and relatively common, there are no known major threats, and it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	On Buton and Malaysia, this species roosts in groups of 1–14 individuals in small tree hollows in lowland mixed-deciduous dipterocarp forest.	There are no major threat to this species, although deforestation is occurring through many parts of its range.	This species is thought to be reasonably common throughout its range ;and the global population is presumed stable.	Stable	This species is found in Thailand, Cambodia, Viet Nam, Peninsular Malaysia, Sumatra, Java, Sulawesi, Buton in Indonesia, and northern Borneo.		Terrestrial	This species occurs in some protected areas throughout its range including, Gunung Halimun National Park on Java, Danau Sentarum National Park in West Kalimantan (Jeanes and Meijaard 2000), and Gunung Palung National Park (Blundell 1996). Further studies are needed into the taxonomy, distribution, abundance and ecology of this species.	Australasian|Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		papillosa	Temminck	1840	1	Monogr. Mamm.	0.236111	Papillose Woolly Bat	<b> malayana </b>Chasen, 1940.	Indonesia, Java, Bantam (restricted by Tate, 1940).	Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Java, Sulawesi, Borneo.	Not listed.	Least Concern	Does not include lenis ; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis , which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).	Kerivoula papillosa	1005302	23	Papillose Woolly Bat	Papillose Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	papillosa	Temminck	1840	1						"L'ïle de Java, dans le district sauvage de Bantam. Se trouve aussi à Sumatra dans le district de Padang [= The island of Java, in the wild district of Bantam. It is also found in Sumatra in the district of Padang]." Restricted by G. H. H. Tate in 1940 to "Java."			papillosa (Temminck, 1840)|malayana Chasen, 1940	NA	NA				Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Kerivoula_papillosa	0	sciname match	Kerivoula_papillosa	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_papillosa	1005302	23	Papillose Woolly Bat	Papillose Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	papillosa	Temminck	1	Vespertilio papillosus	Temminck, C.J. 1840. Livraison 3. Pp. 141–272 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog/page/140/mode/2up	RMNH.MAM.35753, RMNH.MAM.35754	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35753.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35753.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35754	"L'ïle de Java, dans le district sauvage de Bantam. Se trouve aussi à Sumatra dans le district de Padang [= The island of Java, in the wild district of Bantam. It is also found in Sumatra in the district of Padang]." Restricted by G. H. H. Tate in 1940 to "Java."			NA	NA				Thailand|Laos|Vietnam|Cambodia|Malaysia|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia	LC	0	0	0	Kerivoula_papillosa	0	sciname match	Kerivoula_papillosa	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		papillosa	Temminck	1840	1	Monogr. Mamm.	0.236111	Papillose Woolly Bat	malayana Chasen, 1940.	Indonesia, Java, Bantam (restricted by Tate, 1940).	Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Java, Sulawesi, Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10981/22020906/' target='_blank'>Least Concern</a>	Does not include lenis; see Vanitharani et al. (2003). Appears to be a complex of several distinct species (S. Tsang, pers. comm., 2018). See Hill (1983) and Corbet and Hill (1992) for discussion of subspecies. Reviewed in part by Bates and Harrison (1997). Some specimens referred to this species may represent lenis, which apprears to be broadly sympatric with papillosa (Vanitharani et al., 2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula papillosa; Kerivoula papillosa; Kerivoula papillosa; Kerivoula papillosa; Kerivoula papillosa; Kerivoula papillosa; papillosa; malayana; papillosa; malayana; malayana; papillosa; malayana; Kérivoule mamelonnée; Warzen-Wollfledermaus; Querivoulapiloso de Java; Papillose Bat; Papillose Woolly Bat; Papillose Bat; Papillose Woolly Bat; Papillose Woolly Bat; K. papillosa