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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L506	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina	Kerivoula muscina		[MSW2] Subgenus Kerivoula.; [MSW3] See Flannery (1995a) and Bonaccorso (1998).; [HMW] Kerivoula muscina Tate, 1941 , Lake Daviumbu, 6 miles [= 9- 7 km ] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea ]; altitude approximately 20 meters.” Phylogenetic position of Kerivoula muscina 1s uncertain because it has not been included in any genetic studies. Monotypic.; [batnames2022] See Flannery (1995 a ) and Bonaccorso (1998).; [batnames2023] See Flannery (1995 a ) and Bonaccorso (1998).; [batnames2025_1.7] See Flannery (1995a) and Bonaccorso (1998).														muscina				muscina 	muscina 			muscina Tate, 1941		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Fly River trumpet-eared	SE New Guinea bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula muscina	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.	Tate	1941	Bull. Am. Mus. Nat. Hist., 78:586.	Distribution: Confined to central New Guinea.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Fly River trumpet-eared	C New Guinea bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Tate	1941	Bull. Am. Mus. Nat. Hist., 78:586.	Subgenus Kerivoula.	C New Guinea.	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.		TATE	1941	Rostrum of medium proportions, not narrowed anteriorly. Anterior upper and lower premolars nearly circular in section, but middle upper and lower premolars somewhat elongate. Lingual margins of anterior and middle upper molars somewhat rounded, their shelves of moderate width. Size medium (forearm length, 32-33 mm; condylobasal length, 12-13 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	Distribution: Confined to central New Guinea.	No subspecies.		98	species	K. muscina	TATE	1941	Kerivoula	subgenus	Kerivoula muscina				Rostrum of medium proportions, not narrowed anteriorly. Anterior upper and lower premolars nearly circular in section, but middle upper and lower premolars somewhat elongate. Lingual margins of anterior and middle upper molars somewhat rounded, their shelves of moderate width. Size medium (forearm length, 32-33 mm; condylobasal length, 12- 13 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	No subspecies.		9. K. muscina TATE 1941.	9	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula muscina	Kerivoula		muscina	Tate		1941		Bull. Am. Mus. Nat. Hist.	78		586		Fly River Woolly Bat	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.	C New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		See Flannery (1995a) and Bonaccorso (1998).	4C3D87E8FF766AC9FA7694261BDAB3CD	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	901	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF766AC9FA7694261BDAB3CD.xml	Kerivoula muscina	Vespertilionidae	Kerivoula	muscina	Tate	1941	Kérivoule de Fly River @fr | Fly-River Wollfledermaus @de | Querivoula de Fly River @es | Fly River Trumpet-eared Bat @en	Kerivoula muscina Tate, 1941 , Lake Daviumbu, 6 miles [= 9- 7 km ] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea ]; altitude approximately 20 meters.” Phylogenetic position of Kerivoula muscina 1s uncertain because it has not been included in any genetic studies. Monotypic.	Papua New Guinea lowlands, although it is likely present on Papua Province side of New Guinea as well.	Head—body 35-43 mm , tail 327- 44 mm , ear 13-14 mm , hindfoot 7-8 mm , forearm 32-36 mm ; weight 4-5- 4 g . Fur is long and woolly. Dorsal pelage is pale buffy brown (hairs with pale or medium gray brown bases and pale yellowish brown tips); venteris typically pale gray-brown. Pair of orange, subcutaneous, fatty nodes occurs on forehead between eyes. Membranes and ears are blackish brown, and ears are covered in glandular bumps. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavity just below tips on posterior borders; tragus is narrow and tall, with virtually straight anterior margin except for very slight convexity near tip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and sparse fringe of hairs occurs on posterior margin of uropatagium. Skull is similar to that of Hardwicke’s Woolly Bat (K. hardwick), but I? is narrow, and entire dentition is lighter built.	Primarily lowland rainforests to hill forests and some disturbed habitats from sea level up to elevations of ¢. 1600 m .	No information.	A lactating female and two male Fly River Woolly Bats were captured roosting together in April.	The Fly River Woolly Bat roosts in tree hollows and under large leaves.	The Fly River Woolly Bat is known to roost alone or in groups of up to three individuals.	Classified as Least Concern on The IUCN Red List. The Fly River Woolly Bat is difficult to survey but is widespread and likely common throughout its distribution. Habitat destruction from logging and agriculture are major threats, especially because it is usually found in pristine habitats in lowland areas.	Aplin & Armstrong (2017b) | Bonaccorso (1998) | Flannery (1995b) | Flannery & Seri (1990) | Tate (1941e)	https://zenodo.org/record/6398476/files/figure.png	314. Fly River Woolly Bat Kerivoula muscina French: Kérivoule de Fly River / German: Fly-RiverWollfledermaus / Spanish: Querivoula de Fly River Other common names: Fly River Trumpet-eared Bat Taxonomy. Kerivoula muscina Tate, 1941 , Lake Daviumbu, 6 miles [= 9- 7 km ] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea ]; altitude approximately 20 meters.” Phylogenetic position of Kerivoula muscina 1s uncertain because it has not been included in any genetic studies. Monotypic. Distribution. Papua New Guinea lowlands, although it is likely present on Papua Province side of New Guinea as well. Descriptive notes. Head—body 35-43 mm , tail 327- 44 mm , ear 13-14 mm , hindfoot 7-8 mm , forearm 32-36 mm ; weight 4-5- 4 g . Fur is long and woolly. Dorsal pelage is pale buffy brown (hairs with pale or medium gray brown bases and pale yellowish brown tips); venteris typically pale gray-brown. Pair of orange, subcutaneous, fatty nodes occurs on forehead between eyes. Membranes and ears are blackish brown, and ears are covered in glandular bumps. Ears are large and virtually naked, with convex anterior margins, rounded tips, and concavity just below tips on posterior borders; tragus is narrow and tall, with virtually straight anterior margin except for very slight convexity near tip, and has concave posterior margin with small hooked basal lobe. Wings are attached at base of outertoes, and sparse fringe of hairs occurs on posterior margin of uropatagium. Skull is similar to that of Hardwicke’s Woolly Bat (K. hardwick), but I? is narrow, and entire dentition is lighter built. Habitat. Primarily lowland rainforests to hill forests and some disturbed habitats from sea level up to elevations of ¢. 1600 m . Food and Feeding. No information. Breeding. A lactating female and two male Fly River Woolly Bats were captured roosting together in April. Activity patterns. The Fly River Woolly Bat roosts in tree hollows and under large leaves. Movements, Home range and Social organization. The Fly River Woolly Bat is known to roost alone or in groups of up to three individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Fly River Woolly Bat is difficult to survey but is widespread and likely common throughout its distribution. Habitat destruction from logging and agriculture are major threats, especially because it is usually found in pristine habitats in lowland areas. Bibliography. Aplin & Armstrong (2017b), Bonaccorso (1998), Flannery (1995b), Flannery & Seri (1990), Tate (1941e).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula muscina	Kerivoula		muscina	Tate	1941	0	Bull. Am. Mus. Nat. Hist.	87:46:00	Fly River Woolly Bat	None.	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.	C New Guinea.	Not listed.	Least Concern	See Flannery (1995 a ) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula muscina	23	Fly River Woolly Bat	Fly River Trumpet-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	muscina	Tate	1941	0	Kerivoula_muscina	Tate, G. H. H. (1941). Results of the Archbold Expeditions. No. 40. Notes on Vespertilionid bats. Bulletin of the American Museum of Natural History, 78, 586.	http://digitallibrary.amnh.org/bitstream/handle/2246/1782//v2/dspace/ingest/pdfSource/bul/B078a09.pdf?sequence=1&isAllowed=y	AMNH 105315		Lake Daviumbu, 6 miles [= 9.7 km] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea]; altitude approximately 20 meters."			muscina Tate, 1941	NA	NA	Papua New Guinea|Indonesia?	Oceania	Australasia/Oceania	LC	0	0	0	Kerivoula_muscina	0	sciname match	Kerivoula_muscina	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10979	Kerivoula muscina	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	muscina	Tate, 1941		200000000	Kerivoula muscina	Least Concern		2021	2016-07-31 00:00:00 UTC	3.1	English	<p>This species is listed as Least Concern because, despite being difficult to survey, it has been detected in recent times at multiple widely separated sites, is known to inhabit a range of habitat types that are still more or less intact over large parts of Papua New Guinea, and because threats to these habitats are insufficient to suggest that it will enter a threatened category in the foreseeable future.</p>	<p>Most capture records come from areas of lowland rainforest to hill forest habitat but there is one record from lower montane rainforest. All species of Kerivoula show a strong preference for intact closed forest habitats and a low tolerance for habitat degradation and fragmentation. Until shown otherwise, it should be assumed that populations of K. muscina will persist only in relatively intact forest habitat.</p> <p>Nothing is known of the feeding ecology or reproductive biology of the species but it is probably similar to other species of Kerivoula . Its morphology suggests that it is a slow-flying ‘foliage gleaner’, adapted to foraging in dense vegetation. The only information on roost location comes from discovery of a group of three K. muscina inside a curled banana leaf (Flannery and Seri 1990). Other species of Kerivoula have been observed to roost singly or in small groups in tree hollows, epiphytes or abandoned bird nests.</p>	<p>The major threat to this species is forest degradation, clearance and conversion to plantations or gardens. A very high proportion of the remaining habitat for this species in Papua New Guinea is covered by forestry concession leases (Shearman et al. 2009) and there is active forestry activity in many parts of its range. Despite these developments, there are substantial areas of intact habitat that are either too dissected or steep to be a priority for forestry or plantation projects. Should forest loss continue at the present rate, the population of K. muscina is likely to become increasing fragmented, and this should be monitored and taken into account in future reassessments. </p>	<p>This species is difficult to survey and it is not possible to estimate of population size or local population densities with any reliability. It is known from relatively few specimens collected from fewer than 10 sites. Most early captures were probably made by hand (from roosts in foliage) but more recently small numbers of individuals have been captured in harp traps.</p> <p>The high frequency echolocation call of Kerivoula muscina was recorded only recently (K.P. Aplin and K.N. Armstrong, unpublished). Like all other species of Kerivoula , the call of K. muscina has relatively low energy and the species is unlikely to be detected unless it passes very close to an acoustic recorder. The call type of K. muscina is also very similar in frequency and structure to that of Murina florium , hence unambiguous allocation of a recorded call to one or other of the rarely captured species may not be possible. Few recordings of Kerivoula/Murina were made even in the immediate vicinity of captures of K. muscina .</p>	Unknown	<p></p><p> </p><p>This species is endemic to the main island of New Guinea where it has been recorded from several widely distributed sites. All known occurrences are in Papua New Guinea but it is very likely present in parts of Papua Province, Indonesia, although this requires confirmation. It ranges from sea level to 1,600 m Asl, but is probably predominantly a lowland species.</p> <p></p>		Terrestrial	<p>It has been recorded from Varirata National Park, Central Province in Papua New Guinea (Flannery 1995). </p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		muscina	Tate	1941	0	Bull. Am. Mus. Nat. Hist.	87:46:00	Fly River Woolly Bat	None.	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.	C New Guinea.	Not listed.	Least Concern	See Flannery (1995 a ) and Bonaccorso (1998).	Kerivoula muscina	1005300	23	Fly River Woolly Bat	Fly River Trumpet-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	muscina	Tate	1941	0	Kerivoula_muscina	Tate, G. H. H. (1941). Results of the Archbold Expeditions. No. 40. Notes on Vespertilionid bats. Bulletin of the American Museum of Natural History, 78, 586.	http://digitallibrary.amnh.org/bitstream/handle/2246/1782//v2/dspace/ingest/pdfSource/bul/B078a09.pdf?sequence=1&isAllowed=y	AMNH 105315		Lake Daviumbu, 6 miles [= 9.7 km] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea]; altitude approximately 20 meters."			muscina Tate, 1941	NA	NA				Papua New Guinea|Indonesia?	Oceania	Australasia/Oceania	LC	0	0	0	Kerivoula_muscina	0	sciname match	Kerivoula_muscina	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_muscina	1005300	23	Fly River Woolly Bat	Fly River Trumpet-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	muscina	Tate	0	Kerivoula muscina	Tate, G.H.H. 1941-12-31. Notes on vespertilionid bats of the subfamilies Miniopterinae, Murininae, Kerivoulinae, and Nyctophilinae. Bulletin of the American Museum of Natural History 78(9):567-597.	https://hdl.handle.net/2246/1782	AMNH M-105315	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-105315	Lake Daviumbu, 6 miles [= 9.7 km] above mouth of Strickland River, middle Fly River, Western Division [= Province], Papua [New Guinea]; altitude approximately 20 meters."			NA	NA				Papua New Guinea|Indonesia?	Oceania (Continent)	Australasia	LC	0	0	0	Kerivoula_muscina	0	sciname match	Kerivoula_muscina	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		muscina	Tate	1941	0	Bull. Am. Mus. Nat. Hist.	87:46:00	Fly River Woolly Bat	None.	Papua New Guinea, Western Province, Lake Daviumbu, ca. 20 m.	C New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10979/209536068/' target='_blank'>Least Concern</a>	See Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula muscina; Kerivoula muscina; Kerivoula muscina; Kerivoula muscina; Kerivoula muscina; Kerivoula muscina; muscina; Kérivoule de Fly River; Fly-River Wollfledermaus; Querivoula de Fly River; Fly River Trumpet-eared Bat; Fly River Woolly Bat; Fly River Trumpet-eared Bat; Fly River Woolly Bat; Fly River Woolly Bat; K. muscina