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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L501	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis	Kerivoula krauensis		[HMW] Kerivoula krauensis Francis, Kingston & Zubaid, 2007 , “Kuala Lompat, Krau Wildlife Reserve, Pahang , peninsular Malaysia ( 3°43’N , 102°10’E ).” See K. hardwicku. Kerivoula krauensis might be closely related to K. hardwickii or K. minuta . Monotypic.; [MDD2022] recently described; [IUCN] This is a species of Kerivoula described by Francis et al. (2007) from Peninsular Malaysia. It is similar in size and form to Kerivoula hardwickii Miller 1898 or K. intermedia Hill and Francis 1984, but is distinguished by its distinctive colouration as well as several characters of dentition and skull shape (Francis et al. 2007).; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														krauensis	This is a species of Kerivoula described by Francis et al. (2007) from Peninsular Malaysia. It is similar in size and form to Kerivoula hardwickii Miller 1898 or K. intermedia Hill and Francis 1984, but is distinguished by its distinctive colouration as well as several characters of dentition and skull shape (Francis et al. 2007).			krauensis 	krauensis 			krauensis C. M. Francis, Kingston, & Zubaid, 2007						N/A																																								NA																											4C3D87E8FF716ACEFA55945E17E6B279	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	898	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF716ACEFA55945E17E6B279.xml	Kerivoula krauensis	Vespertilionidae	Kerivoula	krauensis	Francis, Kingston & Zubaid	2007	Kérivoule de Krau @fr | Krau-Wollfledermaus @de | Querivoula de Krau @es	Kerivoula krauensis Francis, Kingston & Zubaid, 2007 , “Kuala Lompat, Krau Wildlife Reserve, Pahang , peninsular Malaysia ( 3°43’N , 102°10’E ).” See K. hardwicku. Kerivoula krauensis might be closely related to K. hardwickii or K. minuta . Monotypic.	Scattered localities in extreme S Thailand , Peninsular Malaysia , C & S Sumatra, and WC & EC Borneo.	Head—body 38-40 mm , tail 34-43 mm , ear 12-15 mm , hindfoot 7- 5 mm (one specimen), forearm 28-34 mm ; weight 2-4- 5 g . Fur is long and woolly. Dorsal pelage is shiny gold brown to light buffy brown (hairs dark brown with golden brown to duller buffy-brown tips); venter is whitish buff (hairs dark gray basally with grayish white to grayish brown tips). Ears are moderately short and broadly funnel-shaped, with concavity on posterior margin near tips; tragusis tall and pointed, curving slightly outward. Ears are pale pinkish gray and slightly darker at edges, and face, nose, and lips are pinkish to purplish brown, with yellowish gland on either side of muzzle. Membranes are dark brown, and wing attaches on foot just below base offirst toe. Uropatagium is sparsely covered dorsally in long, dark brown, golden-tipped hairs. Skull is generally similar to that of Hardwicke’s Woolly Bat (K. hardwickit) but slightly smaller, with more inflated braincase, shorter rostrum, shorter canines, and more rounded upper premolars; braincase is large and globular; C! is narrow and moderately tall; and I* and I’ are unicuspid, I* being small and short at less than one-half the height of I*.	Understories of lowland dipterocarp forests (Malay Peninsula), peat swamp forests and lowland dipterocarp forests (Borneo), and montane logged forests ( Sumatra ), typically close to sea level but up to elevations of ¢. 1600 m ( Sumatra ).	No information.	In central Peninsular Malaysia , female Krau Woolly Bats in breeding condition have been caught mostly in February—June (limited sampling in November-December) and two lactating females in August.	In southern Thailand , call shape is steep FM sweep, with start frequencies of 205-241 kHz (mean 224-8 kHz), end frequencies of 44-6 kHz (51-3 kHz), and peak frequencies of 136-8-166-8 kHz (146-7 kHz). In Peninsular Malaysia , average start frequency was 174 kHz, end frequency was 50 kHz, and duration was 2-4 milliseconds, and in Borneo, a single individual had start frequency of 210 kHz, end frequency of 50 kHz, peak frequency of 137 kHz, and duration of 2-3 milliseconds.	No information.	Classified as Data Deficient on The IUCN Red List. The Krau Woolly Bat is currently known from only seven localities spread over a wide area. It appears to be rare based on low capture rates butlikely has a more extensive distribution.	Chiozza (2008d) | Douangboubpha, Bumrungsri, Satasook et al. (2016) | Douangboubpha, Bumrungsri, Soisook etal. (2014) | Francis et al. (2007) | Huang, J.C.C. et al. (2014) | Struebig et al. (2017)	https://zenodo.org/record/6398456/files/figure.png	304. Krau Woolly Bat Kerivoula krauensis French: Kérivoule de Krau / German: Krau-Wollfledermaus / Spanish: Querivoula de Krau Taxonomy. Kerivoula krauensis Francis, Kingston & Zubaid, 2007 , “Kuala Lompat, Krau Wildlife Reserve, Pahang , peninsular Malaysia ( 3°43’N , 102°10’E ).” See K. hardwicku. Kerivoula krauensis might be closely related to K. hardwickii or K. minuta . Monotypic. Distribution. Scattered localities in extreme S Thailand , Peninsular Malaysia , C & S Sumatra, and WC & EC Borneo. Descriptive notes. Head—body 38-40 mm , tail 34-43 mm , ear 12-15 mm , hindfoot 7- 5 mm (one specimen), forearm 28-34 mm ; weight 2-4- 5 g . Fur is long and woolly. Dorsal pelage is shiny gold brown to light buffy brown (hairs dark brown with golden brown to duller buffy-brown tips); venter is whitish buff (hairs dark gray basally with grayish white to grayish brown tips). Ears are moderately short and broadly funnel-shaped, with concavity on posterior margin near tips; tragusis tall and pointed, curving slightly outward. Ears are pale pinkish gray and slightly darker at edges, and face, nose, and lips are pinkish to purplish brown, with yellowish gland on either side of muzzle. Membranes are dark brown, and wing attaches on foot just below base offirst toe. Uropatagium is sparsely covered dorsally in long, dark brown, golden-tipped hairs. Skull is generally similar to that of Hardwicke’s Woolly Bat (K. hardwickit) but slightly smaller, with more inflated braincase, shorter rostrum, shorter canines, and more rounded upper premolars; braincase is large and globular; C! is narrow and moderately tall; and I* and I’ are unicuspid, I* being small and short at less than one-half the height of I*. Habitat. Understories of lowland dipterocarp forests (Malay Peninsula), peat swamp forests and lowland dipterocarp forests (Borneo), and montane logged forests ( Sumatra ), typically close to sea level but up to elevations of ¢. 1600 m ( Sumatra ). Food and Feeding. No information. Breeding. In central Peninsular Malaysia , female Krau Woolly Bats in breeding condition have been caught mostly in February—June (limited sampling in November-December) and two lactating females in August. Activity patterns. In southern Thailand , call shape is steep FM sweep, with start frequencies of 205-241 kHz (mean 224-8 kHz), end frequencies of 44-6 kHz (51-3 kHz), and peak frequencies of 136-8-166-8 kHz (146-7 kHz). In Peninsular Malaysia , average start frequency was 174 kHz, end frequency was 50 kHz, and duration was 2-4 milliseconds, and in Borneo, a single individual had start frequency of 210 kHz, end frequency of 50 kHz, peak frequency of 137 kHz, and duration of 2-3 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Krau Woolly Bat is currently known from only seven localities spread over a wide area. It appears to be rare based on low capture rates butlikely has a more extensive distribution. Bibliography. Chiozza (2008d), Douangboubpha, Bumrungsri, Satasook et al. (2016), Douangboubpha, Bumrungsri, Soisook etal. (2014), Francis et al. (2007), Huang, J.C.C. et al. (2014), Struebig et al. (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula krauensis	Kerivoula		krauensis	Francis, Kingston, & Zubaid	2007	0	Acta Chiropterol.	9(1): 3	Krau Woolly Bat	None.	peninsular Malaysia , Pahang, Kuala Lompat, Krau Wildlife Reserve,	Krau Wildlife Reserve, peninsular Malaysia; Bala Forest, S. Thailand.	Not listed.	Near Threatened		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula krauensis	23	Krau Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	krauensis	Francis, Kingston, & Zubaid	2007	0	Kerivoula_krauensis	Francis, C. M., Kingston, T., & Zubaid, A. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica, 9(1), 3.	https://bioone.org/journals/acta-chiropterologica/volume-9/issue-1/1733-5329(2007)9%5B1%3AANSOKC%5D2.0.CO%3B2/A-new-species-of-Kerivoula-Chiroptera--Vespertilionidae-from-peninsular/10.3161/1733-5329(2007)9[1:ANSOKC]2.0.CO;2.full	BM 1999.294		"Kuala Lompat, Krau Wildlife Reserve, Pahang, peninsular Malaysia (3°43'N, 102°10'E)."	3.7167	102.17	krauensis Francis, Kingston, & Zubaid, 2007	recently described	Francis, C. M., Kingston, T., & Zubaid, A. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica, 9(1), 1-12.	Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Kerivoula_krauensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136572	Kerivoula krauensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	krauensis	Francis, Kingston &; Zubaid, 2007	This is a species of Kerivoula described by Francis et al. (2007) from Peninsular Malaysia. It is similar in size and form to Kerivoula hardwickii Miller 1898 or K. intermedia Hill and Francis 1984, but is distinguished by its distinctive colouration as well as several characters of dentition and skull shape (Francis et al. 2007).	20000000	Kerivoula krauensis	Near Threatened	A2c	2020	2015-12-21 00:00:00 UTC	3.1	English	<p>This bat is listed as Near Threatened (nearly meets criterion A2c) because it is dependent upon primary forest, and there is significant past and continuing deforestation going on within its range such that its global populations suspected to have declined at a rate of 25-30% over the past 15 years (three generations; generation length = 5 years; Pacifici et al. 2013); it is close to qualifying for Vulnerable under criterion A2c. The threats are not resolved and the population decline is projected to continue.</p>	<p>All known examples of this bat have been caught in the understorey of mature lowland rainforest. The type specimen was caught over small pools in the stream bed of a seasonal stream in relatively undisturbed forest about a kilometre from the clearing at Kuala Lompat. Other individuals were caught along forest understorey trails, showing no particular association with streams or swamps. Nothing is known of the species’ roosting ecology, although, like other Kerivoula it has not been found to roost in caves in the area (Francis et al . 2007). Recent studies in Indomalayan region showed that this species also was recorded in logged lowland dipterocarp forest in East Kalimantan, and in peat swamp forest in Brunei Darussalam. Although mainly reported in lowland forest, one individual was captured over a stream in montane area (ca 1,600 m asl) in logged-over forest in Sumatra.</p>	<p>Deforestation for agricultural development and forest fragmentation are considered to be major threats. ;All records of the species are from locations in tall rainforest, where the species is reported to be very rare. There are no records from highly disturbed forest (e.g., intensive logging, forest fires), and notably the species was not captured in an intensive survey of 27 forest fragments around the type locality reserve in peninsular Malaysia (Struebig ;et al . 2008). The species may be able to withstand a moderate logging activities but highly disturbed forests are likely to be poor habitat (Struebig ;et al. 2016).</p><p> </p>	The species is dependent on mature lowland forest and while the abundance and population size of this species are not well known, the global population is suspected to have declined and is continuing to decline due to extensive forest loss in the region.	Decreasing	<p>The type locality for this species is in Peninsular Malaysia: Kuala Lompat, Krau Wildlife Reserve, Pahang Province (3°43’N, 102°10’E). Recent studies extend the range of this species into six new localities throughout Indomalaya: Sekayu Recreational Forest, Terengganu, Peninsular Malaysia (4°43’N, 102°57’E), Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand (5°48’N, 101°49’E), Napal Melintang, Jambi Province, Sumatra (1°23’N, 106°53’E), Way Canguk, Bukit Barisan Selatan National Park, Lampung Province, Sumatra (5°39’N, 104°24’E), Sungai Lesan Protection Forest, Kelai Distric of Berau, East Kalimantan (1°36’N, 117°10’E) and Badas peat swamp forest, Brunei (4°35’N, 114°23’E). This species is recorded from scattered localities on the Malay-Thai peninsula, Borneo and Sumatra. In addition to the type locality at Kuala Lompat, Krau Wildlife Reserve, Pahang state, Malaysia (Francis et al . 2007), it has been reported from two other locations on the peninsula: in <a name="OLE_LINK1">Hulu Terengganu Tambahan </a>Forest Reserve, Terengganu state, Malaysia (Struebig et al . 2016), and Hala-Bala Wildlife Sanctuary in Narathiwat province, Thailand (Douangboubpha et al . 2014). In Borneo, it is known from the Badas peat swamps in Brunei, and Sungai Lesan in East Kalimantan (Struebig et al . 2016). In Sumatra, there are records from logged montane forest at Napal Melintang, Jambi (Struebig et al . 2016), and Bukit Barisan Selatan National Park, Lampung (Huang et al . 2014). It can be expected to also occur in other parts of these regions at very low densities, but this has not yet been confirmed.</p>		Terrestrial	This species has been collected from Krau Wildlife Reserve. Further studies are needed into the taxonomy, distribution, abundance, natural history and threats to this species. ;Extensive forest surveys in Thailand, Indonesia, Malaysia and Brunei over the last 10 years has extent the occurrence of this species to six additional sites, but with only one or two individuals captured at each site.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		krauensis	Francis, Kingston, & Zubaid	2007	0	Acta Chiropterol.	9(1): 3	Krau Woolly Bat	None.	peninsular Malaysia , Pahang, Kuala Lompat, Krau Wildlife Reserve,	Krau Wildlife Reserve, peninsular Malaysia; Bala Forest, S. Thailand.	Not listed.	Near Threatened		Kerivoula krauensis	1005295	23	Krau Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	krauensis	Francis, Kingston, & Zubaid	2007	0	Kerivoula_krauensis	Francis, C. M., Kingston, T., & Zubaid, A. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica, 9(1), 3.	https://bioone.org/journals/acta-chiropterologica/volume-9/issue-1/1733-5329(2007)9%5B1%3AANSOKC%5D2.0.CO%3B2/A-new-species-of-Kerivoula-Chiroptera--Vespertilionidae-from-peninsular/10.3161/1733-5329(2007)9[1:ANSOKC]2.0.CO;2.full	BM 1999.294		"Kuala Lompat, Krau Wildlife Reserve, Pahang, peninsular Malaysia (3°43'N, 102°10'E)."	3.716667	102.1667	krauensis Francis, Kingston, & Zubaid, 2007	recently described	Francis, C. M., Kingston, T., & Zubaid, A. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica, 9(1), 1-12.				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Kerivoula_krauensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_krauensis	1005295	23	Krau Woolly Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	krauensis	C. M. Francis, Kingston, & Zubaid	0	Kerivoula krauensis	Francis, C.M., Kingston, T. and Zubaid, A. 2007-04. A new species of _Kerivoula_ (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica 9(1):1-12.	https://doi.org/10.3161/1733-5329(2007)9[1:ANSOKC]2.0.CO;2	BMNH:Mamm:1999.294	holotype	https://data.nhm.ac.uk/object/60631591-63ef-44de-9d9f-7c9eb0a0eb2c	"Kuala Lompat, Krau Wildlife Reserve, Pahang, peninsular Malaysia (3°43'N, 102°10'E)."	3.716667	102.1667	recently described	Francis, C. M., Kingston, T., & Zubaid, A. (2007). A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia. Acta Chiropterologica, 9(1), 1-12.				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Kerivoula_krauensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		krauensis	Francis, Kingston & Zubaid	2007	0	Acta Chiropterol.	9(1): 3	Krau Woolly Bat	None.	peninsular Malaysia , Pahang, Kuala Lompat, Krau Wildlife Reserve,	Krau Wildlife Reserve, peninsular Malaysia; Bala Forest, S. Thailand.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136572/21992300/' target='_blank'>Near Threatened</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula krauensis; Kerivoula krauensis; Kerivoula krauensis; Kerivoula krauensis; Kerivoula krauensis; krauensis; Kérivoule de Krau; Krau-Wollfledermaus; Querivoula de Krau; Krau Woolly Bat; Krau Woolly Bat; K. krauensis