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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L498	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula hardwickei	Kerivoula hardwickei	Kerivoula hardwickii	Kerivoula hardwickei	Kerivoula hardwickei	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii	Kerivoula hardwickii		[MSW2] Subgenus Kerivoula. Does not include flora; see Hill and Rozendaal (1989).; [MSW3] Does not include flora; see Hill and Rozendaal (1989). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001b) have used the spelling hardwickii.; [HMW] Vespertilio hardwickii Horsfield, 1824 , Java , Indonesia . Kerwoula hardwickii previously included populations now attributed to K. depressa , K. malpasi, K. crypta , K. furva , and K. dongduongana from throughout South Asia and South-east Asia. These species, along with K. kachinensis , K. krauensis , K. titania , and K. minuta , comprise the hardwickii species group, with K. titania occupying the most basal position in the clade. Genetic data places K. hardwickuisister to K.© ahi Genetic and morphological data support recognition of K. depressa , K. furva , and K. dongduongana as distinct species, and limited biogeographical data support recognition of K. erypta and K. malpasi (although they might be conspecific). Distribution of K. hardwickii is ends in South-east Asia at ¢.16” N, which leaves it allopatric with K. furva . Additional studies are needed to further differentiate species in the hardwickii group with genetic and morphological data. Monotypic.; [batnames2022] Does not include flora ; see Hill and Rozendaal (1989). Does not include depressa ; see Kuo et al. (2017). The subspecies crypta . and malpasi may represent one or two species, but additional study is needed as they have not been included in any recent morphological or molecular studies; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b).Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) andSinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993;Sinha, 1999; Hendrichsen et al., 2001b) have used the spelling hardwickii.; [MDD2022] previously included K. crypta, K. malpasi, and K. depressa; [IUCN] This is the new taxonomic concept for Kerivoula hardwickii . Kerivoula depressa Miller, 1905 now considered to represent a separate species, whereas K. crypta Wroughton and Ryley, 1913 and K. malpasi Phillips, 1931, formerly subsumed as subspecies of K. hardwickii , should also be elevated to species rank (Tu et al. 2018). Specimens previously referred to K. hardwickii s.l. with flattened skull found from Indo-Burma and Himalayan regions are now considered to be representatives of a complex of at least three morphologically cryptic species broadly found in sympatry: K. depressa , K. furva Kuo et al. 2017 and K. dongduongana Tu et al. 2018 (see Kuo et al. 2017, Tu et al. 2018). Although specimens currently referred to K. hardwickii from the Philippines may represent two or more cryptic species (Heaney et al. 2010), they are treated as K. hardwickii until further taxonomic studies are completed. In the new concept K. hardwickii s. str., characterized with highly domed skulls, are distributed from the Sunda Islands northward to the Philippines, south Thailand and south Indochina.; [batnames2023] Does not include flora ; see Hill and Rozendaal (1989). Does not include depressa ; see Kuo et al. (2017). Does not include crypta and malpasi ; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001) have used the spelling hardwickii .; [MDD2023] previously included K. crypta, K. malpasi, and K. depressa; [MDD2025_2.0] previously included K. crypta, K. malpasi, and K. depressa; [batnames2025_1.7] Does not include flora; see Hill and Rozendaal (1989). Does not include depressa; see Kuo et al. (2017). Does not include crypta and malpasi; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001) have used the spelling hardwickii.; [MDD2025_2.2] previously included K. crypta, K. malpasi, and K. depressa				flora		crypta, depressa, engana, fusca, malpasi.	malpasi, crypta, depressa, engana, flora, hardwickei		crypta, depressa, engana, fusca, malpasi			hardwickii	hardwickii - crypta, engana, fusca, malpasi	hardwickii, fusca, engana	This is the new taxonomic concept for Kerivoula hardwickii . Kerivoula depressa Miller, 1905 now considered to represent a separate species, whereas K. crypta Wroughton and Ryley, 1913 and K. malpasi Phillips, 1931, formerly subsumed as subspecies of K. hardwickii , should also be elevated to species rank (Tu et al. 2018). Specimens previously referred to K. hardwickii s.l. with flattened skull found from Indo-Burma and Himalayan regions are now considered to be representatives of a complex of at least three morphologically cryptic species broadly found in sympatry: K. depressa , K. furva Kuo et al. 2017 and K. dongduongana Tu et al. 2018 (see Kuo et al. 2017, Tu et al. 2018). Although specimens currently referred to K. hardwickii from the Philippines may represent two or more cryptic species (Heaney et al. 2010), they are treated as K. hardwickii until further taxonomic studies are completed. In the new concept K. hardwickii s. str., characterized with highly domed skulls, are distributed from the Sunda Islands northward to the Philippines, south Thailand and south Indochina.	hardwickii	hardwickii - engana, fusca	hardwickii, fusca, engana 	hardwickii, hardwichii, fusca, engana, hardwickei	hardwickii	hardwickii - engana, fusca	hardwickii (Horsfield, 1824)|hardwichii (Lesson, 1842) [incorrect subsequent spelling]|fusca Dobson, 1871|hardwickei O. Thomas, 1898 [incorrect subsequent spelling]|engana G. S. Miller, 1906|hardwicki W. W. A. Phillips, 1924 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Hardwicke's forest bat	Sri Lanka, India – S China – Java, Lesser Sundas, Philippines, Celebes	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula hardwickei	Indonesia, Java.	Horsfield	1824	Zool. Res. Java, Part 8:28.	Distribution: Ranging from India and Ceylon to southern China and Indo-China, south and east to the Philippines, Celebes, and the Lesser Sundas.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Hardwick's forest bat	Sri Lanka, India – S China – Java, Bali, ? Sumba I, Lesser Sunda Is, Philippines, Sulawesi, Talaud Is; ref. 4.143	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Horsfield	1824	Zool. Res. Java, Part 8:28.	Subgenus Kerivoula. Does not include flora; see Hill and Rozendaal (1989).	India; Sri Lanka; Burma; Thailand; China; W Malaysia; Borneo; Java, Sumatra, Mentawai Isis, Sulawesi, Bali, Lesser Sundas, Kangean Isl and Talaud Isl (Indonesia); Philippines.	Indonesia, Java.		HORSFIELD	1824	Rostrum short, not narrowed anteriorly. Anterior and middle premolars nearly circular in section. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size medium (forearm length, 30-39 mm; condylobasal length, 11-15 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	Distribution: Ranging from India and Ceylon to southern China and Indo-China, south and east to the Philippines, Celebes, and the Lesser Sundas.	Six subspecies are recognized:	K. h. malpasi (Ceylon), K. h. crypta (southern India), K. h. depressa (northern India to southern China and Indo-China), K. h. engana (Mentawei islands off western Sumatra), K. h. flora (Lesser Sundas), K. h. hardwickei (remainder of range).	99	species	K. hardwickei	HORSFIELD	1824	Kerivoula	subgenus	Kerivoula hardwickei				Rostrum short, not narrowed anteriorly. Anterior and middle premolars nearly circular in section. Lingual margins of anterior and middle upper molars more or less square, their shelves broad. Size medium (forearm length, 30-39 mm; condylobasal length, 11-15 mm). Inner upper incisor bicuspid, but secondary cusp small. Fringe of hair on posterior margin of uropatagium poorly developed.	Six subspecies are recognized:		14. K. hardwickei (HORSFIELD 1824).	14	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula hardwickii	Kerivoula		hardwickii	Horsfield	y	1824		Zool. Res. Java	Part 8		p. 4(unno.) of Vespertilio Temminckii acct		Hardwicke's Woolly Bat	Indonesia, Java.	India and Sri Lanka, Burma, Laos, Cambodia, Vietnam, Thailand, China, W Malaysia, Borneo, Java, Sumatra, Nusa Penida, Mentawai Isls, Sulawesi, Bali, Lesser Sundas, Kangean Isl and Talaud Isl (Indonesia), Philippines.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	crypta Wroughton and Ryley, 1913; depressa Miller, 1906; engana Miller, 1906; fusca Dobson, 1871; malpasi Phillips, 1932.	Does not include flora; see Hill and Rozendaal (1989). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001b) have used the spelling hardwickii.	4C3D87E8FF736ACCFA4F942C179AB7B3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	896	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF736ACCFA4F942C179AB7B3.xml	Kerivoula hardwickii	Vespertilionidae	Kerivoula	hardwickii		1824	Kérivoule de Hardwicke @fr | Hardwicke-Wollfledermaus @de | Querivoula de Hardwicke @es | Common Woolly Bat @en | Hardwicke's Forest Bat @en	Vespertilio hardwickii Horsfield, 1824 , Java , Indonesia . Kerwoula hardwickii previously included populations now attributed to K. depressa , K. malpasi, K. crypta , K. furva , and K. dongduongana from throughout South Asia and South-east Asia. These species, along with K. kachinensis , K. krauensis , K. titania , and K. minuta , comprise the hardwickii species group, with K. titania occupying the most basal position in the clade. Genetic data places K. hardwickuisister to K.© ahi Genetic and morphological data support recognition of K. depressa , K. furva , and K. dongduongana as distinct species, and limited biogeographical data support recognition of K. erypta and K. malpasi (although they might be conspecific). Distribution of K. hardwickii is ends in South-east Asia at ¢.16” N, which leaves it allopatric with K. furva . Additional studies are needed to further differentiate species in the hardwickii group with genetic and morphological data. Monotypic.	S Thailand , S Laos , S Vietnam (including Con Son I), Cambodia , Peninsular Malaysia , Singapore , Sumatra, Mentawai Is (Siberut and Sipora), Borneo, Banggi I, Java, Kangean I, Bali I, Nusa Penida I, Lombok I, Sumba I, Sulawesi, Peleng I, Karakelang I, and Philippines (Lubang, Palawan , Samar, Biliran , Leyte , Bohol , and Mindanao Is) including some offshore Is.	Head-body c. 39-41 mm , tail 35-50 mm , ear 9-15 mm , hindfoot 5-7- 10 mm , forearm 26- 5-36 mm ; weight 2-9- 6 g . Fur is smooth, thick, and silky. Dorsal pelage is generally deep brown (hairs dark-based with pale gray or brown tips); venter is paler and grayer; and overall, pelage is lighter than in the Dark Woolly Bat ( K. furva ). Young are grayer. Membranes are brown and semi-translucent, and other bare parts are brown. Ears are funnel-shaped, and fold extends from base of ear to halfway up posterior margin; tragus is long and pointed, with distinct basal notch, and curves slightly outward at tip. Wings are attached at base of toes, and calcar is well developed and curved. Thickened smooth pad occurs at base of thumbs used for gripping. Skull is broadly similar to that of the Dark Woolly Bat but with proportionally higher braincase; skull is domed unlike in the Flatskulled Woolly Bat ( K. depressa ). Dental formula for all species of Kerivoulais 12/3, C1/1,P 3/3, M 3/3 (x2) = 38. Chromosomal complement has 2n = 26 and FN = 46 or 48 (Borneo).	Various forest habitats, including tropical primary forests; dry forests; hill forests; lowland, montane, and ridge-top mossy forests; and peat bogs and heath forests (Borneo) at elevations of 60-1600 m .	Hardwicke’s Woolly Bats are highly maneuverable but slow fliers and likely forage by aerial hawking near water.	A lactating Hardwicke’s Woolly Bat was caught inJanuary in Peninsular Malaysia .	Hardwicke’s Woolly Bat reportedly roosts in foliage, tree hollows, houses, and pitcher plants. In Borneo, the pitcher plant Nepenthes hemsleyana ( Nepenthaceae ) is somewhat mutualistic/symbiotic with Hardwicke’s Woolly Bat; they gain high-quality roosts and give nutrients to plants by defecating in pitchers. Although this benefits both organisms, it might cause the pitcher to wilt faster, which might be energetically costly for the plant. In southern Thailand , call is weak and steep FM sweep, with average start frequency of 169-6 kHz, end frequency of 90-7 kHz, peak frequency of 118-3 kHz, duration of 3-2 milliseconds, and interpulse interval of 15-6 milliseconds. In another study in southern Thailand , average start frequency was 125-7 kHz, end frequency was 104-5 kHz, peak frequency was 114-8 kHz, duration was 0-6 milliseconds, and interpulse interval was 16-1 milliseconds.	Hardwicke’s Woolly Bat probably roosts alone or in small groups.	Classified as Least Concern on The IUCN Red List. Hardwicke’s Woolly Bats are widespread and relatively common, and they do not seem to face any major threats. They might be locally threatened by deforestation from logging and agriculture.	Bates, Struebig et al. (2007) | Douangboubpha et al. (2016) | Esselstyn, Widmann & Heaney (2004) | Hasan & Abdullah (2011) | Heaney et al. (2016) | Hughes et al. (2011) | Khan (2008) | Khan et al. (2010) | Kuo Haochih et al. (2017) | Leong & Lim (2009) | Lim et al. (2015) | Rosell-Ambal, Tabaranza, Heaney, Molur & Srinivasulu (2008) | Schoner et al. (2017) | Tu Vuong Tan et al. (2018) | Yu Wenhua et al. (2018)	https://zenodo.org/record/6398442/files/figure.png	297. Hardwicke’s Woolly Bat Kerivoula hardwickii French: Kérivoule de Hardwicke / German: Hardwicke-Wollfledermaus / Spanish: Querivoula de Hardwicke Other common names: Common Woolly Bat , Hardwicke's Forest Bat Taxonomy. Vespertilio hardwickii Horsfield, 1824 , Java , Indonesia . Kerwoula hardwickii previously included populations now attributed to K. depressa , K. malpasi, K. crypta , K. furva , and K. dongduongana from throughout South Asia and South-east Asia. These species, along with K. kachinensis , K. krauensis , K. titania , and K. minuta , comprise the hardwickii species group, with K. titania occupying the most basal position in the clade. Genetic data places K. hardwickuisister to K.© ahi Genetic and morphological data support recognition of K. depressa , K. furva , and K. dongduongana as distinct species, and limited biogeographical data support recognition of K. erypta and K. malpasi (although they might be conspecific). Distribution of K. hardwickii is ends in South-east Asia at ¢.16” N, which leaves it allopatric with K. furva . Additional studies are needed to further differentiate species in the hardwickii group with genetic and morphological data. Monotypic. Distribution. S Thailand , S Laos , S Vietnam (including Con Son I), Cambodia , Peninsular Malaysia , Singapore , Sumatra, Mentawai Is (Siberut and Sipora), Borneo, Banggi I, Java, Kangean I, Bali I, Nusa Penida I, Lombok I, Sumba I, Sulawesi, Peleng I, Karakelang I, and Philippines (Lubang, Palawan , Samar, Biliran , Leyte , Bohol , and Mindanao Is) including some offshore Is. Descriptive notes. Head-body c. 39-41 mm , tail 35-50 mm , ear 9-15 mm , hindfoot 5-7- 10 mm , forearm 26- 5-36 mm ; weight 2-9- 6 g . Fur is smooth, thick, and silky. Dorsal pelage is generally deep brown (hairs dark-based with pale gray or brown tips); venter is paler and grayer; and overall, pelage is lighter than in the Dark Woolly Bat ( K. furva ). Young are grayer. Membranes are brown and semi-translucent, and other bare parts are brown. Ears are funnel-shaped, and fold extends from base of ear to halfway up posterior margin; tragus is long and pointed, with distinct basal notch, and curves slightly outward at tip. Wings are attached at base of toes, and calcar is well developed and curved. Thickened smooth pad occurs at base of thumbs used for gripping. Skull is broadly similar to that of the Dark Woolly Bat but with proportionally higher braincase; skull is domed unlike in the Flatskulled Woolly Bat ( K. depressa ). Dental formula for all species of Kerivoulais 12/3, C1/1,P 3/3, M 3/3 (x2) = 38. Chromosomal complement has 2n = 26 and FN = 46 or 48 (Borneo). Habitat. Various forest habitats, including tropical primary forests; dry forests; hill forests; lowland, montane, and ridge-top mossy forests; and peat bogs and heath forests (Borneo) at elevations of 60-1600 m . Food and Feeding. Hardwicke’s Woolly Bats are highly maneuverable but slow fliers and likely forage by aerial hawking near water. Breeding. A lactating Hardwicke’s Woolly Bat was caught inJanuary in Peninsular Malaysia . Activity patterns. Hardwicke’s Woolly Bat reportedly roosts in foliage, tree hollows, houses, and pitcher plants. In Borneo, the pitcher plant Nepenthes hemsleyana ( Nepenthaceae ) is somewhat mutualistic/symbiotic with Hardwicke’s Woolly Bat; they gain high-quality roosts and give nutrients to plants by defecating in pitchers. Although this benefits both organisms, it might cause the pitcher to wilt faster, which might be energetically costly for the plant. In southern Thailand , call is weak and steep FM sweep, with average start frequency of 169-6 kHz, end frequency of 90-7 kHz, peak frequency of 118-3 kHz, duration of 3-2 milliseconds, and interpulse interval of 15-6 milliseconds. In another study in southern Thailand , average start frequency was 125-7 kHz, end frequency was 104-5 kHz, peak frequency was 114-8 kHz, duration was 0-6 milliseconds, and interpulse interval was 16-1 milliseconds. Movements, Home range and Social organization. Hardwicke’s Woolly Bat probably roosts alone or in small groups. Status and Conservation. Classified as Least Concern on The IUCN Red List. Hardwicke’s Woolly Bats are widespread and relatively common, and they do not seem to face any major threats. They might be locally threatened by deforestation from logging and agriculture. Bibliography. Bates, Struebig et al. (2007), Douangboubpha et al. (2016), Esselstyn, Widmann & Heaney (2004), Hasan & Abdullah (2011), Heaney et al. (2016), Hughes et al. (2011), Khan (2008), Khan et al. (2010), Kuo Haochih et al. (2017), Leong & Lim (2009), Lim et al. (2015), Rosell-Ambal, Tabaranza, Heaney, Molur & Srinivasulu (2008), Schoner et al. (2017), Tu Vuong Tan et al. (2018), Yu Wenhua et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula hardwickii	Kerivoula		hardwickii	Horsfield	1824	1	Zool. Res. Java	Part 8: p. 4(unno.) of Vespertilio Temminckii acct	Hardwicke's Woolly Bat	 crypta Wroughton and Ryley, 1913; engana Miller, 1906; fusca Dobson, 1871; malpasi Phillips, 1932.	Indonesia, Java.	India and Sri Lanka, Burma, Laos, Cambodia, Vietnam, Thailand, China, W Malaysia, Borneo, Java, Sumatra, Nusa Penida, Mentawai Isls, Sulawesi, Bali, Lesser Sundas, Kangean Isl and Talaud Isl (Indonesia), Philippines.	Not listed.	Least Concern	Does not include flora ; see Hill and Rozendaal (1989). Does not include depressa ; see Kuo et al. (2017). The subspecies crypta . and malpasi may represent one or two species, but additional study is needed as they have not been included in any recent morphological or molecular studies; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b).Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) andSinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993;Sinha, 1999; Hendrichsen et al., 2001b) have used the spelling hardwickii.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula hardwickii	23	Hardwicke's Woolly Bat	Common Woolly Bat|Hardwicke's Forest Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	hardwickii	Horsfield	1824	1	Vespertilio_hardwickii	Horsfield, T. (1824). Zoological researches in Java, and the neighbouring islands. Kingsbury, Parbury, & Allen, London, Part 8, p. 4 (unnumbered) of Vespertilio Temminckii acct.	https://www.biodiversitylibrary.org/item/97435#page/90/mode/1up	BM 1879.11.21.181		Java, Indonesia.			hardwickii (Horsfield, 1824)|fusca Dobson, 1871|engana G. S. Miller, 1906	previously included K. crypta, K. malpasi, and K. depressa	Tu, V. T., Hassanin, A., Furey, N. M., Son, N. T., & Csorba, G. (2018). Four species in one: multigene analyses reveal phylogenetic patterns within Hardwicke's woolly bat, Kerivoula hardwickii-complex (Chiroptera, Vespertilionidae) in Asia. Hystrix, the Italian Journal of Mammalogy, 29(1), 111-121.	Thailand|Myanmar|Laos|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Kerivoula_hardwickii	0	sciname match	Kerivoula_hardwickii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	200000000	Kerivoula hardwickii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	hardwickii	(Horsfield, 1824)	This is the new taxonomic concept for Kerivoula hardwickii . Kerivoula depressa Miller, 1905 now considered to represent a separate species, whereas K. crypta Wroughton and Ryley, 1913 and K. malpasi Phillips, 1931, formerly subsumed as subspecies of K. hardwickii , should also be elevated to species rank (Tu et al. 2018). Specimens previously referred to K. hardwickii s.l. with flattened skull found from Indo-Burma and Himalayan regions are now considered to be representatives of a complex of at least three morphologically cryptic species broadly found in sympatry: K. depressa , K. furva Kuo et al. 2017 and K. dongduongana Tu et al. 2018 (see Kuo et al. 2017, Tu et al. 2018). Although specimens currently referred to K. hardwickii from the Philippines may represent two or more cryptic species (Heaney et al. 2010), they are treated as K. hardwickii until further taxonomic studies are completed. In the new concept K. hardwickii s. str., characterized with highly domed skulls, are distributed from the Sunda Islands northward to the Philippines, south Thailand and south Indochina.	20000000	Kerivoula hardwickii	Least Concern		2020	2019-04-30 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, has a tolerance of a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species was recorded in the understory of various forest types, including evergreen, deciduous, and disturbed and from low-land to montane (Francis 2008). It was found roosting in hollow trees and furled leaves of various plants (Francis, 2008) and especially Nepenthes ;pitchers (Schöner et al. 2017). The species has a relatively slow but very maneuverable flight and forage in highly cluttered places (Kruskop 2011). On Con Dao island (Vietnam), some young, not fully-grown individuals were recorded during the period between late May and early June indicating that females of species can produce litters in April (Kruskop 2011).	There are no major threats to this species as a whole. However, habitat loss and degradation through logging, development, and fire in the region have been known to reduce foraging and roosting sites of this species.	In Southeast Asia, it is a moderately common species in secondary and primary forests (Heaney et al. 2010). The abundance, population size and trends for this species across its range are not known.	Stable	This species was formally known to occur widely throughout Asia. However, it is now considered to be restricted in Southeast Asia with the northern limit in the mainland being around 18ºN (see Tu et al. 2018). In insular Southeast Asia it has been recorded from the islands of Mentawi [Siberut and Sipura], Engano, Sumatra, Java, Bali, Kangean, Nusa Penida, Lombok, Sumba, Sulawesi, Pelang, Banggi, Karakelang (Indonesia), to the island of Borneo (Brunei, Indonesia and Malaysia) (Corbet and Hill 1992), and to the islands of Biliran, Bohol, Leyte, Lubang, Mindanao (Agusan del Norte, Bukidnon, Davao Oriental, Surigao del Sur, and Zamboanga del Sur provinces), and Palawan (Philippines) (Heaney et al. 2010). In the Philippines, it has been recorded between 60 and 1,600 m asl (Heaney et al. 2010).		Terrestrial	This species has been recorded from numerous protected areas throughout Southeast Asia. Additional research is needed on taxonomy, especially within the Philippines.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		hardwickii	Horsfield	1824	1	Zool. Res. Java	Part 8: p. 4(unno.) of Vespertilio Temminckii acct	Hardwicke's Woolly Bat	 crypta Wroughton and Ryley, 1913; engana Miller, 1906; fusca Dobson, 1871; malpasi Phillips, 1932.	Indonesia, Java	Myanmar, Laos, Cambodia, Vietnam, Thailand, China, W Malaysia, Borneo, Java, Sumatra, Nusa Penida, Mentawai Isls, Sulawesi, Bali, Lesser Sundas, Kangean Isl and Talaud Isl (Indonesia), Philippines	Not listed.	Least Concern	Does not include flora ; see Hill and Rozendaal (1989). Does not include depressa ; see Kuo et al. (2017). Does not include crypta and malpasi ; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001) have used the spelling hardwickii .	Kerivoula hardwickii	1005292	23	Hardwicke's Woolly Bat	Common Woolly Bat|Hardwicke's Forest Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	hardwickii	Horsfield	1824	1	Vespertilio_hardwickii	Horsfield, T. (1824). Zoological researches in Java, and the neighbouring islands. Kingsbury, Parbury, & Allen, London, Part 8, p. 4 (unnumbered) of Vespertilio Temminckii acct.	https://www.biodiversitylibrary.org/item/97435#page/90/mode/1up	BM 1879.11.21.181		Java, Indonesia.			hardwickii (Horsfield, 1824)|fusca Dobson, 1871|engana G. S. Miller, 1906	previously included K. crypta, K. malpasi, and K. depressa	Tu, V. T., Hassanin, A., Furey, N. M., Son, N. T., & Csorba, G. (2018). Four species in one: multigene analyses reveal phylogenetic patterns within Hardwicke's woolly bat, Kerivoula hardwickii-complex (Chiroptera, Vespertilionidae) in Asia. Hystrix, the Italian Journal of Mammalogy, 29(1), 111-121.				Thailand|Myanmar|Laos|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia/Oceania	LC	0	0	0	Kerivoula_hardwickii	0	sciname match	Kerivoula_hardwickii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_hardwickii	1005292	23	Hardwicke's Woolly Bat	Common Woolly Bat|Hardwicke's Forest Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	hardwickii	Horsfield	1	Vespertilio hardwickii	Horsfield, T. 1824-09. No. VIII. _Sciurus bicolor_. _Mus setifer_. _Vespertilio Temminckii_. _Cheiromeles torquatus_. _Prinia familiaris_. _Perdix personata_. _Parra superciliosa_. _Anas arcuata_. in Horsfield, T. 1824. Zoological Researches in Java, and the Neighbouring Islands. Kingsbury, Parbury, & Allen, London, not continuously paginated pp.	https://www.biodiversitylibrary.org/page/31111914	BMNH:Mamm:1879.11.21.181	holotype	https://data.nhm.ac.uk/object/928adc94-0a58-441b-8b28-ba5d9009f2b2	Java, Indonesia.			previously included K. crypta, K. malpasi, and K. depressa	Tu, V. T., Hassanin, A., Furey, N. M., Son, N. T., & Csorba, G. (2018). Four species in one: multigene analyses reveal phylogenetic patterns within Hardwicke's woolly bat, Kerivoula hardwickii-complex (Chiroptera, Vespertilionidae) in Asia. Hystrix, the Italian Journal of Mammalogy, 29(1), 111-121.				Thailand|Myanmar|Laos|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia	LC	0	0	0	Kerivoula_hardwickii	0	sciname match	Kerivoula_hardwickii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		hardwickii	Horsfield	1824	1	Zool. Res. Java	Part 8: p. 4(unno.) of Vespertilio Temminckii acct	Hardwicke's Woolly Bat	crypta Wroughton and Ryley, 1913; engana Miller, 1906; fusca Dobson, 1871; malpasi Phillips, 1932.	Indonesia, Java	Myanmar, Laos, Cambodia, Vietnam, Thailand, China, W Malaysia, Borneo, Java, Sumatra, Nusa Penida, Mentawai Isls, Sulawesi, Bali, Lesser Sundas, Kangean Isl and Talaud Isl (Indonesia), Philippines	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/154195594/21973742/' target='_blank'>Least Concern</a>	Does not include flora; see Hill and Rozendaal (1989). Does not include depressa; see Kuo et al. (2017). Does not include crypta and malpasi; see Tu et al. (2018). Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001). Multiple subspecies have been recognized in the past, but recent studies suggest that these are not justified; see Corbet and Hill (1992) and Sinha (1999). This taxon is sometimes spelled hardwickei or hardwicki but most recent authors (e.g., Corbet and Hill, 1992; Koopman, 1993; Sinha, 1999; Hendrichsen et al., 2001) have used the spelling hardwickii.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula hardwickii; Kerivoula hardwickii; Kerivoula hardwickii; Kerivoula hardwickii; Kerivoula hardwickii; Kerivoula hardwickii; crypta; depressa; engana; fusca; malpasi; crypta; engana; fusca; malpasi; hardwickii; fusca; engana; Kérivoule de Hardwicke; Hardwicke-Wollfledermaus; Querivoula de Hardwicke; Common Woolly Bat; Hardwicke's Forest Bat; Hardwicke's Woolly Bat; Common Woolly Bat; Hardwicke's Forest Bat; Hardwicke's Woolly Bat; Hardwicke's Woolly Bat; K. hardwickii