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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L492	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa	Kerivoula cuprosa		[MSW2] Subgenus Kerivoula.; [HMW] Kerivoula cuprosa Thomas, 1912 , “Bitye, Ja River, S.E. Cameroons [= Cameraon). 2000’ [= 610 m ].” Specimens of K. cuprosa have been confused with K. smithii . Monotypic.														cuprosa				cuprosa	cuprosa			cuprosa O. Thomas, 1912		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Copper woolly bat	S Cameroun, Kenya, ? Zaire	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Kerivoula cuprosa	Cameroon, Ja River, Bitye.	Thomas	1912	Ann. Mag. Nat. Hist., ser. 8, 10:41.	Distribution: Known only from forested regions of Ghana, Cameroon, northern Zaire, and Kenya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Copper woolly bat	Ghana, S Cameroun, Kenya, Zaire	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1912	Ann. Mag. Nat. Hist., ser. 8, 10:41.	Subgenus Kerivoula.	Kenya, N Zaire, S Cameroon.	Cameroon, Ja River, Bitye.		THOMAS	1912	Rostrum of medium proportions. Size medium (forearm length, 32-34 mm; condylobasal length, 12-13 mm). Inner upper incisor bicuspid but relatively short. Fringe of hair on posterior margin of uropatagium poorly developed. Outer lower incisor bicuspid.	Distribution: Known only from forested regions of Ghana, Cameroon, northern Zaire, and Kenya.	No subspecies.		98	species	K. cuprosa	THOMAS	1912	Kerivoula	subgenus	Kerivoula cuprosa				Rostrum of medium proportions. Size medium (forearm length, 32-34 mm; condylobasal length, 12-13 mm). In- ner upper incisor bicuspid but relatively short. Fringe of hair on posterior margin of uropatagium poorly developed. Outer lower incisor bicuspid.	No subspecies.		5. K. cuprosa THOMAS 1912.	5	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Kerivoulinae		Kerivoula cuprosa	Kerivoula		cuprosa	Thomas		1912		Ann. Mag. Nat. Hist., ser. 8	10		41		Copper Woolly Bat	Cameroon, Ja River, Bitye.	N Dem. Rep. Congo, S Cameroon. A record of this species from Kenya was based on a specimen subsequently reidentified as smithii (J. Fahr, pers. comm.)	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).			4C3D87E8FF746ACBFF819DE7193DBDEE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	903	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF746ACBFF819DE7193DBDEE.xml	Kerivoula cuprosa	Vespertilionidae	Kerivoula	cuprosa	Thomas	1912	Kérivoule cuivrée @fr | Kupferfarbige Wollfledermaus @de | Querivoulacobrizo @es | Copper Woolly Bat @en	Kerivoula cuprosa Thomas, 1912 , “Bitye, Ja River, S.E. Cameroons [= Cameraon). 2000’ [= 610 m ].” Specimens of K. cuprosa have been confused with K. smithii . Monotypic.	Known from few scattered localities in Guinea , Liberia , Ivory Coast , Cameroon , and DR Congo ; possibly in Ghana .	Head-body ¢. 34-39 mm , tail 40-44 mm , ear 12-14 mm , hindfoot 5-8-7- 5 mm , forearm 30-33 mm ; weight 4- 4-5 g . Pelage is dense, woolly, and frizzled, with many hairs having hooked tips. Dorsal pelage is dark or chocolatebrown, with golden sheen or singed appearance; hairs are dark brown, with terminal one-third bright rusty brown or chocolate-brown; many hairs have pale brown hooked tips. Mid-dorsal hairs are c. 7-8 mm . Dorsal pelage extends onto surfaces of forearm and tibia. Ventral pelage 1s duller brown; hairs are dark grayish brown, with terminal one-third paler and whitish or cream tips on some hairs. Wings and uropatagium are dark brown. Posterior margin of uropatagium has thinly scattered curled hairs but no comb-like fringe of hooked, bristle-like hairs; proximal one-third of dorsal surface is sparsely covered with hairs. Head is high-domed. Muzzle is long, pointed, and hidden by facial pelage. Ears are widely separated, funnel-shaped, short for a vespertilionid, and brown. Tragus is moderately long and narrow, tapering to sharp point, curved outward, and with its base having small lobule above which there is an emargination. Eyes are small. Skull is small (greatest skull lengths 12-:3-13- 1 mm ) and has high-domed cranium, rostrum is narrow, and frontal region is sharply angular to plane of rostrum as in other species of Kerivoula . Upper incisors are shorter than in Spurrell’s Woolly Bat ( K. phalaena ) and Smith’s Woolly Bat ( K. smithii ); I? is bicuspid, with second cusp one-half the length of main cusp; I? is about one-half to two-thirds the height of I? and unicuspid or weakly bicuspid, with secondary cusp oriented posterointernally; I, is tricuspid; and lower premolars are subequal in height, but P, is broader than P, and P,.	Lowland rainforests, swamp forests, and forest-savanna mosaic at elevations up to 600 m . In Ivory Coast , Coppery Woolly Bats were caught in dense vegetation with harp traps placed 1-5 m aboveground.	The Coppery Woolly Bat foragesin cluttered spaces and seems to be a gleaner. Delicate skull suggests consumption of soft-bodied arthropods.	No information.	Wings are broad, and flight is highly maneuverable. Coppery Woolly Bats might roost among leaves. Echolocation calls are steep, broadband FM calls.	Ectoparasites include the mite Olabidocarpus belsorum (Acari, Chirodiscidae ).	Classified as Data Deficient on The IUCN Red List.	ACR (2017) | Fahr (2013ab)	https://zenodo.org/record/6398494/files/figure.png	321. Coppery Woolly Bat Kerivoula cuprosa French: Kérivoule cuivrée / German: Kupferfarbige Wollfledermaus / Spanish: Querivoula cobrizo Other common names: Copper Woolly Bat Taxonomy. Kerivoula cuprosa Thomas, 1912 , “Bitye, Ja River, S.E. Cameroons [= Cameraon). 2000’ [= 610 m ].” Specimens of K. cuprosa have been confused with K. smithii . Monotypic. Distribution. Known from few scattered localities in Guinea , Liberia , Ivory Coast , Cameroon , and DR Congo ; possibly in Ghana . Descriptive notes. Head-body ¢. 34-39 mm , tail 40-44 mm , ear 12-14 mm , hindfoot 5-8-7- 5 mm , forearm 30-33 mm ; weight 4- 4-5 g . Pelage is dense, woolly, and frizzled, with many hairs having hooked tips. Dorsal pelage is dark or chocolatebrown, with golden sheen or singed appearance; hairs are dark brown, with terminal one-third bright rusty brown or chocolate-brown; many hairs have pale brown hooked tips. Mid-dorsal hairs are c. 7-8 mm . Dorsal pelage extends onto surfaces of forearm and tibia. Ventral pelage 1s duller brown; hairs are dark grayish brown, with terminal one-third paler and whitish or cream tips on some hairs. Wings and uropatagium are dark brown. Posterior margin of uropatagium has thinly scattered curled hairs but no comb-like fringe of hooked, bristle-like hairs; proximal one-third of dorsal surface is sparsely covered with hairs. Head is high-domed. Muzzle is long, pointed, and hidden by facial pelage. Ears are widely separated, funnel-shaped, short for a vespertilionid, and brown. Tragus is moderately long and narrow, tapering to sharp point, curved outward, and with its base having small lobule above which there is an emargination. Eyes are small. Skull is small (greatest skull lengths 12-:3-13- 1 mm ) and has high-domed cranium, rostrum is narrow, and frontal region is sharply angular to plane of rostrum as in other species of Kerivoula . Upper incisors are shorter than in Spurrell’s Woolly Bat ( K. phalaena ) and Smith’s Woolly Bat ( K. smithii ); I? is bicuspid, with second cusp one-half the length of main cusp; I? is about one-half to two-thirds the height of I? and unicuspid or weakly bicuspid, with secondary cusp oriented posterointernally; I, is tricuspid; and lower premolars are subequal in height, but P, is broader than P, and P,. Habitat. Lowland rainforests, swamp forests, and forest-savanna mosaic at elevations up to 600 m . In Ivory Coast , Coppery Woolly Bats were caught in dense vegetation with harp traps placed 1-5 m aboveground. Food and Feeding. The Coppery Woolly Bat foragesin cluttered spaces and seems to be a gleaner. Delicate skull suggests consumption of soft-bodied arthropods. Breeding. No information. Activity patterns. Wings are broad, and flight is highly maneuverable. Coppery Woolly Bats might roost among leaves. Echolocation calls are steep, broadband FM calls. Movements, Home range and Social organization. Ectoparasites include the mite Olabidocarpus belsorum (Acari, Chirodiscidae ). Status and Conservation. Classified as Data Deficient on The IUCN Red List. Bibliography. ACR (2017), Fahr (2013ab).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Kerivoula cuprosa	Kerivoula		cuprosa	Thomas	1912	0	Ann. Mag. Nat. Hist.	ser. 8, 10: 41	Copper Woolly Bat	None.	Cameroon, Ja River, Bitye.	N Dem. Rep. Congo, S Cameroon. A record of this species from Kenya was based on a specimen subsequently reidentified as smithii (J. Fahr, pers. comm.)	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Kerivoula cuprosa	23	Coppery Woolly Bat	Copper Woolly Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	KERIVOULINAE	NA	Kerivoula	NA	cuprosa	O. Thomas	1912	0	Kerivoula_cuprosa	Thomas, O. (1912). Mammals from the Ja River, Cameroons. Annals and Magazine of Natural History, ser. 8, 10, 41.	https://www.biodiversitylibrary.org/item/53333#page/57/mode/1up	BM 1912.10.25.1		"Bitye, Ja River, S.E. Cameroons [= Cameroon]. 2000' [= 610 m]."			cuprosa O. Thomas, 1912	NA	NA	Guinea|Liberia|Côte d'Ivoire|Cameroon|Democratic Republic of the Congo|Ghana?	Africa	Afrotropic	DD	0	0	0	Kerivoula_cuprosa	0	sciname match	Kerivoula_cuprosa	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10971	Kerivoula cuprosa	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Kerivoula	cuprosa	Thomas, 1912		20000000	Kerivoula cuprosa	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of continuing problems with its taxonomy as well as absence of recent information on its extent of occurrence, ecological requirements and threats.	This is a species of moist lowland rainforest, tropical dry forest, and swamp forest. Other members of the genus have been found to roost in vegetation.	The threats to this species are not well known. In parts of its range it is presumably threatened to some degree by habitat loss resulting from conversion of land to agricultural use, and harvesting of timber and firewood.	It appears to be a rare, or rarely recorded, species.	Unknown	There are scattered records of this poorly known bat from West and Central Africa. It has apparently been reported from Guinea, Côte d'Ivoire, Liberia, possibly Ghana (though see Koopman et al. 1995; Grubb et al. 1998), Cameroon and the Democratic Republic of the Congo (though see Hayman and Hill 1971). It is found up to 600 m asl. A record of this species from Kenya was based on a specimen subsequently re-identified as Kerivoula smithii (J. Fahr pers. comm.).		Terrestrial	It is unclear if this species is present in any protected areas. Additional studies are needed into the taxonomy, distribution, natural history and threats to this little known bat.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Kerivoula		cuprosa	Thomas	1912	0	Ann. Mag. Nat. Hist.	ser. 8, 10: 41	Copper Woolly Bat	None.	Cameroon, Ja River, Bitye.	N Dem. Rep. Congo, S Cameroon. A record of this species from Kenya was based on a specimen subsequently reidentified as smithii (J. Fahr, pers. comm.)	Not listed.	Data Deficient		Kerivoula cuprosa	1005286	23	Coppery Woolly Bat	Copper Woolly Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	KERIVOULINAE	NA	Kerivoula	NA	cuprosa	O. Thomas	1912	0	Kerivoula_cuprosa	Thomas, O. (1912). Mammals from the Ja River, Cameroons. Annals and Magazine of Natural History, ser. 8, 10, 41.	https://www.biodiversitylibrary.org/item/53333#page/57/mode/1up	BM 1912.10.25.1		"Bitye, Ja River, S.E. Cameroons [= Cameroon]. 2000' [= 610 m]."			cuprosa O. Thomas, 1912	NA	NA				Guinea|Liberia|Côte d'Ivoire|Cameroon|Democratic Republic of the Congo|Ghana?	Africa	Afrotropic	DD	0	0	0	Kerivoula_cuprosa	0	sciname match	Kerivoula_cuprosa	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Kerivoula_cuprosa	1005286	23	Coppery Woolly Bat	Copper Woolly Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Kerivoulinae	NA	Kerivoula	NA	cuprosa	O. Thomas	0	Kerivoula cuprosa	Thomas, O. 1912-07-01. Mammals from the Ja River, Cameroons. Annals and Magazine of Natural History (8)10(55):41-43.	https://www.biodiversitylibrary.org/page/15618238	BMNH:Mamm:1912.10.25.1	holotype	https://data.nhm.ac.uk/object/b1d111a3-1534-48ce-9575-f3189448039a	"Bitye, Ja River, S.E. Cameroons [= Cameroon]. 2000' [= 610 m]."			NA	NA				Guinea|Liberia|Cote d'Ivoire|Cameroon|Democratic Republic of the Congo|Ghana?	Africa	Afrotropic	DD	0	0	0	Kerivoula_cuprosa	0	sciname match	Kerivoula_cuprosa	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Kerivoula		cuprosa	Thomas	1912	0	Ann. Mag. Nat. Hist.	ser. 8, 10: 41	Copper Woolly Bat	None.	Cameroon, Ja River, Bitye.	N Dem. Rep. Congo, S Cameroon. A record of this species from Kenya was based on a specimen subsequently reidentified as smithii (J. Fahr, pers. comm.)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10971/21971772/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Kerivoula cuprosa; Kerivoula cuprosa; Kerivoula cuprosa; Kerivoula cuprosa; Kerivoula cuprosa; Kerivoula cuprosa; cuprosa; Kérivoule cuivrée; Kupferfarbige Wollfledermaus; Querivoulacobrizo; Copper Woolly Bat; Coppery Woolly Bat; Copper Woolly Bat; Copper Woolly Bat; Copper Woolly Bat; K. cuprosa