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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L480	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus petersi	Pipistrellus petersi	Pipistrellus petersi	Pipistrellus petersi	Pipistrellus petersi	Falsistrellus petersi	Hypsugo petersi	Hypsugo petersi	Hypsugo petersi	Falsistrellus petersi	Hypsugo petersi	Hypsugo petersi	Hypsugo petersi	Hypsugo petersi	Hypsugo petersi		[MSW2] Subgenus Falsistrellus.; [MSW3] May be conspecific with affinis; see Francis and Hill (1986) and Corbet and Hill (1992).; [HMW] Vesperugo petersi A. B. Meyer, 1899 , Minahassa, northern Sulawesi , Indonesia . See Hypsugo affinis . Monotypic.; [batnames2022] Formerly included in Falsitrellus ; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis ; see Francis and Hill (1986) and Corbet and Hill (1992).; [MDD2022] moved from Falsistrellus to Hypsugo; [IUCN] The taxonomic status of this species requires review.; [batnames2023] Formerly included in Falsitrellus ; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis ; see Francis and Hill (1986) and Corbet and Hill (1992).; [MDD2023] moved from Falsistrellus to Hypsugo; [MDD2025_2.0] moved from Falsistrellus to Hypsugo; [batnames2025_1.7] Formerly included in Falsitrellus; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis; see Francis and Hill (1986) and Corbet and Hill (1992).; [MDD2025_2.2] moved from Falsistrellus to Hypsugo					(in P. affinis ?)									petersi	The taxonomic status of this species requires review.			petersi 	petersi 			petersi (A. B. Meyer, 1899)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Peters' pipistrelle	N Celebes, ? Buru, S Moluccas	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus petersi	Indonesia, Sulawesi, North Sulawesi, Minahassa.	Meyer	1899	Abh. Zool. Anthrop.-Ethnology. Mus. Dresden, 7(7):13.	Distribution: Ranging from Borneo and the Philippines through Celebes to the Moluccas.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Peters' pipistrelle	N Borneo – S Moluccas; ; ref. 4.131	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Meyer	1899	Abh. Zool. Anthrop.-Ethnology. Mus. Dresden, 7(7):13.	Subgenus Falsistrellus.	Borneo; Sulawesi; Buru and Amboina (Molucca Isis); Philippines.	Indonesia, Sulawesi, North Sulawesi, Minahassa.		MEYER	1899	Size medium (forearm length, 36-42 mm). Rostrum fairly broad. Forehead almost flat. Basicranial pits poorly developed.	Distribution: Ranging from Borneo and the Philippines through Celebes to the Moluccas.	No subspecies.		113	species	P. petersi	MEYER	1899	Pipistrellus	subgenus	Pipistrellus petersi				Size medium (forearm length, 36-42 mm). Rostrum fairly broad. Forehead almost flat. Basicranial pits poorly developed.	No subspecies.		23. P. petersi (MEYER 1899) [affinis group],	23	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Falsistrellus petersi	Falsistrellus		petersi	A. Meyer	y	1899		Abh. Zool. Anthrop.-Ethnology. Mus. Dresden	7	7	13		Peters's Pipistrelle	Indonesia, Sulawesi, N Sulawesi, Minahassa.	Borneo; Sulawesi; Buru and Amboina (Molucca Isls); Philippines.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc) as Pipistrellus petersi.		May be conspecific with affinis; see Francis and Hill (1986) and Corbet and Hill (1992).	4C3D87E8FFCC6A73FF8595A91B28B7D7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	813	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFCC6A73FF8595A91B28B7D7.xml	Hypsugo petersi	Vespertilionidae	Hypsugo	petersi	A. B. Meyer	1899	Vespere de Peters @fr | Peters-Zwergfledermaus @de | Hypsugo de Peters @es | North Wallacean Pipistrelle @en | Wallacean Chocolate Pipistrelle @en | Woolly Pipistrelle @en	Vesperugo petersi A. B. Meyer, 1899 , Minahassa, northern Sulawesi , Indonesia . See Hypsugo affinis . Monotypic.	Philippines (Mountain and Benguet provinces on Luzon I, and Surigao del Norte and Davao del Sur provinces on Mindanao I), Borneo (Crocker Range in Sabah , and Kubah National Park in Sarawak ), NE & C Sulawesi , and Moluccas (Buru and Ambon Is). A record from Mt Isarog (Camarines Sur Province on Luzon I) is assigned to the Pipustrellusjavanicus complex.	Head-body ¢.49-53 mm, tail 35-39 mm, ear 14-16 mm, hindfoot 8-9 mm, forearm 38-5—41-9 mm; weight 6-9 g. Dorsal fur of Peters’s Pipistrelle is somewhat woolly and long; upperparts are dark chocolate-brown with paler tips, giving frosted appearance; ventral fur is paler with brown tips; small amount of pale brown fur extends onto ventral surface of femur and adjacent to anus. Ears are blackish, broad, and rounded, taller than wide, with short, broad, blunt tragus. Muzzle is dark brown, and appears narrow and elongated compared to the Javan Pipistrelle ( Pipustrellus javanicus ), with less pronounced swellings on side; nostrils are simple (not tubular). Wing membranes are blackish and naked, except small area of pale brown fur extending onto ventral surface of femur and adjacent to anus. Tail tip extends c.1-5 mm beyond tail membrane; calcar is well developed, with slight keel extending medially from hindfoot. Adult males have long, pendulous penis. Compared to Peters’s Pipistrelle, the Javan Pipistrelle has shorter ears, shorter, broader muzzle, and dorsal pelage lacks frosted appearance. Baculum of a specimen from Buru, Moluccas , is similar to that of the Chocolate Pipistrelle ( Hypsugo affinis ) of India , broad and proximally widened with no distal expansion, and ventrally fluted. Skull is long and narrow, with slender zygomatic arches; occipital crests are well developed; sagittal and lambdoidal crests are present but low; forehead is almost flat. I? has prominent anterior and posterior cusps; C' is unicuspid with prominent lingual cingulum; P* is large and in contact with P* and lower incisors are strongly imbricated. Condylo-canine lengths are 13-6-14-7 mm, and maxillary tooth row lengths are 5-3-5-8 mm.	In Borneo, Peters’s Pipistrelle is known from lower montane moss forest in Sabah . Most records are from high elevation (above 1000 m ), but in Kubah National Park, Sarawak , it has been recorded in lowland forest. In the Philippines , specimens were caught in primary and heavily disturbed mossy forests and pine forests at high elevations (c.1200-2130 m). Nothing is known of habitat requirements in Sulawesi , Buru, and Ambon islands.	Diet is composed of small flying insects.	No information.	Peters’s Pipistrelle is nocturnal. In Borneo (Crocker Range) and the Philippines it was found roosting under the eaves of houses. Another specimen was captured on Mount Amuyao (LLuzon) in an area dominated by old-growth mossy forest. Peters’s Pipistrelles have been seen flying into wooden buildings.	No information.	Classified as Data Deficient on The IUCN Red List (as Falsistrellus petersi ), as there are only a few scattered records over a wide area. It may be declining because of widespread habitat loss in the region, but precise habitat requirements are not known. The species occurs in Kubah National Park. Further studies are needed into its distribution, abundance, ecology, and threats.	Boitani et al. (2006) | Corbet & Hill (1992) | Francis & Hill (1986) | Gorfol & Csorba (2018) | Gorfal, Furey et al. (2019) | Gorfol, Struebig et al. (2016) | Heaney, Balete, Alviola et al. (2012) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Hill & Harrison (1987) | Koopman (1994) | Kumaran et al. (2011) | Meyer (1899) | Payne et al. (1985) | Ruedi, Eger et al. (2017) | Sedlock et al. (2008) | Simmons (2005) | Tate (1942b)	https://zenodo.org/record/6397988/files/figure.png	104. Peters’s Pipistrelle Hypsugo petersi French: Vespere de Peters / German: Peters-Zwergfledermaus / Spanish: Hypsugo de Peters Other common names: North Wallacean Pipistrelle , Wallacean Chocolate Pipistrelle , Woolly Pipistrelle Taxonomy. Vesperugo petersi A. B. Meyer, 1899 , Minahassa, northern Sulawesi , Indonesia . See Hypsugo affinis . Monotypic. Distribution. Philippines (Mountain and Benguet provinces on Luzon I, and Surigao del Norte and Davao del Sur provinces on Mindanao I), Borneo (Crocker Range in Sabah , and Kubah National Park in Sarawak ), NE & C Sulawesi , and Moluccas (Buru and Ambon Is). A record from Mt Isarog (Camarines Sur Province on Luzon I) is assigned to the Pipustrellusjavanicus complex. Descriptive notes. Head-body ¢.49-53 mm, tail 35-39 mm, ear 14-16 mm, hindfoot 8-9 mm, forearm 38-5—41-9 mm; weight 6-9 g. Dorsal fur of Peters’s Pipistrelle is somewhat woolly and long; upperparts are dark chocolate-brown with paler tips, giving frosted appearance; ventral fur is paler with brown tips; small amount of pale brown fur extends onto ventral surface of femur and adjacent to anus. Ears are blackish, broad, and rounded, taller than wide, with short, broad, blunt tragus. Muzzle is dark brown, and appears narrow and elongated compared to the Javan Pipistrelle ( Pipustrellus javanicus ), with less pronounced swellings on side; nostrils are simple (not tubular). Wing membranes are blackish and naked, except small area of pale brown fur extending onto ventral surface of femur and adjacent to anus. Tail tip extends c.1-5 mm beyond tail membrane; calcar is well developed, with slight keel extending medially from hindfoot. Adult males have long, pendulous penis. Compared to Peters’s Pipistrelle, the Javan Pipistrelle has shorter ears, shorter, broader muzzle, and dorsal pelage lacks frosted appearance. Baculum of a specimen from Buru, Moluccas , is similar to that of the Chocolate Pipistrelle ( Hypsugo affinis ) of India , broad and proximally widened with no distal expansion, and ventrally fluted. Skull is long and narrow, with slender zygomatic arches; occipital crests are well developed; sagittal and lambdoidal crests are present but low; forehead is almost flat. I? has prominent anterior and posterior cusps; C' is unicuspid with prominent lingual cingulum; P* is large and in contact with P* and lower incisors are strongly imbricated. Condylo-canine lengths are 13-6-14-7 mm, and maxillary tooth row lengths are 5-3-5-8 mm. Habitat. In Borneo, Peters’s Pipistrelle is known from lower montane moss forest in Sabah . Most records are from high elevation (above 1000 m ), but in Kubah National Park, Sarawak , it has been recorded in lowland forest. In the Philippines , specimens were caught in primary and heavily disturbed mossy forests and pine forests at high elevations (c.1200-2130 m). Nothing is known of habitat requirements in Sulawesi , Buru, and Ambon islands. Food and Feeding. Diet is composed of small flying insects. Breeding. No information. Activity patterns. Peters’s Pipistrelle is nocturnal. In Borneo (Crocker Range) and the Philippines it was found roosting under the eaves of houses. Another specimen was captured on Mount Amuyao (LLuzon) in an area dominated by old-growth mossy forest. Peters’s Pipistrelles have been seen flying into wooden buildings. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List (as Falsistrellus petersi ), as there are only a few scattered records over a wide area. It may be declining because of widespread habitat loss in the region, but precise habitat requirements are not known. The species occurs in Kubah National Park. Further studies are needed into its distribution, abundance, ecology, and threats. Bibliography. Boitani et al. (2006), Corbet & Hill (1992), Francis & Hill (1986), Gorfol & Csorba (2018), Gorfal, Furey et al. (2019), Gorfol, Struebig et al. (2016), Heaney, Balete, Alviola et al. (2012), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Hill & Harrison (1987), Koopman (1994), Kumaran et al. (2011), Meyer (1899), Payne et al. (1985), Ruedi, Eger et al. (2017), Sedlock et al. (2008), Simmons (2005), Tate (1942b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hypsugo petersi	Hypsugo		petersi	A. Meyer	1899	1	Abh. Zool. Anthrop.-Ethnology. Mus. Dresden	7(7): 13	Peters' Pipistrelle	None.	Indonesia, Sulawesi, N Sulawesi, Minahassa.	Borneo; Sulawesi; Buru and Amboina (Molucca Isls); Philippines.	Not listed.	Data Deficient under Falsistrellus petersi 	Formerly included in Falsitrellus ; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis ; see Francis and Hill (1986) and Corbet and Hill (1992).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hypsugo petersi	23	Peters's Pipistrelle	North Wallacean Pipistrelle|Wallacean Chocolate Pipistrelle|Woolly Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	petersi	A. B. Meyer	1899	1						Minahassa, northern Sulawesi, Indonesia.			petersi (A. B. Meyer, 1899)	moved from Falsistrellus to Hypsugo	Górfól, T., & Csorba, G. (2018). Integrative taxonomy places Asian species of Falsistrellus (Chiroptera: Vespertilionidae) into Hypsugo. Mammalian Biology, 93(1), 56-63.	Philippines|Indonesia|Malaysia	Asia|Oceania	Indomalaya|Australasia/Oceania	DD	0	0	0	Hypsugo_petersi	0	sciname match	Falsistrellus_petersi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17359	Falsistrellus petersi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Falsistrellus	petersi	(Meyer, 1899)	The taxonomic status of this species requires review.	20000000	Falsistrellus petersi	Data Deficient		2016	2015-12-24 00:00:00 UTC	3.1	English	This species is assessed as Data Deficient as there are only a few scattered records over a wide area. It may be declining because of widespread habitat loss in the region, but precise habitat requirements are not known. Taxonomic revision of this species is required.	<p>There is little information available about the ecology of this species. A colony was found roosting under the eaves of an isolated house in lower montane forest in Sabah (Payne et al . 1985). Specimens from the Philippines were caught in disturbed and undisturbed forests and in buildings in high elevations (Heaney et al. 2016).</p>	The threats to this species are not known.	The abundance and population size of this species are not known.	Unknown	This species is found on Borneo, Sulawesi, Buru and Amboina islands (Molucca Islands, Indonesia), and the Philippines. In the Philippines the species has been recorded from Luzon (Benguet province) and Mindanao (Davao del Sur, and Surigao del Norte provinces) (Heaney et al . 1998).		Terrestrial	The species occurs in Kubah National Park, Sarawak, Malaysia (Jayaraj et al. 2011) Further studies are needed into the distribution, abundance, ecology and threats to this species.	Australasian|Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hypsugo		petersi	A. Meyer	1899	1	Abh. Zool. Anthrop.-Ethnology. Mus. Dresden	7(7): 13	Peters' Pipistrelle	None.	Indonesia, Sulawesi, N Sulawesi, Minahassa.	Borneo; Sulawesi; Buru and Amboina (Molucca Isls); Philippines.	Not listed.	Data Deficient under Falsistrellus petersi 	Formerly included in Falsitrellus ; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis ; see Francis and Hill (1986) and Corbet and Hill (1992).	Hypsugo petersi	1005725	23	Peters's Pipistrelle	North Wallacean Pipistrelle|Wallacean Chocolate Pipistrelle|Woolly Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	petersi	A. B. Meyer	1899	1						Minahassa, northern Sulawesi, Indonesia.			petersi (A. B. Meyer, 1899)	moved from Falsistrellus to Hypsugo	Górfól, T., & Csorba, G. (2018). Integrative taxonomy places Asian species of Falsistrellus (Chiroptera: Vespertilionidae) into Hypsugo. Mammalian Biology, 93(1), 56-63.				Philippines|Indonesia|Malaysia	Asia|Oceania	Indomalaya|Australasia/Oceania	DD	0	0	0	Hypsugo_petersi	0	sciname match	Falsistrellus_petersi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hypsugo_petersi	1005725	23	Peters's Pipistrelle	North Wallacean Pipistrelle|Wallacean Chocolate Pipistrelle|Woolly Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo	NA	petersi	A. B. Meyer	1	Vesperugo petersi	Meyer, A.B. 1899. Säugethiere vom Celébes- und Philippinen-Archipel. II. Celébes-Sammlungen der Herren Sarasin. Abhandlungen und Berichte des königlichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 7(7):1-55.	https://www.biodiversitylibrary.org/page/33471547	MTD B3805 (lost!) (= MTD B2524)	holotype		Minahassa, northern Sulawesi, Indonesia.			moved from Falsistrellus to Hypsugo	Górfól, T., & Csorba, G. (2018). Integrative taxonomy places Asian species of Falsistrellus (Chiroptera: Vespertilionidae) into Hypsugo. Mammalian Biology, 93(1), 56-63.				Philippines|Indonesia|Malaysia	Asia|Oceania (Continent)	Indomalaya|Australasia	DD (as Falsistrellus petersi)	0	0	0	Hypsugo_petersi	0	sciname match	Falsistrellus_petersi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hypsugo		petersi	A. Meyer	1899	1	Abh. Zool. Anthrop.-Ethnology. Mus. Dresden	7(7): 13	Peters' Pipistrelle	None.	Indonesia, Sulawesi, N Sulawesi, Minahassa.	Borneo; Sulawesi; Buru and Amboina (Molucca Isls); Philippines.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17359/22125873/' target='_blank'>Data Deficient as Falsistrellus petersi</a>	Formerly included in Falsitrellus; transferred to Hypsugo by Görföl and Csorba (2018). May be conspecific with affinis; see Francis and Hill (1986) and Corbet and Hill (1992).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Falsistrellus petersi; Hypsugo petersi; Hypsugo petersi; Hypsugo petersi; Falsistrellus petersi; Hypsugo petersi; petersi; Vespere de Peters; Peters-Zwergfledermaus; Hypsugo de Peters; North Wallacean Pipistrelle; Wallacean Chocolate Pipistrelle; Woolly Pipistrelle; Peters's Pipistrelle; North Wallacean Pipistrelle; Wallacean Chocolate Pipistrelle; Woolly Pipistrelle; Peters's Pipistrelle; Peters' Pipistrelle; H. petersi