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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L476	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus macrotis	Pipistrellus macrotis	Pipistrellus macrotis	Pipistrellus macrotis	Pipistrellus macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis	Hypsugo macrotis		[MSW2] Subgenus Pipistrellus. Listed as a subspecies of imbricatus by Medway (1969:39); but see Tate (1942fl:249) and Corbet and Hill (1980:67). Includes curtatus and vordermanni, but see Hill (1983).; [MSW3] Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942a) and Corbet and Hill (1980). May include vordermanni; see Corbet and Hill (1992).; [HMW] Vespertilio macrotis Temminck, 1840 , “le district de Padang, ile de Sumatra ,” Indonesia . Hypsugo macrotis has been considered conspecific with H. imbricatus , but they are generally recognized as distinct species based on morphological data. It appears to be closely related to H. vordermanni , which is morphologically very similar. Monotypic.; [batnames2022] Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942 a ) and Corbet and Hill (1980). May include vordermanni ; see Corbet and Hill (1992).; [IUCN] <p>The species may be conspecific with vordermanni (Corbet and Hill 1992).</p>; [batnames2023] Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942 a ) and Corbet and Hill (1980). May include vordermanni ; see Corbet and Hill (1992).; [batnames2025_1.7] Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942a) and Corbet and Hill (1980). May include vordermanni; see Corbet and Hill (1992).						curtatus, vordermanni.	vordermanni, curtatus, macrotis		curtatus			macrotis 	macrotis - curtatus	macrotis, curtatus	The species may be conspecific with vordermanni (Corbet and Hill 1992).</p>	macrotis 	macrotis - curtatus	macrotis, curtatus 	macrotis, curtatus 	macrotis	macrotis - curtatus	macrotis (Temminck, 1840)|curtatus (G. S. Miller, 1911)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Sumatra etc.	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus macrotis	Indonesia, Sumatra, Padang.	Temminck	1840	Monogr. Mamm., 2:218.	Distribution: Ranging from Malaya to Bali and Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Malaya, Sumatra, Bali; ref. 4.14	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1840	Monogr. Mamm., 2:218.	Subgenus Pipistrellus. Listed as a subspecies of imbricatus by Medway (1969:39); but see Tate (1942fl:249) and Corbet and Hill (1980:67). Includes curtatus and vordermanni, but see Hill (1983).	W Malaysia, Sumatra, Borneo, Bali, adjacent small islands.	Indonesia, Sumatra, Padang.		TEMMINCK	1835	Size relatively small (forearm length, 29-35 mm). Inner upper incisor more or less unicuspid. Anterior upper premolar greatly reduced and displaced medially. Rostrum relatively short but fairly slender. Forehead fairly concave. Braincase relatively inflated. Basicranial pits well developed. A small postorbital process present on zygomatic arch.	Distribution: Ranging from Malaya to Bali and Borneo.	Three subspecies are recognized:	P. m. macrotis (Malaya to Bali), P. m. vordermanni (Borneo and Billiton), P. m. curtatus (West Sumatran islands).	115	species	P. macrotis	TEMMINCK	1835	Pipistrellus	subgenus	Pipistrellus macrotis				Size relatively small (forearm length, 29-35 mm). Inner upper incisor more or less unicuspid. An- terior upper premolar greatly reduced and displaced medially. Rostrum relatively short but fairly slender. Forehead fairly concave. Braincase relatively inflated. Basicranial pits well developed. A small postorbital process present on zygomatic arch.	Three subspecies are recognized:		36. P. macrotis (TEMMINCK 1835) [savii group].	36	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo macrotis	Hypsugo		macrotis	Temminck	y	1840		Monogr. Mamm.	2		218		Big-eared Pipistrelle	Indonesia, Sumatra, Padang.	W Malaysia, Sumatra, Bali, adjacent small islands.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Pipistrellus macrotis.	curtatus Miller, 1911.	Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942a) and Corbet and Hill (1980). May include vordermanni; see Corbet and Hill (1992).	4C3D87E8FFC36A7CFF499A36188EBEE4	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	814	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFC36A7CFF499A36188EBEE4.xml	Hypsugo macrotis	Vespertilionidae	Hypsugo	macrotis		1840	espére a grandes oreilles @fr | Grofsohr-Zwergfledermaus @de | Hypsugo orejudo @es	Vespertilio macrotis Temminck, 1840 , “le district de Padang, ile de Sumatra ,” Indonesia . Hypsugo macrotis has been considered conspecific with H. imbricatus , but they are generally recognized as distinct species based on morphological data. It appears to be closely related to H. vordermanni , which is morphologically very similar. Monotypic.	W Peninsular Malaysia , Singapore , NE Sumatra, and Padang, Enggano, Bali, and Lombok Is.	Head-body ¢. 50 mm , tal ¢.20 mim, ear c¢.13-5 mm, forearm 31-7-34-5 mm; weight 4-6 g. Dorsal pelage of the Big-eared Pipistrelle is reddish brown (hairs with dark brown bases); ventral pelage is a little paler. Ears are comparatively long, broad, and subtriangular with broadly rounded tip; tragus is short, broad, and hatchet-shaped. Wing membranes are translucent white with brown tinge. Skull is larger and shorter than in Vordermann’s Pipistrelle ( Hypsugo vordermanni ); rostrum is short; forehead has evenly sloping profile. P* is minute and completely displaced inwardly; C' and P* are in contact.	The Big-eared Pipistrelle has been reported foraging over open mudflats in coastal regions in Peninsular Malaysia and in lowland rainforest.	The Big-eared Pipistrelle has been observed foraging 3 m around the edge and canopy ofa forest.	Big-eared Pipistrelles form maternity colonies during the breeding season with 10-20 individuals.	A maternity colony in Malaysia was recorded roosting between a pipeline and a pillar and Big-eared Pipistrelles were observed switching roosts in response to a feral cat disturbing their roost.	Big-eared Pipistrelles roost in colonies.	Classified as Data Deficient on The IUCN Red List. The Bigeared Pipistrelle is considered a rare species throughout its scattered distribution, but it may be more common than currently known.	Francis (2008a) | Gorfol, Bates et al. (2016) | Lee & Teo (2018) | Lim, L.S. et al. (2016) | McKenzie, Gunnell et al. (1995)	https://zenodo.org/record/6397998/files/figure.png	109. Big-eared Pipistrelle Hypsugo macrotis French: espére a grandes oreilles / German: Grofsohr-Zwergfledermaus / Spanish: Hypsugo orejudo Taxonomy. Vespertilio macrotis Temminck, 1840 , “le district de Padang, ile de Sumatra ,” Indonesia . Hypsugo macrotis has been considered conspecific with H. imbricatus , but they are generally recognized as distinct species based on morphological data. It appears to be closely related to H. vordermanni , which is morphologically very similar. Monotypic. Distribution. W Peninsular Malaysia , Singapore , NE Sumatra, and Padang, Enggano, Bali, and Lombok Is. Descriptive notes. Head-body ¢. 50 mm , tal ¢.20 mim, ear c¢.13-5 mm, forearm 31-7-34-5 mm; weight 4-6 g. Dorsal pelage of the Big-eared Pipistrelle is reddish brown (hairs with dark brown bases); ventral pelage is a little paler. Ears are comparatively long, broad, and subtriangular with broadly rounded tip; tragus is short, broad, and hatchet-shaped. Wing membranes are translucent white with brown tinge. Skull is larger and shorter than in Vordermann’s Pipistrelle ( Hypsugo vordermanni ); rostrum is short; forehead has evenly sloping profile. P* is minute and completely displaced inwardly; C' and P* are in contact. Habitat. The Big-eared Pipistrelle has been reported foraging over open mudflats in coastal regions in Peninsular Malaysia and in lowland rainforest. Food and Feeding. The Big-eared Pipistrelle has been observed foraging 3 m around the edge and canopy ofa forest. Breeding. Big-eared Pipistrelles form maternity colonies during the breeding season with 10-20 individuals. Activity patterns. A maternity colony in Malaysia was recorded roosting between a pipeline and a pillar and Big-eared Pipistrelles were observed switching roosts in response to a feral cat disturbing their roost. Movements, Home range and Social organization. Big-eared Pipistrelles roost in colonies. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Bigeared Pipistrelle is considered a rare species throughout its scattered distribution, but it may be more common than currently known. Bibliography. Francis (2008a), Gorfol, Bates et al. (2016), Lee & Teo (2018), Lim, L.S. et al. (2016), McKenzie, Gunnell et al. (1995).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hypsugo macrotis	Hypsugo		macrotis	Temminck	1840	1	Monogr. Mamm.	0.2347	Big-eared Pipistrelle	 curtatus Miller, 1911.	Indonesia, Sumatra, Padang.	W Malaysia, Sumatra, Bali, adjacent small islands.	Not listed.	Data Deficient	Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942 a ) and Corbet and Hill (1980). May include vordermanni ; see Corbet and Hill (1992).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hypsugo macrotis	23	Big-eared Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	macrotis	Temminck	1840	1	Vespertilio_macrotis	Temminck, C. J. (1840). Monographies de Mammalogie, ou description de quelques genres de mammifÃ©res, dont les espÃ©ces ont Ã©tÃ© observÃ©es dens les diffÃ©rens musÃ©es de l'Europe. G. Dufour and E. d'Ocagne, Paris, Vol. 2, 218.	https://books.google.com/books?hl=en&lr=&id=Lw9jAAAAcAAJ&oi=fnd&pg=PA1&dq=monographies+de+mammalogie+temminck+vol.+2&ots=o25ZP2qQxx&sig=jNZE7B2jrj-W7sB7y_Bz5_qjud8#v=onepage&q=monographies%20de%20mammalogie%20temminck%20vol.%202&f=false	RMNH MAM.35469-35471 [syntypes]		"le district de Padang, ile de Sumatra," Indonesia.			macrotis (Temminck, 1840)|curtatus (G. S. Miller, 1911)	NA	NA	Malaysia|Singapore|Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_macrotis	0	sciname match	Hypsugo_macrotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17349	Hypsugo macrotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hypsugo	macrotis	Temminck, 1840	<p>The species may be conspecific with vordermanni (Corbet and Hill 1992).</p>	20000000	Hypsugo macrotis	Data Deficient		2016	2015-12-24 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of recent information on its population status, ecological requirements and major threats.	The species has been caught foraging over open mud flats in coastal areas in Peninsular Malaysia (Francis, C.M. pers. comm). It was also observed foraging 3 m around the edge and canopy of lowland rainforests (McKenzie et al . 1995) and a specimen possibly belonging to this species were mist-netted in a plantation in Jambi, East Sumatra, Indonesia (Darras, K., Lobite, N.L. and Aryanti, E.S. pers. comm). A maternity colony was found in a crevice between a pillar and a pipeline of a building in Seremban, Malaysia. It is not known whether it is tolerant of any disturbance of its habitat.	Deforestation due to agriculture, plantations, logging and forest fires is occurring within the species' range, but it is not known whether this species is forest dependent.	The abundance and population size of this species are not known.	Unknown	This species occurs in western Malaysia, Indonesia (Sumatra, Padang, Bali, Lombok, and adjacent small islands). It is not known whether it occurs in Java.		Terrestrial	It is not known if the species is present in any protected areas. Further studies are needed into the distribution, abundance, threats and ecology of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hypsugo		macrotis	Temminck	1840	1	Monogr. Mamm.	0.234722	Big-eared Pipistrelle	 curtatus Miller, 1911.	Indonesia, Sumatra, Padang.	W Malaysia, Sumatra, Bali, adjacent small islands.	Not listed.	Data Deficient	Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942 a ) and Corbet and Hill (1980). May include vordermanni ; see Corbet and Hill (1992).	Hypsugo macrotis	1005722	23	Big-eared Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	macrotis	Temminck	1840	1	Vespertilio_macrotis	Temminck, C. J. (1840). Monographies de Mammalogie, ou description de quelques genres de mammifÃ©res, dont les espÃ©ces ont Ã©tÃ© observÃ©es dens les diffÃ©rens musÃ©es de l'Europe. G. Dufour and E. d'Ocagne, Paris, Vol. 2, 218.	https://books.google.com/books?hl=en&lr=&id=Lw9jAAAAcAAJ&oi=fnd&pg=PA1&dq=monographies+de+mammalogie+temminck+vol.+2&ots=o25ZP2qQxx&sig=jNZE7B2jrj-W7sB7y_Bz5_qjud8#v=onepage&q=monographies%20de%20mammalogie%20temminck%20vol.%202&f=false	RMNH MAM.35469-35471 [syntypes]		"le district de Padang, ile de Sumatra," Indonesia.			macrotis (Temminck, 1840)|curtatus (G. S. Miller, 1911)	NA	NA				Malaysia|Singapore|Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_macrotis	0	sciname match	Hypsugo_macrotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hypsugo_macrotis	1005722	23	Big-eared Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo	NA	macrotis	Temminck	1	Vespertilio macrotis	Temminck, C.J. 1840. Livraison 3. Pp. 141â€“272 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog/page/140/mode/2up	RMNH.MAM.35448, RMNH.MAM.35449, RMNH.MAM.35469, RMNH.MAM.35470, RMNH.MAM.35471	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35448 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35449 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35469 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35470 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35471	"le district de Padang, ile de Sumatra," Indonesia.			NA	NA				Malaysia|Singapore|Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_macrotis	0	sciname match	Hypsugo_macrotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hypsugo		macrotis	Temminck	1840	1	Monogr. Mamm.	0.234722	Big-eared Pipistrelle	curtatus Miller, 1911.	Indonesia, Sumatra, Padang.	W Malaysia, Sumatra, Bali, adjacent small islands.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17349/22127259/' target='_blank'>Data Deficient</a>	Listed as a subspecies of imbricatus by Medway (1969), but see Tate (1942a) and Corbet and Hill (1980). May include vordermanni; see Corbet and Hill (1992).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hypsugo macrotis; Hypsugo macrotis; Hypsugo macrotis; Hypsugo macrotis; Hypsugo macrotis; Hypsugo macrotis; curtatus; curtatus; macrotis; curtatus; espére a grandes oreilles; Grofsohr-Zwergfledermaus; Hypsugo orejudo; Big-eared Pipistrelle; Big-eared Pipistrelle; Big-eared Pipistrelle; H. macrotis
