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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L473	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus kitcheneri	Pipistrellus kitcheneri	Pipistrellus kitcheneri	Pipistrellus kitcheneri	Pipistrellus kitchneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri	Hypsugo kitcheneri		[MSW2] Subgenus Hypsugo. Listed as a subspecies of imbricatus by Chasen (1940); but see Tate (1942fl:221) and Medway (1977:55).; [MSW3] Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942a) and Medway (1977). May be conspecific with lophurus; see Francis and Hill (1986).; [HMW] Pipistrellus kitchener: Thomas, 1915 , “Boentok, Barito River [ Central Kalimantan , Borneo]. Alt. 20’ [= 6 m ].” Hypsugo kitcheneri may be conspecific with H. lophurus , although the bacula of the two are morphologically distinct. Monotypic.; [batnames2022] Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942 a ) and Medway (1977). May be conspecific with lophurus ; see Francis and Hill (1986).; [IUCN] Francis and Hill (1986) argue that this species may be conspecific with lophurus .; [batnames2023] Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942 a ) and Medway (1977). May be conspecific with lophurus ; see Francis and Hill (1986).; [batnames2025_1.7] Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942a) and Medway (1977). May be conspecific with lophurus; see Francis and Hill (1986).														kitcheneri	Francis and Hill (1986) argue that this species may be conspecific with lophurus .			kitcheneri	kitcheneri			kitcheneri (O. Thomas, 1915)|kitchneri (Koopman, 1994) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus kitcheneri	Malaysia, Kalimantan, Barito River.	Thomas	1916	Ann. Mag. Nat. Hist., ser. 8, 15:229.	Distribution: Confined to Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1916	Ann. Mag. Nat. Hist., ser. 8, 15:229.	Subgenus Hypsugo. Listed as a subspecies of imbricatus by Chasen (1940); but see Tate (1942fl:221) and Medway (1977:55).	Borneo.	Borneo, Kalimantan Tengah, Barito River.		THOMAS	1915	Size fairly small (forearm length, 34-38 mm). Anterior upper premolar greatly reduced and displaced medially. Rostrum fairly broad. Forehead almost flat. Basicranial pits well developed.	Distribution: Confined to Borneo.	No subspecies.		113	species	P. kitchneri	THOMAS	1915	Pipistrellus	subgenus	Pipistrellus kitchneri				Size fairly small (forearm length, 34-38 mm). Anterior upper premolar greatly reduced and displaced medially. Rostrum fairly broad. Forehead almost flat. Basicranial pits well developed.	No subspecies.		21. P. kitchneri THOMAS 1915 [pulveratus group],	21	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo kitcheneri	Hypsugo		kitcheneri	Thomas	y	1915		Ann. Mag. Nat. Hist., ser. 8	15		229		Red-brown Pipistrelle	Borneo, Kalimantan Tengah, Barito River.	Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt) as Pipistrellus kitcheneri.		Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942a) and Medway (1977). May be conspecific with lophurus; see Francis and Hill (1986).	4C3D87E8FFC36A7CFF4B975D1F64BEE6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	814	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFC36A7CFF4B975D1F64BEE6.xml	Hypsugo kitcheneri	Vespertilionidae	Hypsugo	kitcheneri		1915	Vespere de Kitchener @fr | Kitchener Zwergfledermaus @de | Hypsugo de Kitchener @es	Pipistrellus kitchener: Thomas, 1915 , “Boentok, Barito River [ Central Kalimantan , Borneo]. Alt. 20’ [= 6 m ].” Hypsugo kitcheneri may be conspecific with H. lophurus , although the bacula of the two are morphologically distinct. Monotypic.	NC, SW & SE Borneo; possibly in extreme N Borneo.	Head-body ¢. 56 mm , tail c. 41 mm , ear c. 14 mm , forearm 35-38 mm. Dorsal pelage of the Red-brown Pipistrelle is rich reddish brown (hairs with black bases); ventral pelage is slightly paler and grayer. Membranes are brown. Ears are short and subtriangular; tragus is broad and curved forward. Skull is moderately large and slightly broadened; rostrum is comparatively elongate; zygomatic arches are moderately thick with distinct postorbital processes; basioccipital pits are deep and distinct. C' is unicuspid; P? is small and intruded, I? has large posterior cusp; lower incisors are weakly or not imbricated; I, is slightly larger than I; and P, is slightly over one-half the area and two-thirds the height of P.	Recorded from sea level to elevations of 1500 m .	No information.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List. Virtually nothing is known regarding ecology and threats of the Red-brown Pipistrelle; further studies are needed. Recorded from Mount Palung and Kayan Mentarang national parks.	Francis & Hill (1986) | Gorfol, Kingston & Hutson (2016) | Medway (1977) | Payne et al. (1985) | Thomas (1915a)	https://zenodo.org/record/6397994/files/figure.png	107. Red-brown Pipistrelle Hypsugo kitcheneri French: Vespere de Kitchener / German: KitchenerZwergfledermaus / Spanish: Hypsugo de Kitchener Taxonomy. Pipistrellus kitchener: Thomas, 1915 , “Boentok, Barito River [ Central Kalimantan , Borneo]. Alt. 20’ [= 6 m ].” Hypsugo kitcheneri may be conspecific with H. lophurus , although the bacula of the two are morphologically distinct. Monotypic. Distribution. NC, SW & SE Borneo; possibly in extreme N Borneo. Descriptive notes. Head-body ¢. 56 mm , tail c. 41 mm , ear c. 14 mm , forearm 35-38 mm. Dorsal pelage of the Red-brown Pipistrelle is rich reddish brown (hairs with black bases); ventral pelage is slightly paler and grayer. Membranes are brown. Ears are short and subtriangular; tragus is broad and curved forward. Skull is moderately large and slightly broadened; rostrum is comparatively elongate; zygomatic arches are moderately thick with distinct postorbital processes; basioccipital pits are deep and distinct. C' is unicuspid; P? is small and intruded, I? has large posterior cusp; lower incisors are weakly or not imbricated; I, is slightly larger than I; and P, is slightly over one-half the area and two-thirds the height of P. Habitat. Recorded from sea level to elevations of 1500 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Virtually nothing is known regarding ecology and threats of the Red-brown Pipistrelle; further studies are needed. Recorded from Mount Palung and Kayan Mentarang national parks. Bibliography. Francis & Hill (1986), Gorfol, Kingston & Hutson (2016), Medway (1977), Payne et al. (1985), Thomas (1915a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hypsugo kitcheneri	Hypsugo		kitcheneri	Thomas	1915	1	Ann. Mag. Nat. Hist.	ser. 8, 15: 229	Red-brown Pipistrelle	None.	Borneo, Kalimantan Tengah, Barito River.	Borneo.	Not listed.	Data Deficient	Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942 a ) and Medway (1977). May be conspecific with lophurus ; see Francis and Hill (1986).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hypsugo kitcheneri	23	Red-brown Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	kitcheneri	O. Thomas	1915	1	Pipistrellus_kitcheneri	Thomas, O. (1915). On bats of the genera Nyctalus, Tylonycteris, and Pipistrellus. Annals and Magazine of Natural History, Ser. 8, 15, 229.	https://www.biodiversitylibrary.org/item/78262#page/241/mode/1up	BM 1910.4.5.55		"Boentok, Barito River [Central Kalimantan, Borneo]. Alt. 20' [= 6 m]."			kitcheneri (O. Thomas, 1915)	NA	NA	Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_kitcheneri	0	sciname match	Hypsugo_kitcheneri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	17346	Hypsugo kitcheneri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hypsugo	kitcheneri	Thomas, 1915	Francis and Hill (1986) argue that this species may be conspecific with lophurus .	20000000	Hypsugo kitcheneri	Data Deficient		2016	2015-12-24 00:00:00 UTC	3.1	English	Listed as Data Deficient as it is known only from a couple of locations and there is no further information regarding its distribution, abundance, ecology or threats. It may be conspecific with lophurus .	There is little information available about the ecology and habitat preferences of this species. Presumably it inhabits lowland and montane forest.	The threats to this species are not known, but habitat loss may be a threat if it inhabits lowland forest.	The abundance and population size of this species are not known.	Unknown	<p>The type locality is in Kalimantan Tengah (Barito River), in Indonesian Borneo. It was also reported from Gunung Palung National Park (Blundell 1996), and from Kayan Mentarang National Park (Puri 1997, Jayasilan et al . 2003). It was collected also in Sandakan, Sabah, Malaysia and Buntok, Central Kalimantan, Indonesia (Payne et al . 1985) and Laham, Indonesia (Medway 1977).</p>		Terrestrial	The species has been reported from protected areas; Gunung Palung National Park (Blundell 1996), and Kayan Mentarang National Park (Puri 1997, Jayasilan et al. 2003). Further studies are needed into the taxonomy, distribution, abundance, ecology and threats to this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hypsugo		kitcheneri	Thomas	1915	1	Ann. Mag. Nat. Hist.	ser. 8, 15: 229	Red-brown Pipistrelle	None.	Borneo, Kalimantan Tengah, Barito River.	Borneo.	Not listed.	Data Deficient	Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942 a ) and Medway (1977). May be conspecific with lophurus ; see Francis and Hill (1986).	Hypsugo kitcheneri	1005719	23	Red-brown Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	kitcheneri	O. Thomas	1915	1	Pipistrellus_kitcheneri	Thomas, O. (1915). On bats of the genera Nyctalus, Tylonycteris, and Pipistrellus. Annals and Magazine of Natural History, Ser. 8, 15, 229.	https://www.biodiversitylibrary.org/item/78262#page/241/mode/1up	BM 1910.4.5.55		"Boentok, Barito River [Central Kalimantan, Borneo]. Alt. 20' [= 6 m]."			kitcheneri (O. Thomas, 1915)	NA	NA				Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_kitcheneri	0	sciname match	Hypsugo_kitcheneri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hypsugo_kitcheneri	1005719	23	Red-brown Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo	NA	kitcheneri	O. Thomas	1	Pipistrellus kitcheneri	Thomas, O. 1915-02-01. On bats of the genera _Nyctalus_, _Tylonycteris_, and _Pipistrellus_. Annals and Magazine of Natural History (8)15(86):225-232.	https://www.biodiversitylibrary.org/page/24255799	BMNH:Mamm:1910.4.5.55	holotype	https://data.nhm.ac.uk/object/2b0c8048-a641-469e-b71a-b5e4a9019796	"Boentok, Barito River [Central Kalimantan, Borneo]. Alt. 20' [= 6 m]."			NA	NA				Indonesia	Asia	Indomalaya	DD	0	0	0	Hypsugo_kitcheneri	0	sciname match	Hypsugo_kitcheneri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hypsugo		kitcheneri	Thomas	1915	1	Ann. Mag. Nat. Hist.	ser. 8, 15: 229	Red-brown Pipistrelle	None.	Borneo, Kalimantan Tengah, Barito River.	Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/17346/22127626/' target='_blank'>Data Deficient</a>	Listed as a subspecies of imbricatus by Chasen (1940), but see Tate (1942a) and Medway (1977). May be conspecific with lophurus; see Francis and Hill (1986).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hypsugo kitcheneri; Hypsugo kitcheneri; Hypsugo kitcheneri; Hypsugo kitcheneri; Hypsugo kitcheneri; Hypsugo kitcheneri; kitcheneri; Vespere de Kitchener; Kitchener Zwergfledermaus; Hypsugo de Kitchener; Red-brown Pipistrelle; Red-brown Pipistrelle; Red-brown Pipistrelle; H. kitcheneri