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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L471	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon	Hypsugo dolichodon		[HMW] Hypsugo dolichodon Gorfol et al., 2014 , “ 14°57'N , 107°08'E , Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l.” The relationships of Hypsugo dolichodon are uncertain, but it appears to be close to H. petersi and H. pulveratus , based on limited genetic data. Specimens from Cambodia and Myanmar were originally identified as H. affinis , but are now considered to represent this species. Monotypic.; [IUCN] The taxon was described in 2014 based on specimens from Lao PDR and Vietnam. It belongs to the â€œpulveratus -groupâ€ of Hypsugo .														dolichodon	The taxon was described in 2014 based on specimens from Lao PDR and Vietnam. It belongs to the â€œpulveratus -groupâ€ of Hypsugo .			dolichodon 	dolichodon 			dolichodon T. GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & C. M. Francis, 2014						N/A																																								NA																											4C3D87E8FFCC6A73FF869B731BB7BCB3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	813	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFCC6A73FF869B731BB7BCB3.xml	Hypsugo dolichodon	Vespertilionidae	Hypsugo	dolichodon	Gorfol	2014	Vespere a longues dents @fr | Langzahn-Zwergfledermaus @de | Hypsugo de dientes largos @es	Hypsugo dolichodon Gorfol et al., 2014 , “ 14°57'N , 107°08'E , Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l.” The relationships of Hypsugo dolichodon are uncertain, but it appears to be close to H. petersi and H. pulveratus , based on limited genetic data. Specimens from Cambodia and Myanmar were originally identified as H. affinis , but are now considered to represent this species. Monotypic.	Known from scattered localities in SE Myanmar , W & S Cambodia , SE Laos , and S Vietnam .	Hindfoot 5-9-7 mm, forearm 35-2-38-4 mm. No other specific measurements are available. Dorsal pelage of the Long-toothed Pipistrelle is blackish brown (hairs with paler tips), and ventral pelage lighter (hairs blackish brown on basal one-half and dirty white on distal one-half). Bare portions of face, ears, limbs, and wings are nearly black. Ears are relatively short and wide and somewhat rounded at tip; tragus is short and squarish. Tail extends a little beyond margin of uropatagium; calcar extends halfway to tail from ankle, and has narrow Postcalcarial lobe—Tless defined than in the Chinese Pipistrelle ( Hypsugo pulveratus ). Baculum has more developed base with enlarged lateral projection, narrowing shaft, and shallower ventral depression on tip when compared to the Chocolate Pipistrelle ( H. affinis ). Skull is large with relatively robust, long, narrow rostrum; rostral profile of skull is almost flat; braincase is flattened; there is no sagittal crest; lambdoid crest is pronounced; and basioccipital pits are shallow but visible. I* is bicuspid; C' and C, are relatively slender and long; P? is small and intruded but C' and P* are still separated; and lower molars are myotodont.	The Long-toothed Pipistrelle has been reported in lowland evergreen rainforest at elevations of 50-400 m throughout much ofits range, and is associated with limestone outcrops. Reported at 978 m elevation in limestone caves surrounded by evergreen rainforest and cultivated areas in Cambodia .	No information.	No information.	Long-toothed Pipistrelles roost in caves.	No information.	Not assessed on The [UCN Red List. The Long-toothed Pipistrelle is known only from a few localities and does not seem to be very common. It may be threatened by habitat destruction due to limestone extraction, and general human disturbance. Further studies are needed, particularly into general ecology and threats.	Gorfol & Csorba (2018) | Gorfél, Csorba et al. (2014) | Gorfol, Furey et al. (2019) | Lim, L.S. et al. (2016)	https://zenodo.org/record/6397990/files/figure.png	105. Long-toothed Pipistrelle Hypsugo dolichodon French: Vespere a longues dents / German: Langzahn-Zwergfledermaus / Spanish: Hypsugo de dientes largos Taxonomy. Hypsugo dolichodon Gorfol et al., 2014 , “ 14°57'N , 107°08'E , Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l.” The relationships of Hypsugo dolichodon are uncertain, but it appears to be close to H. petersi and H. pulveratus , based on limited genetic data. Specimens from Cambodia and Myanmar were originally identified as H. affinis , but are now considered to represent this species. Monotypic. Distribution. Known from scattered localities in SE Myanmar , W & S Cambodia , SE Laos , and S Vietnam . Descriptive notes. Hindfoot 5-9-7 mm, forearm 35-2-38-4 mm. No other specific measurements are available. Dorsal pelage of the Long-toothed Pipistrelle is blackish brown (hairs with paler tips), and ventral pelage lighter (hairs blackish brown on basal one-half and dirty white on distal one-half). Bare portions of face, ears, limbs, and wings are nearly black. Ears are relatively short and wide and somewhat rounded at tip; tragus is short and squarish. Tail extends a little beyond margin of uropatagium; calcar extends halfway to tail from ankle, and has narrow Postcalcarial lobe—Tless defined than in the Chinese Pipistrelle ( Hypsugo pulveratus ). Baculum has more developed base with enlarged lateral projection, narrowing shaft, and shallower ventral depression on tip when compared to the Chocolate Pipistrelle ( H. affinis ). Skull is large with relatively robust, long, narrow rostrum; rostral profile of skull is almost flat; braincase is flattened; there is no sagittal crest; lambdoid crest is pronounced; and basioccipital pits are shallow but visible. I* is bicuspid; C' and C, are relatively slender and long; P? is small and intruded but C' and P* are still separated; and lower molars are myotodont. Habitat. The Long-toothed Pipistrelle has been reported in lowland evergreen rainforest at elevations of 50-400 m throughout much ofits range, and is associated with limestone outcrops. Reported at 978 m elevation in limestone caves surrounded by evergreen rainforest and cultivated areas in Cambodia . Food and Feeding. No information. Breeding. No information. Activity patterns. Long-toothed Pipistrelles roost in caves. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCN Red List. The Long-toothed Pipistrelle is known only from a few localities and does not seem to be very common. It may be threatened by habitat destruction due to limestone extraction, and general human disturbance. Further studies are needed, particularly into general ecology and threats. Bibliography. Gorfol & Csorba (2018), Gorfél, Csorba et al. (2014), Gorfol, Furey et al. (2019), Lim, L.S. et al. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hypsugo dolichodon	Hypsugo		dolichodon	G&ouml;rf&ouml;l Csorba, Eger, Son & Francis	2014	0	Zootaxa	3887(2): 241	Long-toothed Pipistrelle	None.	Laos, Attapu Province, 10 km E from Ban Paam, Xe Kaman proposed dam site.	Laos and S Vietnam	Not listed (new species).	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hypsugo dolichodon	23	Long-toothed Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	dolichodon	GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & Francis	2014	0	Hypsugo_dolichodon	GÃ¶rfÃ¶l, T., Csorba, G., Eger, J., Nguyen Truong Son, & Francis, C. M. (2014). Canines make the difference: a new species of Hypsugo (Chiroptera: Vespertilionidae) from Laos and Vietnam. Zootaxa, 3887(2), 241.	https://www.mapress.com/j/zt/article/view/zootaxa.3887.2.6	ROM 110459		"14Â°57'N, 107Â°08'E, Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l."	14.95	107.13	dolichodon GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & Francis, 2014	NA	NA	Myanmar|Cambodia|Laos|Vietnam	Asia	Indomalaya	DD	0	0	0	Hypsugo_dolichodon	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Hypsugo dolichodon	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hypsugo	dolichodon	GÃ¶rfÃ¶l, Csorba, Eger, Truong-Son &; Francis, 2014	The taxon was described in 2014 based on specimens from Lao PDR and Vietnam. It belongs to the â€œpulveratus -groupâ€ of Hypsugo .	90000000	Hypsugo dolichodon	Data Deficient		2019	2016-07-02 00:00:00 UTC	3.1	English	Listed as Data Deficient as it is known only from about 20 specimens and the information on its distribution, abundance, ecology or threats is very limited.	<span lang="EN-US">The Lao specimens were captured in mist nets near Xe Kaman, a moderately large, fast-flowing river 50â€“100 m across, flowing through lowland evergreen rainforest in the Dong Amphan National Biodiversity Area. All of the Vietnamese specimens came from Cat Tien National Park. One specimen was collected near the headquarters of the Park, close to the Dong Nai River. This area is generally described as lowland rain forest with dipterocarp trees (GÃ¶rfÃ¶l et al. 2014). <span lang="EN-US">Hypsugo dolichodon is closely associated with cave ecosystems. A single specimen was collected from Saddan-Sin Cave in Mon State, Myanmar (identified as Falsistrellus affinis by Bates et al. , 2005). The cave is situated in a large isolated limestone outcrop and is surrounded by paddy fields and toddy palms. Two additional specimens were collected at Indian Single Rock Temple Cave. Several forested limestone outcrops surrounded by rice fields are present in the surrounding area. Remnants of evergreen, deciduous and mangrove forests are also present (GÃ¶rfÃ¶l et al. 2018).<span lang="EN-US"> <span lang="EN-US">Aside from a single male caught in evergreen forest, the five other specimens from Phnom Kulen National Park, Siem Reap Province, Cambodia were caught at a southwest-facing cave entrance in an area of semi-evergreen forest with scattered rocky outcrops and boulders. All three specimens from Kampot were caught at the entrance of Bat Khteas Cave, Chhngauk Hill, Kampot Province. The isolated karst hill where the cave is located is surrounded by seasonal wet rice cultivation, with many local farmers also cultivating small amounts of durian, banana, mango, coconuts, guava and cashew (Chheang et al. 2013, Phauk et al. 2013).</span></span>	The species is at least partly a cave-dweller, hence the disturbance of its roosting sites may be a threat. Also, one of the localities is a riverine habitat and it may be destroyed by dam construction works.	The species is known from only about 20 specimens from four countries and seven localities.	Unknown	The known distribution of this species is limited to Laos and southern Vietnam (GÃ¶rfÃ¶l et al. 2014), Cambodia and Myanmar (GÃ¶rfÃ¶l et al. 2018). It has been found at Bat Khteas Cave, Chhngauk Hill, Kampot Province and in Phnom Kulen National Park, Siem Reap Province, both in Cambodia (Chheang et al. 2013, Phauk et al. 2013). It is suspected that the species is more widely distributed in the Indo-Burma subregion (GÃ¶rfÃ¶l et al. 2018).		Terrestrial	The species occurs in three protected areas, but this not always ensures its protection. One of its localities may be destroyed by constructional works of a river dam.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hypsugo		dolichodon	G&ouml;rf&ouml;l Csorba, Eger, Son & Francis	2014	0	Zootaxa	3887(2): 241	Long-toothed Pipistrelle	None.	Laos, Attapu Province, 10 km E from Ban Paam, Xe Kaman proposed dam site.	Laos and S Vietnam	Not listed (new species).	Data Deficient		Hypsugo dolichodon	1005717	23	Long-toothed Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	dolichodon	GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & Francis	2014	0	Hypsugo_dolichodon	GÃ¶rfÃ¶l, T., Csorba, G., Eger, J., Nguyen Truong Son, & Francis, C. M. (2014). Canines make the difference: a new species of Hypsugo (Chiroptera: Vespertilionidae) from Laos and Vietnam. Zootaxa, 3887(2), 241.	https://www.mapress.com/j/zt/article/view/zootaxa.3887.2.6	ROM 110459		"14Â°57'N, 107Â°08'E, Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l."	14.95	107.1333	dolichodon GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & Francis, 2014	NA	NA				Myanmar|Cambodia|Laos|Vietnam	Asia	Indomalaya	DD	0	0	0	Hypsugo_dolichodon	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hypsugo_dolichodon	1005717	23	Long-toothed Pipistrelle		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo	NA	dolichodon	T. GÃ¶rfÃ¶l, Csorba, Eger, Nguyen Truong Son, & C. M. Francis	0	Hypsugo dolichodon	GÃ¶rfÃ¶l, T., Csorba, G., Eger, J.L., Son, N.T. and Francis, C.M. 2014-11-24. Canines make the difference: a new species of _Hypsugo_ (Chiroptera: Vespertilionidae) from Laos and Vietnam. Zootaxa 3887(2):239-250.	https://doi.org/10.11646/zootaxa.3887.2.6	ROM:MAM:110459	holotype	http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-110459	"14Â°57'N, 107Â°08'E, Xe Kaman proposed dam site, 10 km east from Ban Paam, Attapu Province, Lao PDR, 150 m a.s.l."	14.95	107.1333	NA	NA				Myanmar|Cambodia|Laos|Vietnam	Asia	Indomalaya	DD	0	0	0	Hypsugo_dolichodon	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hypsugo		dolichodon	G&ouml;rf&ouml;l Csorba, Eger, Son & Francis	2014	0	Zootaxa	3887(2): 241	Long-toothed Pipistrelle	None.	Laos, Attapu Province, 10 km E from Ban Paam, Xe Kaman proposed dam site.	Laos and S Vietnam	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85201719/85201728/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hypsugo dolichodon; Hypsugo dolichodon; Hypsugo dolichodon; Hypsugo dolichodon; Hypsugo dolichodon; dolichodon; Vespere a longues dents; Langzahn-Zwergfledermaus; Hypsugo de dientes largos; Long-toothed Pipistrelle; Long-toothed Pipistrelle; H. dolichodon
