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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L468	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Pipistrellus ariel	Pipistrellus ariel	Pipistrellus ariel	Pipistrellus ariel	Pipistrellus ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel	Hypsugo ariel		[MSW2] Subgenus Hypsugo.; [MSW3] Reviewed by Harrison and Bates (1991). May include arabicus and bodenheimeri, see Benda et al. (2002).; [HMW] Pipistrellus ariel Thomas, 1904 , “Eastern Egyptian desert, lat. 22° N ., long. 35° E . Lat. [ Sudan ]. 2000 feet [= 610 m ].” The genus Hypsugo is often subsumed into Pipistrellus , and several species have repeatedly been switched around as belonging to Pipistrellus , Neoromicia , or Hypsugo . Genetic data place the currently listed members of Hypsugo within Vespertilionini along with Neoromicia but to the exclusion of Pipustrellus (which is in Pipistrellini). Hypsugo as currently defined remains paraphyletic and, based on limited species sampling, Eurasian species (true Hypsugo ) appear to form a monophyletic clade sister to the Australasian genera of Vespertilionini , whereas the African species may be better placed in Nycticeinops ( H. eisentrauti and H. crassulus ) or Neoromicia (H. anchieta and H. bemainty ). These projected transfers are withheld for the present, as to date not all African species of Hypsugo ( H. musciculus ) and Neoromicia have been assessed phylogenetically, so the full picture is not yet clear. Hypsugo ariel is often placed in Pipustrellus, but genetic data clearly situate it in Hypsugo , within the Eurasian (true Hypsugo ) clade, close to H. arabicus and H. savii . Hypsugo ariel includes bodenheimeri as a synonym, based on genetic and morphological data. Monotypic.; [batnames2022] Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).; [MDD2022] includes bodenheimeri; [batnames2023] Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).; [MDD2023] includes bodenheimeri; [MDD2025_2.0] includes bodenheimeri; [batnames2025_1.7] Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).; [MDD2025_2.2] includes bodenheimeri											bodenheimeri	ariel 	ariel - bodenheimeri	ariel, bodenheimeri		ariel 	ariel - bodenheimeri 	ariel, bodenheimeri 	ariel, bodenheimeri 	ariel	ariel - bodenheimeri	ariel (O. Thomas, 1904)|bodenheimeri (D. L. Harrison, 1960)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Desert pipistrelle	Egypt, Sudan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Pipistrellus ariel	Sudan, Kassala Province, Wadi Alagi (22° N. and 35° E.), 2000 ft. (610 m).	Thomas	1904	Ann. Mag. Nat. Hist., ser. 7, 14:157.	Distribution: A poorly known species apparently ranging from extreme southern Egypt to central Sudan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Desert pipistrelle	Egypt, Sudan, Israel; ref. 4.130	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1904	Ann. Mag. Nat. Hist., ser. 7, 14:157.	Subgenus Hypsugo.	Egypt, N Sudan.	Sudan, Kassala Province, Wadi Alagi (22"N, 35°E), 2,000 ft. (610 m).		THOMAS	1904	Size relatively small (forearm length, 28-31 mm). Inner upper incisor unicuspid. Anterior upper premolar greatly reduced and displaced medially. Rostrum relatively slender. Forehead distinctly concave.	Distribution: A poorly known species apparently ranging from extreme southern Egypt to central Sudan.	No subspecies.		115	species	P. ariel	THOMAS	1904	Pipistrellus	subgenus	Pipistrellus ariel				Size relatively small (forearm length, 28-31 mm). Inner upper incisor unicuspid. Anterior upper premolar greatly reduced and displaced medially. Rostrum relatively slender. Forehead distinctly concave.	No subspecies.		38. P. ariel THOMAS 1904 [savii group].	38	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo ariel	Hypsugo		ariel	Thomas	y	1904		Ann. Mag. Nat. Hist., ser. 7	14		157		Fairy Pipistrelle	Sudan, Kassala Province, Wadi Alagi (22°N, 35°E), 2,000 ft. (610 m).	Israel, Jordan, N Sudan, possibly Egypt.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable as Pipistrellus ariel.		Reviewed by Harrison and Bates (1991). May include arabicus and bodenheimeri, see Benda et al. (2002).	4C3D87E8FFC86A77FF8197F118B1BE07	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	809	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFC86A77FF8197F118B1BE07.xml	Hypsugo ariel	Vespertilionidae	Hypsugo	ariel		1904	Vespére ariel @fr | Elfen-Zwergfledermaus @de | Hypsugo ariel @es | Ariel Pipistrelle @en | Bodenheimer's Pipistrelle @en | Desert Pipistrelle @en | Egyptian Desert Pipistrelle @en	Pipistrellus ariel Thomas, 1904 , “Eastern Egyptian desert, lat. 22° N ., long. 35° E . Lat. [ Sudan ]. 2000 feet [= 610 m ].” The genus Hypsugo is often subsumed into Pipistrellus , and several species have repeatedly been switched around as belonging to Pipistrellus , Neoromicia , or Hypsugo . Genetic data place the currently listed members of Hypsugo within Vespertilionini along with Neoromicia but to the exclusion of Pipustrellus (which is in Pipistrellini). Hypsugo as currently defined remains paraphyletic and, based on limited species sampling, Eurasian species (true Hypsugo ) appear to form a monophyletic clade sister to the Australasian genera of Vespertilionini , whereas the African species may be better placed in Nycticeinops ( H. eisentrauti and H. crassulus ) or Neoromicia (H. anchieta and H. bemainty ). These projected transfers are withheld for the present, as to date not all African species of Hypsugo ( H. musciculus ) and Neoromicia have been assessed phylogenetically, so the full picture is not yet clear. Hypsugo ariel is often placed in Pipustrellus, but genetic data clearly situate it in Hypsugo , within the Eurasian (true Hypsugo ) clade, close to H. arabicus and H. savii . Hypsugo ariel includes bodenheimeri as a synonym, based on genetic and morphological data. Monotypic.	Palestine , S Israel , SWJordan, NE Egypt ( Sinai ), NE & SC Sudan , W Saudi Arabia , Yemen , and SW Oman ; it may occur in SE Egypt , and there is a purported record from C Kenya , but these need verification.	Head-body 35-45 mm, tail 32-44 mm, ear 9-14-1 mm, hindfoot 5-6 mm, forearm 28-1-33-7 mm; weight 2:3—4-8 g. Pelage of the Fairy Pipistrelle is not dense; dorsally it is pale pinkish brown to pale grayish cream or off-white cream, with underlying color showing through (hairs are blackish brown basally); ventrally it is generally paler brownish cream to off-white (hairs with blackish-brown base and pale grayish-cream tip). Bare face, ears, limbs, and digits are pinkish to dark pinkish brown. Ears are relatively short with convex inner margins, concave outer margins, and rounded tip; tragus has maximum breadth thatis only slightly less than length of its anterior margin, straight anterior margin, smoothly convex posterior margin with distinct lobule basally, and rounded tip. Membranes are pale brown and semitranslucent (uropatagium is apparently lighter than wings); Postcalcarial lobe is very narrow and tail projects slightly beyond uropatagium (last two vertebrae). Penisis relatively short with glans covered in long fine hairs. Baculum is short (1-4-1-8 mm) with upturned tip; shaft is wide in dorsal view, gradually widening just before narrowed and pointed tip, and thinner in lateral view; base is bulbous and non-bifurcated. Skull is small and gracile; braincase is relatively narrow and moderately high; rostrum is flattened and relatively narrow; forehead when viewed laterally is strongly concave; sagittal crest is extremely reduced and lambdoidal crests are absent. I” varies between unicuspid and bicuspid; I? is about two-thirds length of I?; P* is minute and sometimes not visible above gum and displaced lingually; C' and P* are in contact; and lower molars are myotodont. Dental formula for all members of Hypsugois 12/3, C 1/1, P 2/2, M 3/3 (x2) = 34 (but see H. alaschanicus and H. musciculus ).	Rocky areas, deserts, and semi-deserts, where it often occurs around oases and other water bodies. Also known to occur around farms and irrigated areas in Jordan . Recorded at elevations of 345-1224 m.	Foraging individuals were captured in Jordan in a rocky canyon, at a small spring, and in an old village garden, suggesting Fairy Pipistrelles mostly forage in open areas, although they have also been observed foraging around trees. They seem to fly close to the ground while foraging and are fairly maneuverable. Fifty-five fecal pellets in Jordan contained a variety of insects, including Coleoptera , Lepidoptera , Auchenorrhyncha , Heteroptera , Hymenoptera , Nematocera , Neuroptera , and Trichoptera, being dominated by small Lepidoptera in Al Ghal, and Coleoptera in Wadi Al Hasa. Digestive tracts in Sinai , Egypt , showed high proportions (80% by volume) of small Coleoptera (including 60% Scarabaeidae and 20% Tenebrionidae ), along with smaller amounts of Brachycera (10%), Lepidoptera (7%), and Auchenorrhyncha (3%). Fairy Pipistrelles appear to shift their diet based on availability. They have also been recorded feeding on arachnids, which suggests that they may glean prey occasionally.	In Jordan , a single pregnant female was captured in May with two well-developed fetuses, suggesting that births probably occur around late May in that region. In Palestine and Israel , the Fairy Pipistrelle appears to give birth in late April to May, as a pregnant female was captured in April with two embryos, and lactating females were captured from May to July. In Israel , subadults were first captured in June and made up 50% of individuals captured in July and August.	The Fairy Pipistrelle is known to roost in cracks and crevices in rocks during the day, foraging by night. In the Negev HighlandsofIsrael, this species expressed a bimodal nightly activity pattern, being most active during the first and last hoursof the night. It is active throughout the year, but during a summer night, there may be as many as 20 times more individuals active than on winter nights. Fairy Pipistrelles hibernate during winter, but arouse to forage and have been captured at temperatures as low as 11°C. Search-call shapeis a steep FM/QCFsweep with a very short CF portion; records fromJordan had start frequencies of47-7-75 kHz, end frequencies of 30-47-4 kHz, peak frequencies of 43-4—49 kHz, durations of 2-4-7-2 milliseconds, and interpulse intervals of 45-193 milliseconds. In Sinai , Egypt , start frequencies were 48-3-87 kHz, end frequencies 41-6-45-7 kHz, peak frequencies 44-3—49-1 kHz, durations 2-6-5-9 milliseconds, and interpulse intervals 31-8-186 milliseconds. In Israel , start frequencies were 45-83.9 kHz (mean 56-3 kHz), end frequencies 42-9—49-8 kHz (mean 46-1 kHz), peak frequencies 42-8-53-7 kHz (mean 48-2 kHz), and duration averaged 3-2 milliseconds.	A colony of 20 Fairy Pipistrelles was observed leavingits castle ruin roost in Saudi Arabia . This species probably roosts in relatively small groups or solitarily.	Classified as Data Deficient on The IUCN Red List (as Pipistrellus ariel ). The full distribution of the species is still uncertain, and very little is known of threats. It may be threatened by agricultural expansion and general habitat loss.	ACR (2018) | Aloufi et al. (2016) | Benda & Aulagnier (2008) | Benda, Al-Jumaily et al. (2011) | Benda, Dietz et al. (2008) | Benda, Lucan et al. (2010) | Benda, Nasheret al. (2017) | Benda, Reiter et al. (2016) | Gaucher & Harrison (1995) | Hackett et al. (2017) | Makin & Harrison (1988) | Mayer et al. (2007) | Razgour et al. (2011) | Riskin (2001) | Van Cakenberghe & Happold (2013f) | Whitaker et al. (1994) | Yom-Tov (1993) | Yom-Tov & Kadmon (1998) | Yom-Tov etal. (1992)	https://zenodo.org/record/6397968/files/figure.png	94. Fairy Pipistrelle Hypsugo ariel French: Vespére ariel / German: Elfen-Zwergfledermaus / Spanish: Hypsugo ariel Other common names: Ariel Pipistrelle , Bodenheimer's Pipistrelle , Desert Pipistrelle , Egyptian Desert Pipistrelle Taxonomy. Pipistrellus ariel Thomas, 1904 , “Eastern Egyptian desert, lat. 22° N ., long. 35° E . Lat. [ Sudan ]. 2000 feet [= 610 m ].” The genus Hypsugo is often subsumed into Pipistrellus , and several species have repeatedly been switched around as belonging to Pipistrellus , Neoromicia , or Hypsugo . Genetic data place the currently listed members of Hypsugo within Vespertilionini along with Neoromicia but to the exclusion of Pipustrellus (which is in Pipistrellini). Hypsugo as currently defined remains paraphyletic and, based on limited species sampling, Eurasian species (true Hypsugo ) appear to form a monophyletic clade sister to the Australasian genera of Vespertilionini , whereas the African species may be better placed in Nycticeinops ( H. eisentrauti and H. crassulus ) or Neoromicia (H. anchieta and H. bemainty ). These projected transfers are withheld for the present, as to date not all African species of Hypsugo ( H. musciculus ) and Neoromicia have been assessed phylogenetically, so the full picture is not yet clear. Hypsugo ariel is often placed in Pipustrellus, but genetic data clearly situate it in Hypsugo , within the Eurasian (true Hypsugo ) clade, close to H. arabicus and H. savii . Hypsugo ariel includes bodenheimeri as a synonym, based on genetic and morphological data. Monotypic. Distribution. Palestine , S Israel , SWJordan, NE Egypt ( Sinai ), NE & SC Sudan , W Saudi Arabia , Yemen , and SW Oman ; it may occur in SE Egypt , and there is a purported record from C Kenya , but these need verification. Descriptive notes. Head-body 35-45 mm, tail 32-44 mm, ear 9-14-1 mm, hindfoot 5-6 mm, forearm 28-1-33-7 mm; weight 2:3—4-8 g. Pelage of the Fairy Pipistrelle is not dense; dorsally it is pale pinkish brown to pale grayish cream or off-white cream, with underlying color showing through (hairs are blackish brown basally); ventrally it is generally paler brownish cream to off-white (hairs with blackish-brown base and pale grayish-cream tip). Bare face, ears, limbs, and digits are pinkish to dark pinkish brown. Ears are relatively short with convex inner margins, concave outer margins, and rounded tip; tragus has maximum breadth thatis only slightly less than length of its anterior margin, straight anterior margin, smoothly convex posterior margin with distinct lobule basally, and rounded tip. Membranes are pale brown and semitranslucent (uropatagium is apparently lighter than wings); Postcalcarial lobe is very narrow and tail projects slightly beyond uropatagium (last two vertebrae). Penisis relatively short with glans covered in long fine hairs. Baculum is short (1-4-1-8 mm) with upturned tip; shaft is wide in dorsal view, gradually widening just before narrowed and pointed tip, and thinner in lateral view; base is bulbous and non-bifurcated. Skull is small and gracile; braincase is relatively narrow and moderately high; rostrum is flattened and relatively narrow; forehead when viewed laterally is strongly concave; sagittal crest is extremely reduced and lambdoidal crests are absent. I” varies between unicuspid and bicuspid; I? is about two-thirds length of I?; P* is minute and sometimes not visible above gum and displaced lingually; C' and P* are in contact; and lower molars are myotodont. Dental formula for all members of Hypsugois 12/3, C 1/1, P 2/2, M 3/3 (x2) = 34 (but see H. alaschanicus and H. musciculus ). Habitat. Rocky areas, deserts, and semi-deserts, where it often occurs around oases and other water bodies. Also known to occur around farms and irrigated areas in Jordan . Recorded at elevations of 345-1224 m. Food and Feeding. Foraging individuals were captured in Jordan in a rocky canyon, at a small spring, and in an old village garden, suggesting Fairy Pipistrelles mostly forage in open areas, although they have also been observed foraging around trees. They seem to fly close to the ground while foraging and are fairly maneuverable. Fifty-five fecal pellets in Jordan contained a variety of insects, including Coleoptera , Lepidoptera , Auchenorrhyncha , Heteroptera , Hymenoptera , Nematocera , Neuroptera , and Trichoptera, being dominated by small Lepidoptera in Al Ghal, and Coleoptera in Wadi Al Hasa. Digestive tracts in Sinai , Egypt , showed high proportions (80% by volume) of small Coleoptera (including 60% Scarabaeidae and 20% Tenebrionidae ), along with smaller amounts of Brachycera (10%), Lepidoptera (7%), and Auchenorrhyncha (3%). Fairy Pipistrelles appear to shift their diet based on availability. They have also been recorded feeding on arachnids, which suggests that they may glean prey occasionally. Breeding. In Jordan , a single pregnant female was captured in May with two well-developed fetuses, suggesting that births probably occur around late May in that region. In Palestine and Israel , the Fairy Pipistrelle appears to give birth in late April to May, as a pregnant female was captured in April with two embryos, and lactating females were captured from May to July. In Israel , subadults were first captured in June and made up 50% of individuals captured in July and August. Activity patterns. The Fairy Pipistrelle is known to roost in cracks and crevices in rocks during the day, foraging by night. In the Negev HighlandsofIsrael, this species expressed a bimodal nightly activity pattern, being most active during the first and last hoursof the night. It is active throughout the year, but during a summer night, there may be as many as 20 times more individuals active than on winter nights. Fairy Pipistrelles hibernate during winter, but arouse to forage and have been captured at temperatures as low as 11°C. Search-call shapeis a steep FM/QCFsweep with a very short CF portion; records fromJordan had start frequencies of47-7-75 kHz, end frequencies of 30-47-4 kHz, peak frequencies of 43-4—49 kHz, durations of 2-4-7-2 milliseconds, and interpulse intervals of 45-193 milliseconds. In Sinai , Egypt , start frequencies were 48-3-87 kHz, end frequencies 41-6-45-7 kHz, peak frequencies 44-3—49-1 kHz, durations 2-6-5-9 milliseconds, and interpulse intervals 31-8-186 milliseconds. In Israel , start frequencies were 45-83.9 kHz (mean 56-3 kHz), end frequencies 42-9—49-8 kHz (mean 46-1 kHz), peak frequencies 42-8-53-7 kHz (mean 48-2 kHz), and duration averaged 3-2 milliseconds. Movements, Home range and Social organization. A colony of 20 Fairy Pipistrelles was observed leavingits castle ruin roost in Saudi Arabia . This species probably roosts in relatively small groups or solitarily. Status and Conservation. Classified as Data Deficient on The IUCN Red List (as Pipistrellus ariel ). The full distribution of the species is still uncertain, and very little is known of threats. It may be threatened by agricultural expansion and general habitat loss. Bibliography. ACR (2018), Aloufi et al. (2016), Benda & Aulagnier (2008), Benda, Al-Jumaily et al. (2011), Benda, Dietz et al. (2008), Benda, Lucan et al. (2010), Benda, Nasheret al. (2017), Benda, Reiter et al. (2016), Gaucher & Harrison (1995), Hackett et al. (2017), Makin & Harrison (1988), Mayer et al. (2007), Razgour et al. (2011), Riskin (2001), Van Cakenberghe & Happold (2013f), Whitaker et al. (1994), Yom-Tov (1993), Yom-Tov & Kadmon (1998), Yom-Tov etal. (1992).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Hypsugo ariel	Hypsugo		ariel	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 14: 157	Fairy Pipistrelle	 bodenheimeri Harrison 1960.	Sudan, Kassala Province, Wadi Alagi (22°N, 35°E), 2,000 ft. (610 m).	Israel, Jordan, N Sudan, possibly Egypt.	Not listed.	Data Deficient	Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hypsugo ariel	23	Fairy Pipistrelle	Ariel Pipistrelle|Bodenheimer's Pipistrelle|Desert Pipistrelle|Egyptian Desert Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	ariel	O. Thomas	1904	1	Pipistrellus_ariel	Thomas, O. (1904). On some small mammals collected by Mr. A. M. Mackilligin in the Eastern desert of Egypt. Annals and Magazine of Natural History, Ser. 7, 14, 157.	https://www.biodiversitylibrary.org/item/88431#page/175/mode/1up	BM 1904.11.4.7		"Eastern Egyptian desert, lat. 22° N., long. 35° E. Lat. [Sudan]. 2000 feet [= 610 m]."	22	35	ariel (O. Thomas, 1904)|bodenheimeri (D. L. Harrison, 1960)	includes bodenheimeri	Benda, P., Dietz, C., Andreas, M., Hotový, J., Lučan, R. K., Maltby, A., ... & Vallo, P. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 6. Bats of Sinai (Egypt) with some taxonomic, ecological and echolocation data on that fauna. Acta Societatis Zoologicae Bohemicae, 72(1-2), 1-103.	Oman|Yemen|Saudi Arabia|Israel|Palestine|Jordan|Egypt|Sudan|Kenya?	Africa|Asia	Afrotropic|Palearctic	DD	0	0	0	Hypsugo_ariel	0	sciname match	Hypsugo_ariel	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	200000000	Hypsugo ariel	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Hypsugo	ariel	(Thomas, 1904)		20000000	Hypsugo ariel	Data Deficient		2020	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of recent information on its extent of occurrence, status and ecological requirements.	Inhabits rocky areas, desert and semi-desert. Occurs in oases and forages near the ground and around trees. Roosts in cracks and crevices. In Jordan this species occurs around farms and irrigated areas (Z. Amr pers. comm. 2005).	Loss of natural habitat because of agricultural intensification. The species is affected by pesticides, although this is not considered a major threat (Z. Amr pers. comm. 2005).	The population is apparently small but is probably stable (Z. Amr pers. comm. 2005).	Unknown	Recorded from Israel, Jordan and Egypt (including the Sinai) within the Mediterranean (Dietz, 2005). Outside the area the range extends to Saudi Arabia, and Yemen (including Socotra, but that may be treated as a separate species). In Africa is recorded from northern Sudan (type locality, see Simmons in Wilson and Reeder 2005). Approximately five localities known in northeast Africa, including Egypt and Sudan.		Terrestrial	None in place. Further research on the taxonomy, population ecology and trends is required and the extent of the range requires further clarification.	Afrotropical|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Hypsugo		ariel	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 14: 157	Fairy Pipistrelle	 bodenheimeri Harrison 1960.	Sudan, Kassala Province, Wadi Alagi (22°N, 35°E), 2,000 ft. (610 m).	Israel, Jordan, N Sudan, possibly Egypt.	Not listed.	Data Deficient	Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).	Hypsugo ariel	1005715	23	Fairy Pipistrelle	Ariel Pipistrelle|Bodenheimer's Pipistrelle|Desert Pipistrelle|Egyptian Desert Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	VESPERTILIONINI	Hypsugo	NA	ariel	O. Thomas	1904	1	Pipistrellus_ariel	Thomas, O. (1904). On some small mammals collected by Mr. A. M. Mackilligin in the Eastern desert of Egypt. Annals and Magazine of Natural History, Ser. 7, 14, 157.	https://www.biodiversitylibrary.org/item/88431#page/175/mode/1up	BM 1904.11.4.7		"Eastern Egyptian desert, lat. 22° N., long. 35° E. Lat. [Sudan]. 2000 feet [= 610 m]."	22	35	ariel (O. Thomas, 1904)|bodenheimeri (D. L. Harrison, 1960)	includes bodenheimeri	Benda, P., Dietz, C., Andreas, M., Hotový, J., Lučan, R. K., Maltby, A., ... & Vallo, P. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 6. Bats of Sinai (Egypt) with some taxonomic, ecological and echolocation data on that fauna. Acta Societatis Zoologicae Bohemicae, 72(1-2), 1-103.				Oman|Yemen|Saudi Arabia|Israel|Palestine|Jordan|Egypt|Sudan|Kenya?	Africa|Asia	Afrotropic|Palearctic	DD	0	0	0	Hypsugo_ariel	0	sciname match	Hypsugo_ariel	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hypsugo_ariel	1005715	23	Fairy Pipistrelle	Ariel Pipistrelle|Bodenheimer's Pipistrelle|Desert Pipistrelle|Egyptian Desert Pipistrelle	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Vespertilionini	Hypsugo	NA	ariel	O. Thomas	1	Pipistrellus ariel	Thomas, O. 1904-08-01. On some small Mammals collected by Mr. A. M. Mackilligin in the Eastern Desert of Egypt. Annals and Magazine of Natural History (7)14(80):155-159.	https://www.biodiversitylibrary.org/page/27733884	BMNH:Mamm:1904.11.4.7	holotype	https://data.nhm.ac.uk/object/f6e0d4ce-08f0-4ed4-93b0-b994bb17f28f	"Eastern Egyptian desert, lat. 22° N., long. 35° E. Lat. [Sudan]. 2000 feet [= 610 m]."	22	35	includes bodenheimeri	Benda, P., Dietz, C., Andreas, M., Hotový, J., Lučan, R. K., Maltby, A., ... & Vallo, P. (2008). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 6. Bats of Sinai (Egypt) with some taxonomic, ecological and echolocation data on that fauna. Acta Societatis Zoologicae Bohemicae, 72(1-2), 1-103.				Oman|Yemen|Saudi Arabia|Israel|Palestine|Jordan|Egypt|Sudan|Kenya?	Africa|Asia	Afrotropic|Palearctic	DD	0	0	0	Hypsugo_ariel	0	sciname match	Hypsugo_ariel	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Hypsugo		ariel	Thomas	1904	1	Ann. Mag. Nat. Hist.	ser. 7, 14: 157	Fairy Pipistrelle	bodenheimeri Harrison 1960.	Sudan, Kassala Province, Wadi Alagi (22°N, 35°E), 2,000 ft. (610 m).	Israel, Jordan, N Sudan, possibly Egypt.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/171619155/22071929/' target='_blank'>Data Deficient</a>	Includes bodenheimeri (see Mayer et al. 2007; Benda et al. 2008); Reviewed by Harrison and Bates (1991). May include arabicus; see Benda et al. (2002).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hypsugo ariel; Hypsugo ariel; Hypsugo ariel; Hypsugo ariel; Hypsugo ariel; Hypsugo ariel; bodenheimeri; bodenheimeri; ariel; bodenheimeri; Vespére ariel; Elfen-Zwergfledermaus; Hypsugo ariel; Ariel Pipistrelle; Bodenheimer's Pipistrelle; Desert Pipistrelle; Egyptian Desert Pipistrelle; Fairy Pipistrelle; Ariel Pipistrelle; Bodenheimer's Pipistrelle; Desert Pipistrelle; Egyptian Desert Pipistrelle; Fairy Pipistrelle; Fairy Pipistrelle; H. ariel