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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L462	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Lonchophylla thomasi	Lonchophylla thomasi	Lonchophylla thomasi	Lonchophylla thomasi	Lonchophylla thomasi	Lonchophylla thomasi	Hsunycteris thomasi	Hsunycteris thomasi	Hsunycteris thomasi	Lonchophylla thomasi	Hsunycteris thomasi	Hsunycteris thomasi	Hsunycteris thomasi	Hsunycteris thomasi	Hsunycteris thomasi		[MSW2] Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978:1, 2) and Koopman (1978b).; [MSW3] Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978) and Koopman (1978b).; [HMW] Lonchophylla thomas J. A. Allen, 1904 , “Cuidad [sic] Bolivar ,” Venezuela . Hsunycteris thomas : 1s considered paraphyletic to H. pattoni , but no formal subdivisions (subspecies) have been recognized. It is also a nomenclatural matter because both genetic lineages are found in Bolivar State near type locality. Monotypic.; [batnames2022] Specimens of this species have frequently been confused with concava , mordax , and Lionycteris  spurrelli ; see Taddei et al. (1978) and Koopman (1978).; [MDD2022] moved from Lonchophylla to the recently described Hsunycteris; [IUCN] Specimens of this species have frequently been confused with concava , mordax ;and Lionycteris spurrelli .; [batnames2023] Specimens of this species have frequently been confused with concava , mordax , and Lionycteris  spurrelli ; see Taddei et al. (1978) and Koopman (1978).; [MDD2023] moved from Lonchophylla to the recently described Hsunycteris; [MDD2025_2.0] moved from Lonchophylla to the recently described Hsunycteris; [batnames2025_1.7] Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978) and Koopman (1978).; [MDD2025_2.2] moved from Lonchophylla to the recently described Hsunycteris														thomasi	Specimens of this species have frequently been confused with concava , mordax ;and Lionycteris spurrelli .			thomasi	thomasi			thomasi (J. A. Allen, 1904)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Panama, Peru, Bolivia, Brazil, Surinam	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Lonchophylla thomasi	Venezuela, Bolivar, Ciudad Bolivar.	J. A. Allen	1904	Bull. Am. Mus. Nat. Hist., 20:230.	Distribution: Ranging from eastern Panama to eastern Peru and Amazonian Brazil but west of the Andes not south of Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Thomas's long-tongued bat	Panama, Peru, Bolivia, Brazil, Guianas	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	J. A. Allen	1904	Bull. Am. Mus. Nat. Hist., 20:230.	Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978:1, 2) and Koopman (1978b).	E Panama, Colombia, Venezuela, Guianas, Amazonian Brazil, Peru, Bolivia.	Venezuela, Bolivar, Ciudad Bolivar.		J. A. ALLEN	1904	Size relatively small (forearm length, 31-34 mm; condylobasal length, 18-21 mm; maxillary toothrow length, 6.2-7.0 mm). Lingual cusp oflast upper premolar well-developed, the tooth thus T-shaped. Posterior cusp of anterior lower premolar well developed and hooklike. Posterior palatal emargination V-shaped.	Distribution: Ranging from eastern Panama to eastern Peru and Amazonian Brazil but west of the Andes not south of Ecuador.	No subspecies.		78	species	L. thomasi	J. A. ALLEN	1904	Lonchophylla	genus	Lonchophylla thomasi				Size relatively small (forearm length, 31-34 mm; condylobasal length, 18-21 mm; maxillary toothrow length, 6.2-7.0 mm). Lingual cusp oflast upper premolar well-developed, the tooth thus T-shaped. Posterior cusp of anterior lower premolar well developed and hooklike. Posterior palatal emargination V-shaped.	No subspecies.		1. L. thomasi J. A. ALLEN 1904.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Glossophaginae	Lonchophyllini	Lonchophylla thomasi	Lonchophylla		thomasi	J. A. Allen		1904		Bull. Am. Mus. Nat. Hist.	20		230		Thomas's Nectar Bat	Venezuela, Bolivar, Ciudad Bolivar.	E Panama, Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, Peru, Bolivia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978) and Koopman (1978b).	03A687BCFF9DFF9D13FAFCE1FC6EF951	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	528	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF9DFF9D13FAFCE1FC6EF951.xml	Hsunycteris thomasi	Phyllostomidae	Hsunycteris	thomasi		1904	Lonchophylle de Thomas @fr | Thomas-Nektarfledermaus @de | Sunicterio de Thomas @es | Thomas's Small Nectar Bat @en	Lonchophylla thomas J. A. Allen, 1904 , “Cuidad [sic] Bolivar ,” Venezuela . Hsunycteris thomas : 1s considered paraphyletic to H. pattoni , but no formal subdivisions (subspecies) have been recognized. It is also a nomenclatural matter because both genetic lineages are found in Bolivar State near type locality. Monotypic.	C & E Panama , Colombia , Venezuela (mostly S of Orinoco River), the Guianas, Amazonian Brazil , E Ecuador , E Peru , and Bolivia .	Head-body 45-56 mm, tail 4-11 mm, ear 11-18 mm, hindfoot 7-11 mm, forearm 29-7-34-3 mm; weight 5-10 g. Thomas’s Nectar Bat is among the smaller species in the subfamily. Dorsal pelage is dark brown to clove-brown, with hairs 5-7 mm and basal two-thirds whitish. Ventral coloris similar to dorsum but slightly paler; it generally darkens from abdomen to neck, and hairs are indistinctly bicolored. Ears and wing membranes are blackish brown, and wings attach to ankles. Tail membraneis only sparsely haired. Muzzle is elongated, with lower jaw extending slightly beyond upperjaw. Spear of noseleaf is relatively large (c. 5 mm tall). Forearms are naked. Rostrum is narrow and about as long as braincase. Rostrum is only slightly inflated above M', lateral outlines are gently convex, and postorbital region is inflated, typically lacking lateral projections. In palatal view, obvious gaps occur between I' and I°. I' is more than double the height of I’. Lower incisors are small but always present. Chromosomal complement has 2n = 30-36 and FN = 34-48, which suggests an accelerated chromosomal evolution.	Rainforest in lowlands and foothills up to elevations of ¢. 1300 m . Thomas's Nectar Bat prefers evergreen forests but can be found in clearings, apparently because of availability of floral resources in gardens and plantations. In French Guiana , it was caught primarily in ground-level (0-2 m) mist nets and rarely in canopy nets, suggesting thatit typically flies in lower foreststrata.	Thomas’s Nectar Bats from south-eastern Peru fed mostly on pollen and insects. Pollen of Caryocar villosum ( Caryocaraceae ), seeds of Vismia sp. ( Hypericaceae ), and beetles remains were found in stomachs of Brazilian specimens. It might feed on banana flowers.	Pregnant Thomas’s Nectar Bats have been found in February and June-October, lactating females in January-March and October, and subadults in January-February.	Thomas’s Nectar Bat roosts in tree hollows and hollow logs and on undersides of buttresses offallen trees.	Small colonies of males and females numbering up to four individuals have been found to roosting together.	Classified as Least Concern on The IUCN Red List (as Lonchophylla thomas ).	Allen (1904) | Baker et al. (1982) | Davalos & Jansa (2004) | Griffiths & Gardner (2008b) | Handley (1976) | Nogueira, Dias & Peracchi (2007) | Parlos et al. (2014) | Ribeiro et al. (2003) | Simmons & Voss (1998 Velazco et al. (2017) | Woodman (2007) | Woodman & Timm (2006)	https://zenodo.org/record/6458782/files/figure.png	86. Thomas's Nectar Bat Hsunycteris thomasi French: Lonchophylle de Thomas / German: Thomas-Nektarfledermaus / Spanish: Sunicterio de Thomas Other common names: Thomas's Small Nectar Bat Taxonomy. Lonchophylla thomas J. A. Allen, 1904 , “Cuidad [sic] Bolivar ,” Venezuela . Hsunycteris thomas : 1s considered paraphyletic to H. pattoni , but no formal subdivisions (subspecies) have been recognized. It is also a nomenclatural matter because both genetic lineages are found in Bolivar State near type locality. Monotypic. Distribution. C & E Panama , Colombia , Venezuela (mostly S of Orinoco River), the Guianas, Amazonian Brazil , E Ecuador , E Peru , and Bolivia . Descriptive notes. Head-body 45-56 mm, tail 4-11 mm, ear 11-18 mm, hindfoot 7-11 mm, forearm 29-7-34-3 mm; weight 5-10 g. Thomas’s Nectar Bat is among the smaller species in the subfamily. Dorsal pelage is dark brown to clove-brown, with hairs 5-7 mm and basal two-thirds whitish. Ventral coloris similar to dorsum but slightly paler; it generally darkens from abdomen to neck, and hairs are indistinctly bicolored. Ears and wing membranes are blackish brown, and wings attach to ankles. Tail membraneis only sparsely haired. Muzzle is elongated, with lower jaw extending slightly beyond upperjaw. Spear of noseleaf is relatively large (c. 5 mm tall). Forearms are naked. Rostrum is narrow and about as long as braincase. Rostrum is only slightly inflated above M', lateral outlines are gently convex, and postorbital region is inflated, typically lacking lateral projections. In palatal view, obvious gaps occur between I' and I°. I' is more than double the height of I’. Lower incisors are small but always present. Chromosomal complement has 2n = 30-36 and FN = 34-48, which suggests an accelerated chromosomal evolution. Habitat. Rainforest in lowlands and foothills up to elevations of ¢. 1300 m . Thomas's Nectar Bat prefers evergreen forests but can be found in clearings, apparently because of availability of floral resources in gardens and plantations. In French Guiana , it was caught primarily in ground-level (0-2 m) mist nets and rarely in canopy nets, suggesting thatit typically flies in lower foreststrata. Food and Feeding. Thomas’s Nectar Bats from south-eastern Peru fed mostly on pollen and insects. Pollen of Caryocar villosum ( Caryocaraceae ), seeds of Vismia sp. ( Hypericaceae ), and beetles remains were found in stomachs of Brazilian specimens. It might feed on banana flowers. Breeding. Pregnant Thomas’s Nectar Bats have been found in February and June-October, lactating females in January-March and October, and subadults in January-February. Activity patterns. Thomas’s Nectar Bat roosts in tree hollows and hollow logs and on undersides of buttresses offallen trees. Movements, Home range and Social organization. Small colonies of males and females numbering up to four individuals have been found to roosting together. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Lonchophylla thomas ). Bibliography. Allen (1904), Baker et al. (1982), Davalos & Jansa (2004), Griffiths & Gardner (2008b), Handley (1976), Nogueira, Dias & Peracchi (2007), Parlos et al. (2014), Ribeiro et al. (2003), Simmons & Voss (1998 Velazco et al. (2017), Woodman (2007), Woodman & Timm (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Hsunycteris thomasi	Hsunycteris		thomasi	J. A. Allen	1904	1	Bull. Am. Mus. Nat. Hist.	0.9931	Thomas's Nectar Bat	None.	Venezuela, Bolivar, Ciudad Bolivar.	E Panama, Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, Peru, Bolivia.	Not listed.	Least Concern as Lonchophylla thomasi 	Specimens of this species have frequently been confused with concava , mordax , and Lionycteris  spurrelli ; see Taddei et al. (1978) and Koopman (1978).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hsunycteris thomasi	23	Thomas's Nectar Bat	Thomas's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	thomasi	J. A. Allen	1904	1	Lonchophylla_thomasi	Allen, J. A. (1904). New bats from tropical America with note on species of Otopterus. Bulletin of the American Museum of Natural History, 20, 230.	https://digitallibrary.amnh.org/bitstream/handle/2246/668//v2/dspace/ingest/pdfSource/bul/B020a20.pdf?sequence=1&isAllowed=y	AMNH 16120		"Cuidad [sic] Bolivar," Venezuela.			thomasi (J. A. Allen, 1904)	moved from Lonchophylla to the recently described Hsunycteris	Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.	Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Hsunycteris_thomasi	0	manual	Lonchophylla_thomasi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	12269	Lonchophylla thomasi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lonchophylla	thomasi	J.A. Allen, 1904	Specimens of this species have frequently been confused with concava , mordax ;and Lionycteris spurrelli .	20000000	Lonchophylla thomasi	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is widespread, relatively common and thought to occur in large subpopulations. This species is unlikely to be declining at a rate which would qualify it for inclusion in any threatened category.	It is strongly associated with streams and moist areas. Although tolerant of man-made clearings, this species prefers multistratal tropical evergreen forests (Eisenberg and Redford 1999). It feeds on nectar, insects and probably fruit and pollen. This species probably roosts in small groups in hollow trees and in small caves (Emmons and Feer 1997).	Deforestation occurs within the range, although this is not a major threat.	It is poorly known though it may be relatively common.	Unknown	This species is distributed in South America and occurs in Bolivia, Colombia, Venezuela, the Guyana Shield, eastern Peru, Ecuador and northern Brazil (Eisenberg and Redford 1999). Principally it was caught below 850 m asl in Venezuela (Handley 1976). It is also in Panama (Handley 1976).	This species is not used.	Terrestrial	The recommended conservation action is to maintain the habitat. This species occurs in a number of protected areas throughout its range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Hsunycteris		thomasi	J. A. Allen	1904	1	Bull. Am. Mus. Nat. Hist.	0.993056	Thomas's Nectar Bat	None.	Venezuela, Bolivar, Ciudad Bolivar.	E Panama, Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, Peru, Bolivia.	Not listed.	Least Concern as Lonchophylla thomasi 	Specimens of this species have frequently been confused with concava , mordax , and Lionycteris  spurrelli ; see Taddei et al. (1978) and Koopman (1978).	Hsunycteris thomasi	1004927	23	Thomas's Nectar Bat	Thomas's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	thomasi	J. A. Allen	1904	1	Lonchophylla_thomasi	Allen, J. A. (1904). New bats from tropical America with note on species of Otopterus. Bulletin of the American Museum of Natural History, 20, 230.	https://digitallibrary.amnh.org/bitstream/handle/2246/668//v2/dspace/ingest/pdfSource/bul/B020a20.pdf?sequence=1&isAllowed=y	AMNH 16120		"Cuidad [sic] Bolivar," Venezuela.			thomasi (J. A. Allen, 1904)	moved from Lonchophylla to the recently described Hsunycteris	Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC	0	0	0	Hsunycteris_thomasi	0	manual	Lonchophylla_thomasi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hsunycteris_thomasi	1004927	23	Thomas's Nectar Bat	Thomas's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Lonchophyllinae	Hsunycterini	Hsunycteris	NA	thomasi	J. A. Allen	1	Lonchophylla thomasi	Allen, J.A. 1904-06-29. New bats from tropical America, with note on species of _Otopterus_. Bulletin of the American Museum of Natural History 20(20):227-237.	https://hdl.handle.net/2246/668	AMNH M-16120	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-16120	"Cuidad [sic] Bolivar," Venezuela.			moved from Lonchophylla to the recently described Hsunycteris	Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Brazil|Ecuador|Peru|Bolivia	North America|South America	Neotropic	LC (as Lonchophylla thomasi)	0	0	0	Hsunycteris_thomasi	0	manual	Lonchophylla_thomasi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Hsunycteris		thomasi	J. A. Allen	1904	1	Bull. Am. Mus. Nat. Hist.	0.993056	Thomas's Nectar Bat	None.	Venezuela, Bolivar, Ciudad Bolivar.	E Panama, Colombia, Venezuela, Guianas, Amazonian Brazil, Ecuador, Peru, Bolivia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/12269/22039689/' target='_blank'>Least Concern as Lonchophylla thomasi</a>	Specimens of this species have frequently been confused with concava, mordax, and Lionycteris spurrelli; see Taddei et al. (1978) and Koopman (1978).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Lonchophylla thomasi; Hsunycteris thomasi; Hsunycteris thomasi; Hsunycteris thomasi; Lonchophylla thomasi; Hsunycteris thomasi; thomasi; Lonchophylle de Thomas; Thomas-Nektarfledermaus; Sunicterio de Thomas; Thomas's Small Nectar Bat; Thomas's Nectar Bat; Thomas's Small Nectar Bat; Thomas's Nectar Bat; Thomas's Nectar Bat; H. thomasi