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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L461	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Hsunycteris pattoni	Hsunycteris pattoni	Hsunycteris pattoni	Lonchophylla pattoni	Hsunycteris pattoni	Hsunycteris pattoni	Hsunycteris pattoni	Hsunycteris pattoni	Hsunycteris pattoni		[HMW] Lonchophylla patton: Woodman & Timm, 2006 , “Reserva Cusco Amazonico (ca. 12º33' S , 69°03' W ), 200 m : el evation; north bank of the Rio Madre de Dios ; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru .” Hsunycteris pattoni was previously confused with “ Lonchophylla ” thomasi , now H. thomasi . Phylogenetic analyses of molecular data show H. thomasi is paraphyletic in relation to H. pattoni . Monotypic.; [MDD2022] recently described; moved from Lonchophylla to the recently described Hsunycteris; [MDD2023] recently described; moved from Lonchophylla to the recently described Hsunycteris; [MDD2025_2.0] recently described; moved from Lonchophylla to the recently described Hsunycteris; [MDD2025_2.2] recently described; moved from Lonchophylla to the recently described Hsunycteris														pattoni				pattoni 	pattoni 			pattoni (Woodman & Timm, 2006)						N/A																																								NA																											03A687BCFF9AFF9D165BF95DFA97F156	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	527	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF9AFF9D165BF95DFA97F156.xml	Hsunycteris pattoni	Phyllostomidae	Hsunycteris	pattoni		2006	Lonchophylle de Patton @fr | Patton-Nektarfledermaus @de | Sunicterio de Patton @es | Patton's Small Nectar Bat @en	Lonchophylla patton: Woodman & Timm, 2006 , “Reserva Cusco Amazonico (ca. 12º33' S , 69°03' W ), 200 m : el evation; north bank of the Rio Madre de Dios ; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru .” Hsunycteris pattoni was previously confused with “ Lonchophylla ” thomasi , now H. thomasi . Phylogenetic analyses of molecular data show H. thomasi is paraphyletic in relation to H. pattoni . Monotypic.	Known from scattered localities in Amazon Basin, including SE Colombia ( Amazonas Department ), W & C Brazil (Amazonas and Para states), E Ecuador (Zamora, Sucumbios , Orellana , and Pastaza provinces), E Peru ( Loreto and Madre de Dios departments), and NE Bolivia (La Paz and Beni departments).	Head-body 51-59 mm, tail 5-10 mm, ear 12-17 mm, hindfoot 7-11 mm, forearm 31-34 mm; weight 5-12 g. Patton’s Nectar Bat is among the smaller species in the subfamily. Dorsal hairs are 6-7 mm and distinctly bicolored, with paler basal two-thirds or so tending toward pale vinaceous fawn and distal segment medium chocolate-brown to reddish brown, some hairs with slightly paler tips. Venter is somewhat paler than dorsum, proximal one-half of hairs are pale, and distal segments approach yellowish brown to reddish brown. Muzzle is elongated, with lower jaw extending slightly beyond upper jaw. Dermal papillae on chin are large and not separated by basal cleft. Weakly developed central rib of noseleaf extends to tip. Ears and wing membranes are blackish brown, and wing attaches to ankle. Forearms are naked. Tail membrane is only sparsely haired. Rostrum is narrow and only slightly longer than braincase. In dorsal view of skull, rostrum is short and narrow, only slightly inflated above first upper molars; postorbital region is inflated,typically lacking lateral projections; and posterior margin of infraorbital foramen projects beyond lateral outline of rostrum. I' is more than double the size of I*; in palatal view, obvious gaps occur between I' and I?; and P* has conspicuous rooted lingual cusp. Molars are small. On following pages: 86. Thomas's Nectar Bat ( Hsunycteris thomas); 87. Chestnut Long-tongued Bat ( Lionycteris spurrell); 88. Brazilian Nectar Bat ( Lonchophylla mordax ); 89. Chocoan Nectar Bat ( Lonchophylla chocoanal; 90. Goldman's Nectar Bat ( Lonchophylla concava ); 91. Pacific Forest Long-tongued Bat ( Lonchophylla fornicata ); 92. Orces's Long-tongued Bat ( Lonchophylla orcesi ); 93. Western Nectar Bat ( Lonchophylla hesperia ); 94. Eastern Cordilleran Nectar Bat ( Lonchophylla orienticollina ); 95. Handley’s Nectar Bat ( Lonchophylla handleyi ); 96. Orange Nectar Bat ( Lonchophylla robusta ); 97. Dekeyser's Nectar Bat ( Lonchophylla dekeyseri ), 98. Pale-bellied Nectar Bat ( Lonchophylla inexpectata ); 99. Bokermann’s Nectar Bat ( Lonchophylla bokermanni ); 100. Peracchi's Nectar Bat ( Lonchophylla peracchii ); 101. Long-snouted Bat ( Platalina genovensium ); 102. Vieira's Long-tongued Bat ( Xeronycteris vieirai ).	Amazonian lowlands mostly below elevations of 300 m but up to almost 1000 m in eastern Ecuador . Specific habitats of Patton’s Nectar Bat are mainly mature and secondary rainforests but also clearings and plantations. It might prefer to fly in lower forest strata.	No information.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List (as Lonchophylla patton ). Available information about Patton’s Nectar Bat is inadequate to make reliable assessments of conservation risks.	Griffiths & Gardner (2008b) | Mantilla-Meluk, Jiménez-Ortega & Baker (2009a) | Mantilla-Meluk, Ramirez-Chaves et al. (2010) | Parlos et al. (2014) | Tirira (2012a) | Velazco et al. (2017) | Woodman & Timm (2006)	https://zenodo.org/record/6458780/files/figure.png	85. Patton’s Nectar Bat Hsunycteris pattoni French: Lonchophylle de Patton / German: Patton-Nektarfledermaus / Spanish: Sunicterio de Patton Other common names: Patton's Small Nectar Bat Taxonomy. Lonchophylla patton: Woodman & Timm, 2006 , “Reserva Cusco Amazonico (ca. 12º33' S , 69°03' W ), 200 m : el evation; north bank of the Rio Madre de Dios ; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru .” Hsunycteris pattoni was previously confused with “ Lonchophylla ” thomasi , now H. thomasi . Phylogenetic analyses of molecular data show H. thomasi is paraphyletic in relation to H. pattoni . Monotypic. Distribution. Known from scattered localities in Amazon Basin, including SE Colombia ( Amazonas Department ), W & C Brazil (Amazonas and Para states), E Ecuador (Zamora, Sucumbios , Orellana , and Pastaza provinces), E Peru ( Loreto and Madre de Dios departments), and NE Bolivia (La Paz and Beni departments). Descriptive notes. Head-body 51-59 mm, tail 5-10 mm, ear 12-17 mm, hindfoot 7-11 mm, forearm 31-34 mm; weight 5-12 g. Patton’s Nectar Bat is among the smaller species in the subfamily. Dorsal hairs are 6-7 mm and distinctly bicolored, with paler basal two-thirds or so tending toward pale vinaceous fawn and distal segment medium chocolate-brown to reddish brown, some hairs with slightly paler tips. Venter is somewhat paler than dorsum, proximal one-half of hairs are pale, and distal segments approach yellowish brown to reddish brown. Muzzle is elongated, with lower jaw extending slightly beyond upper jaw. Dermal papillae on chin are large and not separated by basal cleft. Weakly developed central rib of noseleaf extends to tip. Ears and wing membranes are blackish brown, and wing attaches to ankle. Forearms are naked. Tail membrane is only sparsely haired. Rostrum is narrow and only slightly longer than braincase. In dorsal view of skull, rostrum is short and narrow, only slightly inflated above first upper molars; postorbital region is inflated,typically lacking lateral projections; and posterior margin of infraorbital foramen projects beyond lateral outline of rostrum. I' is more than double the size of I*; in palatal view, obvious gaps occur between I' and I?; and P* has conspicuous rooted lingual cusp. Molars are small. On following pages: 86. Thomas's Nectar Bat ( Hsunycteris thomas); 87. Chestnut Long-tongued Bat ( Lionycteris spurrell); 88. Brazilian Nectar Bat ( Lonchophylla mordax ); 89. Chocoan Nectar Bat ( Lonchophylla chocoanal; 90. Goldman's Nectar Bat ( Lonchophylla concava ); 91. Pacific Forest Long-tongued Bat ( Lonchophylla fornicata ); 92. Orces's Long-tongued Bat ( Lonchophylla orcesi ); 93. Western Nectar Bat ( Lonchophylla hesperia ); 94. Eastern Cordilleran Nectar Bat ( Lonchophylla orienticollina ); 95. Handley’s Nectar Bat ( Lonchophylla handleyi ); 96. Orange Nectar Bat ( Lonchophylla robusta ); 97. Dekeyser's Nectar Bat ( Lonchophylla dekeyseri ), 98. Pale-bellied Nectar Bat ( Lonchophylla inexpectata ); 99. Bokermann’s Nectar Bat ( Lonchophylla bokermanni ); 100. Peracchi's Nectar Bat ( Lonchophylla peracchii ); 101. Long-snouted Bat ( Platalina genovensium ); 102. Vieira's Long-tongued Bat ( Xeronycteris vieirai ). Habitat. Amazonian lowlands mostly below elevations of 300 m but up to almost 1000 m in eastern Ecuador . Specific habitats of Patton’s Nectar Bat are mainly mature and secondary rainforests but also clearings and plantations. It might prefer to fly in lower forest strata. Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List (as Lonchophylla patton ). Available information about Patton’s Nectar Bat is inadequate to make reliable assessments of conservation risks. Bibliography. Griffiths & Gardner (2008b), Mantilla-Meluk, Jiménez-Ortega & Baker (2009a), Mantilla-Meluk, Ramirez-Chaves et al. (2010), Parlos et al. (2014), Tirira (2012a), Velazco et al. (2017), Woodman & Timm (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Hsunycteris pattoni	Hsunycteris		pattoni	Woodman and Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 455	Patton&apos;s Nectar Bat	None.	14 km E of Puerto Maldonado; Tambopata Province; Madre de Dios Department; Peru	Colombia, SE Peru, Bolivia	Not listed.	Data Deficient as Lonchophylla pattoni 		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hsunycteris pattoni	23	Patton's Nectar Bat	Patton's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	pattoni	Woodman & Timm	2006	1	Lonchophylla_pattoni	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 455.	https://repository.si.edu/bitstream/handle/10088/2642/2006_Lonchophylla2.pdf?sequence=1&isAllowed=y	MUSM 24350/KU 144232		"Reserva Cusco Amazónico (ca. 12°33' S, 69°03' W), 200 m elevation; north bank of the Río Madre de Dios; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru."	-12.55	-69.05	pattoni (Woodman & Timm, 2006)	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.	Colombia|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	DD	0	0	0	Hsunycteris_pattoni	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Lonchophylla pattoni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lonchophylla	pattoni	Woodman &; Timm, 2006		90000000	Lonchophylla pattoni	Data Deficient		2018	2016-11-03 00:00:00 UTC	3.1	English	The species is listed as Data Deficient because the available information is inadequate to make a reliable assessment of its risk of extinction based on its distribution and/or population status. Confirmed records exist for SE Peru and NW Bolivia, where protected areas cover part of its distribution. Further study of all records from the lowlands of western Amazonia could reveal its real distribution and threats.	This species is known from tropical lowland forests below 1,000 m in the western Amazonia. Vegetation in the type locality consists primarily of undisturbed, evergreen, lowland tropical forest transitional between Moist Tropical Forest and Dry Tropical ;Forest (Woodman and Timm 2006). In Colombia, the area enclosing the collecting locality is characterized by primary stratified seasonally inundated forest (Mantilla-Meluk et al. 2010). It seems to prefer forested areas, either primary or secondary, but with a good cover, rather than agricultural transformed land as seen in other species of the genus.	Threats to this species are unknown.	The original description (Woodman and Timm 2006) included just one specimen from the type locality. Subsequently, other records have been added from Ecuador, Colombia, Peru and Bolivia (Mantilla-Meluk et al.  2009, 2010, Parlos et al.  2014). An analysis of abundance has not been performed so far, although the species seems to be rare.	Unknown	Lonchophylla pattoni ; is known from three localities in western Amazonia, from southeastern Peru, eastern Ecuador, and southeastern Colombia (Woodman and Timm 2006, Mantilla-Meluk et al.  2009, 2010). Additional records include one locality in SE Peru and two localities from eastern Bolivia (Parlos et al.  2014). Phylogenetic analyses indicate that this and other species formerly in Lonchophylla actually belong in a new genus, ;Hsunycteris ; (Parlos et al.  2014).		Terrestrial	The lowland forest of southeastern Peru and northwestern Bolivia, where the species has been found, are protected under several natural reserves and national parks, thus providing some overall protection. However, a lot of information about general aspects of its biology needs to be compiled for a proper assessment. Further study of all records from the lowlands of western Amazonia could reveal its real distribution and threats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Hsunycteris		pattoni	Woodman and Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 455	Patton&apos;s Nectar Bat	None.	14 km E of Puerto Maldonado; Tambopata Province; Madre de Dios Department; Peru	Colombia, SE Peru, Bolivia	Not listed.	Data Deficient as Lonchophylla pattoni 		Hsunycteris pattoni	1004926	23	Patton's Nectar Bat	Patton's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	pattoni	Woodman & Timm	2006	1	Lonchophylla_pattoni	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 455.	https://repository.si.edu/bitstream/handle/10088/2642/2006_Lonchophylla2.pdf?sequence=1&isAllowed=y	MUSM 24350/KU 144232		"Reserva Cusco Amazónico (ca. 12°33' S, 69°03' W), 200 m elevation; north bank of the Río Madre de Dios; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru."	-12.55	-69.05	pattoni (Woodman & Timm, 2006)	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Colombia|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	DD	0	0	0	Hsunycteris_pattoni	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hsunycteris_pattoni	1004926	23	Patton's Nectar Bat	Patton's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Lonchophyllinae	Hsunycterini	Hsunycteris	NA	pattoni	Woodman & Timm	1	Lonchophylla pattoni	Woodman, N. and Timm, R.M. 2006-12-22. Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new _Lonchophylla_ from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington 119(4):437-476.	https://doi.org/10.2988/0006-324X(2006)119[437:CAPRON]2.0.CO;2	MUSM 24350 (= KU:M:144232)	holotype	http://portal.vertnet.org/o/ku/kum?id=96091e3f-1ed8-11e3-bfac-90b11c41863e	"Reserva Cusco Amazónico (ca. 12°33' S, 69°03' W), 200 m elevation; north bank of the Río Madre de Dios; 14 km east of Puerto Maldonado; Tambopata Province; Madre de Dios Department; southeastern Peru."	-12.55	-69.05	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Colombia|Ecuador|Peru|Bolivia|Brazil	South America	Neotropic	DD (as Lonchophylla pattoni)	0	0	0	Hsunycteris_pattoni	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Hsunycteris		pattoni	Woodman & Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 455	Patton&apos;s Nectar Bat	None.	14 km E of Puerto Maldonado; Tambopata Province; Madre de Dios Department; Peru	Colombia, SE Peru, Bolivia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88149229/88149238/' target='_blank'>Data Deficient as Lonchophylla pattoni</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hsunycteris pattoni; Hsunycteris pattoni; Hsunycteris pattoni; Lonchophylla pattoni; Hsunycteris pattoni; pattoni; Lonchophylle de Patton; Patton-Nektarfledermaus; Sunicterio de Patton; Patton's Small Nectar Bat; Patton's Nectar Bat; Patton's Small Nectar Bat; Patton&apos;s Nectar Bat; H. pattoni