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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L459	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Hsunycteris cadenai	Hsunycteris cadenai	Hsunycteris cadenai	Lonchophylla cadenai	Hsunycteris cadenai	Hsunycteris cadenai	Hsunycteris cadenai	Hsunycteris cadenai	Hsunycteris cadenai		[HMW] Lonchophylla cadena Woodman & Timm, 2006 , “Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Rio Zabaletas, across from the village of Zabaletas ( 3°44’ N , 76°57' W ); Valle del Cauca Department ; Colombia .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining evidence from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved Lonchophylla as paraphyletic. Based on these findings, they described Hsunycteris and included three named species ( L. thomasi , L. cadenai , and L. pattoni ) and one unnamed species. P. M. Velazco and collaborators in 2017 described H. dashe from north-eastern Peru ; it was previously confused with Lionycteris spurrelli . Monotypic.; [batnames2022] For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).; [MDD2022] recently described; moved from Lonchophylla to the recently described Hsunycteris; [batnames2023] For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).; [MDD2023] recently described; moved from Lonchophylla to the recently described Hsunycteris; [MDD2025_2.0] recently described; moved from Lonchophylla to the recently described Hsunycteris; [batnames2025_1.7] For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).; [MDD2025_2.2] recently described; moved from Lonchophylla to the recently described Hsunycteris														cadenai				cadenai	cadenai			cadenai (Woodman & Timm, 2006)						N/A																																								NA																											03A687BCFF9AFF9A1354FDF4FBC8F88B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	527	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFF9AFF9A1354FDF4FBC8F88B.xml	Hsunycteris cadenai	Phyllostomidae	Hsunycteris	cadenai		2006	Lonchophylle de Cadena @fr | Cadena-Nektarfledermaus @de | Sunicterio de Cadena @es | Cadena’s Small Nectar Bat @en	Lonchophylla cadena Woodman & Timm, 2006 , “Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Rio Zabaletas, across from the village of Zabaletas ( 3°44’ N , 76°57' W ); Valle del Cauca Department ; Colombia .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining evidence from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved Lonchophylla as paraphyletic. Based on these findings, they described Hsunycteris and included three named species ( L. thomasi , L. cadenai , and L. pattoni ) and one unnamed species. P. M. Velazco and collaborators in 2017 described H. dashe from north-eastern Peru ; it was previously confused with Lionycteris spurrelli . Monotypic.	Trans-Andean, known from a few localities in W Colombia ( Risaralda and Valle del Cauca departments) and NW Ecuador ( Esmeraldas Province ).	Head-body 52-58 mm, tail 7-10 mm, ear 14-15 mm, hindfoot 8-10 mm, forearm 31-32-7 mm; weight 7 g . Cadena’s Nectar Bat is among the smaller species in the subfamily Lonchophyllinae . Dorsal hairs are typically 4-7 mm, bicolored with paler light ocherous buff to ocherous tawny bases (c.60% of length) and darker tips from dark brown on lower back to cinnamon-brown near head; ventral hairs are buckthorn brown to cinnamon-brown, typically unicolored on abdomen but having darker tips near chest. Muzzle is elongated, with lowerjaw extending slightly beyond upperjaw. Weakly developed central rib of noseleaf extends to tip. Ears and wing membranes are blackish brown, and wings attach to ankles. Forearms are naked. Tail membraneis only sparsely haired. Similar to other nectarfeeding bats, rostrum is narrow but only slightly shorter than braincase. In dorsal view of skull, rostrum is short, moderately broad, and inflated above M'; lateral margins are convex; postorbital region is strongly inflated, typically with small lateral projections; and posterior margin of infraorbital foramen typically projects beyond lateral outline of rostrum. I' is more than double size of I’. In palatal view, gaps are obvious between I' and I?, and P* has conspicuous rooted lingual cusp. Dental formula for all species of Hsunycterisis12/2,C1/1,P2/3,M 3/3 ( x2 ) = 34. Chromosomal complement has 2n = 36 and FN = 50, with multiple acrocentric autosomes.	Pre-montane rainforests of Pacific coastal plain at elevations below 200 m (most records) but also up to ¢. 2400 m . Cadena’s Nectar Bat seems to prefer primary and secondary rainforests but is also found in clearings, gardens, and banana plantations. A single record comes from Andean montane forests on eastern slope of Western Andes Range. Many individuals have been caught in ground-level (0-3 m) mist nets, indicating preference to fly in low forest strata.	Cadena’s Nectar Batis primarily nectarivorous but also eats pollen, insects, and banana flowers.	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List (as Lonchophylla cadenai ). Available information is inadequate to make reliable assessments of conservation risks to Cadena’s Nectar Bat.	Baker et al. (1982) | Carrera et al. (2010) | Davalos & Jansa (2004) | Griffiths & Gardner (2008b) | Parlos et al. (2014) | Ribeiro et al. (2003) | Velazco et al. (2017) | Woodman & Timm (2006)	https://zenodo.org/record/6458776/files/figure.png	83. Cadena’s Nectar Bat Hsunycteris cadenai French: Lonchophylle de Cadena / German: Cadena-Nektarfledermaus / Spanish: Sunicterio de Cadena Other common names: Cadena’s Small Nectar Bat Taxonomy. Lonchophylla cadena Woodman & Timm, 2006 , “Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Rio Zabaletas, across from the village of Zabaletas ( 3°44’ N , 76°57' W ); Valle del Cauca Department ; Colombia .” Molecular and morphological analyses have recovered Lonchophylla (sensu lato) as a paraphyletic assemblage. Combining evidence from nuclear and mitochondrial genes, karyotypes, and skull morphology, J. A. Parlos and collaborators in 2014 also retrieved Lonchophylla as paraphyletic. Based on these findings, they described Hsunycteris and included three named species ( L. thomasi , L. cadenai , and L. pattoni ) and one unnamed species. P. M. Velazco and collaborators in 2017 described H. dashe from north-eastern Peru ; it was previously confused with Lionycteris spurrelli . Monotypic. Distribution. Trans-Andean, known from a few localities in W Colombia ( Risaralda and Valle del Cauca departments) and NW Ecuador ( Esmeraldas Province ). Descriptive notes. Head-body 52-58 mm, tail 7-10 mm, ear 14-15 mm, hindfoot 8-10 mm, forearm 31-32-7 mm; weight 7 g . Cadena’s Nectar Bat is among the smaller species in the subfamily Lonchophyllinae . Dorsal hairs are typically 4-7 mm, bicolored with paler light ocherous buff to ocherous tawny bases (c.60% of length) and darker tips from dark brown on lower back to cinnamon-brown near head; ventral hairs are buckthorn brown to cinnamon-brown, typically unicolored on abdomen but having darker tips near chest. Muzzle is elongated, with lowerjaw extending slightly beyond upperjaw. Weakly developed central rib of noseleaf extends to tip. Ears and wing membranes are blackish brown, and wings attach to ankles. Forearms are naked. Tail membraneis only sparsely haired. Similar to other nectarfeeding bats, rostrum is narrow but only slightly shorter than braincase. In dorsal view of skull, rostrum is short, moderately broad, and inflated above M'; lateral margins are convex; postorbital region is strongly inflated, typically with small lateral projections; and posterior margin of infraorbital foramen typically projects beyond lateral outline of rostrum. I' is more than double size of I’. In palatal view, gaps are obvious between I' and I?, and P* has conspicuous rooted lingual cusp. Dental formula for all species of Hsunycterisis12/2,C1/1,P2/3,M 3/3 ( x2 ) = 34. Chromosomal complement has 2n = 36 and FN = 50, with multiple acrocentric autosomes. Habitat. Pre-montane rainforests of Pacific coastal plain at elevations below 200 m (most records) but also up to ¢. 2400 m . Cadena’s Nectar Bat seems to prefer primary and secondary rainforests but is also found in clearings, gardens, and banana plantations. A single record comes from Andean montane forests on eastern slope of Western Andes Range. Many individuals have been caught in ground-level (0-3 m) mist nets, indicating preference to fly in low forest strata. Food and Feeding. Cadena’s Nectar Batis primarily nectarivorous but also eats pollen, insects, and banana flowers. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List (as Lonchophylla cadenai ). Available information is inadequate to make reliable assessments of conservation risks to Cadena’s Nectar Bat. Bibliography. Baker et al. (1982), Carrera et al. (2010), Davalos & Jansa (2004), Griffiths & Gardner (2008b), Parlos et al. (2014), Ribeiro et al. (2003), Velazco et al. (2017), Woodman & Timm (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Hsunycteris cadenai	Hsunycteris		cadenai	Woodman and Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 462	Cadena&apos;s Nectar Bat	None.	across from the village of Zabaletas, Valle del Cauca Department; Colombia	SW Colombia, perhaps NW Ecuador	Not listed.	Data Deficient as Lonchophylla cadenai 	For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hsunycteris cadenai	23	Cadena's Nectar Bat	Cadena's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	cadenai	Woodman & Timm	2006	1	Lonchophylla_cadenai	Woodman, N. & Timm, R. M. (2006). Characters and Phylogenetic Relationships of Nectar-feeding Bats, with Descriptions of New Lonchophylla from Western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119, 462.	https://repository.si.edu/bitstream/handle/10088/2642/2006_Lonchophylla2.pdf?sequence=1&isAllowed=y	USNM 483359		"Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Río Zabaletas, across from the village of Zabaletas (3°44' N, 76°57' W); Valle del Cauca Department; Colombia."	3.7333	-76.95	cadenai (Woodman & Timm, 2006)	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.	Colombia|Ecuador	South America	Neotropic	DD	0	0	0	Hsunycteris_cadenai	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Lonchophylla cadenai	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Lonchophylla	cadenai	Woodman &; Timm, 2006		90000000	Lonchophylla cadenai	Data Deficient		2018	2016-11-01 00:00:00 UTC	3.1	English	The species is listed as Data Deficient because the available information is inadequate to make a reliable assessment of its risk of extinction based on its distribution and/or population status. Although the Andean and lowland forests on the Pacific region are subject to various threats, the distribution of this species ;also includes montane forests and nearby protected areas. In addition, a further study of all records from that region could reveal a larger distribution.	The behaviour and ecology of this species ;are poorly known. It is known from an elevational zone between 75 and 2,400 m in the western slope of the Colombian Andes, in areas dominated by sub-Andean and Andean mountainous forests, as well as piedmont forests (Mantilla-Meluk et al.  2010) as well as in the lowlands forests surrounded by a mosaic of primary and secondary forests and agricultural lands, in northwestern Ecuador (Carrera et al.  2010, as L. thomasi ). These sites are on the Pacific coastal plain in Premontane Rain Forest (Woodman and Timm 2006).	The Pacific slope of the western Colombian Andes contains a mosaic of primary and secondary forests and agricultural lands. The Ecuadorian Pacific lowlands have been severely transformed into agricultural lands. In general, this is a region subject to changes in land use (agriculture), although this is not uniform and some areas still include well-preserved forests.	The original description (Woodman and Timm 2006) included a few specimens and localities from western Colombia, an analysis of abundance was not performed. The study by Mantilla-Meluk et al.  (2010) listed two more individuals but could not identify additional representatives of the species. However, at northwestern Ecuador it was a relatively common species (Carrera et al.  2010, Parlos et al.  2014). As other species in the genus, this one seems to be generally uncommonly netted (Woodman and Timm 2006).	Unknown	Lonchophylla cadenai ;is only known from Valle del Cauca and Risaralda departments, western Colombia (Woodman and Timm 2006, Mantilla-Meluk et al.  2010). Specimens from Esmeraldas Province, northwestern corner of Ecuador, also represent this species (Parlos et al.  2014). Recent phylogenetic analyses indicate that this and other species in the genus actually belong in a new genus, ;Hsunycteris (Parlos et al.  2014).		Terrestrial	The species is distributed nearby some protected areas. ;However, further studies are needed into its distribution, habitat, ecology, and threats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Hsunycteris		cadenai	Woodman and Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 462	Cadena&apos;s Nectar Bat	None.	across from the village of Zabaletas, Valle del Cauca Department; Colombia	SW Colombia, perhaps NW Ecuador	Not listed.	Data Deficient as Lonchophylla cadenai 	For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).	Hsunycteris cadenai	1004924	23	Cadena's Nectar Bat	Cadena's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	LONCHOPHYLLINAE	HSUNYCTERINI	Hsunycteris	NA	cadenai	Woodman & Timm	2006	1	Lonchophylla_cadenai	Woodman, N. & Timm, R. M. (2006). Characters and Phylogenetic Relationships of Nectar-feeding Bats, with Descriptions of New Lonchophylla from Western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119, 462.	https://repository.si.edu/bitstream/handle/10088/2642/2006_Lonchophylla2.pdf?sequence=1&isAllowed=y	USNM 483359		"Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Río Zabaletas, across from the village of Zabaletas (3°44' N, 76°57' W); Valle del Cauca Department; Colombia."	3.733333	-76.95	cadenai (Woodman & Timm, 2006)	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Colombia|Ecuador	South America	Neotropic	DD	0	0	0	Hsunycteris_cadenai	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hsunycteris_cadenai	1004924	23	Cadena's Nectar Bat	Cadena's Small Nectar Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Lonchophyllinae	Hsunycterini	Hsunycteris	NA	cadenai	Woodman & Timm	1	Lonchophylla cadenai	Woodman, N. and Timm, R.M. 2006-12-22. Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new _Lonchophylla_ from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington 119(4):437-476.	https://doi.org/10.2988/0006-324X(2006)119[437:CAPRON]2.0.CO;2	USNM:MAMM:483359	holotype	http://n2t.net/ark:/65665/384d53a4b-5cd8-47ad-8f47-ae2e0b7ad8a2	"Twenty-nine km SE of Buenaventura, 75 m elevation; east bank of Río Zabaletas, across from the village of Zabaletas (3°44' N, 76°57' W); Valle del Cauca Department; Colombia."	3.733333	-76.95	recently described; moved from Lonchophylla to the recently described Hsunycteris	Woodman, N., & Timm, R. M. (2006). Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini). Proceedings of the Biological Society of Washington, 119(4), 437-476.|Parlos, J. A., Timm, R. M., Swier, V. J., Zeballos, H., & Baker, R. J. (2014). Evaluation of the paraphyletic assemblages within Lonchophyllinae, with description of a new tribe and genus. Occasional Papers of the Museum of Texas Tech University, 320, 1-23.				Colombia|Ecuador	South America	Neotropic	DD (as Lonchophylla cadenai)	0	0	0	Hsunycteris_cadenai	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Hsunycteris		cadenai	Woodman & Timm	2006	1	Proc. Biol. Soc. Wash.	119(4): 462	Cadena&apos;s Nectar Bat	None.	across from the village of Zabaletas, Valle del Cauca Department; Colombia	SW Colombia, perhaps NW Ecuador	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88149262/88149265/' target='_blank'>Data Deficient as Lonchophylla cadenai</a>	For discussion of possible Ecuadorian specimens see Woodman and Timm (2006).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hsunycteris cadenai; Hsunycteris cadenai; Hsunycteris cadenai; Hsunycteris cadenai; cadenai; Lonchophylle de Cadena; Cadena-Nektarfledermaus; Sunicterio de Cadena; Cadena’s Small Nectar Bat; Cadena's Nectar Bat; Cadena's Small Nectar Bat; Cadena&apos;s Nectar Bat; H. cadenai