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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L458	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Eptesicus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus	Histiotus velatus		[HMW] Plecotus velatus 1. Geoffroy Saint-Hilaire, 1824 , Curitiba, Parana , Brazil . Taxonomic status of Histiotus velatus is somewhat uncertain; in the past, more than one form seemed to exist in its wide distribution. Taxonomic revision is needed. Monotypic.; [batnames2022] Subgenus Histiotus .; [MDD2022] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2023] None.; [MDD2023] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2025_1.7] None.						miotis.			miotis			velatus	velatus - miotis	velatus, euryotis, miotis		velatus	velatus - miotis	velatus, euryotis, miotis	velatus, euryotis, miotis	velatus	velatus - miotis	velatus (I. Geoffroy Saint-Hilaire, 1824)|euryotis (Temminck, 1840) [not used as valid]|miotis O. Thomas, 1916		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Brazil	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Histiotus velatus	Brazil, Parana, Curitiba.	I. Geoffroy	1824	Ann. Sei. Nat. Zool., 3:446.	Distribution: Confined to eastern Brazil and eastern Paraguay.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Brazil, Paraguay	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	I. Geoffroy	1824	Ann. Sci. Nat. Zool., 3:446.		E Brazil, Paraguay.	Brazil, Parana, Curitiba.		I. GEOFFROY	1824	Ears more or less triangular. Connecting band well developed. Skull relatively narrow. Forearm length, 45-49 mm.	Distribution: Confined to eastern Brazil and eastern Paraguay.	No subspecies currently recognized.		122	species	H. velatus	I. GEOFFROY	1824	Histiotus	genus	Histiotus velatus				Ears more or less triangular. Connecting band well developed. Skull relatively narrow. Forearm length, 45-49 mm.	No subspecies currently recognized.		3. H. velatus (I. GEOFFROY 1824).	3	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Histiotus velatus	Histiotus		velatus	I. Geoffroy	y	1824		Ann. Sci. Nat. Zool.	3		446		Tropical Big-eared Brown Bat	Brazil, Parana, Curitiba.	E Brazil, Bolivia, Paraguay, NW Argentina	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	miotis Thomas, 1916.		4C3D87E8FFA86A17FF829E7B16E6BE27	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	841	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFA86A17FF829E7B16E6BE27.xml	Histiotus velatus	Vespertilionidae	Histiotus	velatus		1824	Sérotine voilée @fr | Tropische Grof Rohrfledermaus @de | Histiotus tropical @es	Plecotus velatus 1. Geoffroy Saint-Hilaire, 1824 , Curitiba, Parana , Brazil . Taxonomic status of Histiotus velatus is somewhat uncertain; in the past, more than one form seemed to exist in its wide distribution. Taxonomic revision is needed. Monotypic.	Peru , Bolivia , C & S Brazil , S Paraguay , and NE Argentina .	Head-body 58-2-70-8 mm, tail 46-60 mm, ear 24-3 33-5 mm, hindfoot 6-11 mm, forearm 42-50 mm; weight 9-17 g. Tibia lengths are 18-8-20-2 mm. Dorsal hairs of the Tropical Big-eared Brown Bat are slightly bicolored, with blackish brown bases and lighter cinnamon-brown tips; ventral hairs have dark brown bases and grayish to dark chestnut tips. Ears are very large, triangular, brown to dark brown, and connected by well-developed membrane (c. 3 mm high) across forehead; tragus is long, broad, and rounded at tip. Membranes are naked and brown to dark brown. Calcar is well developed. Skull is robust, narrower than observed in congeners; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally and narrower than in congeners; caudal spine is well developed; and tympanic bullae are large and rounded.	Wide variety of habitats including moist forests, savannas in grasslands, evergreen forests, semideciduous forests, open and shrub savannas, plantations, and urban areas from sea level up to elevations of ¢. 2400 m .	Tropical Big-eared Brown Bats are insectivorous. In south-eastern Brazil , a small group preyed on flying winged termites (Isoptera). Feces contained fragments of Lepidoptera , Hymenoptera , Coleoptera , and Araneae ( Pholcidae and Araneoidea).	In southern Brazil , pregnant Tropical Big-eared Brown Bats were captured in October-December, one female captured in October had two embryos, and young were observed in late October-November and early December. Available data suggest that parturition is synchronized and takes place once a year in spring.	Tropical Big-eared Brown Bats are nocturnal. Individuals were observed leaving roosts at sunset, and captures occurred during the first two hours after sunset. Roosts have been found in buildings, such as inside of churches and roofs. Study of a colony in southern Brazil showed that individuals were “asleep” ¢.75% of the time and “alert” ¢.17% of the time during the day; sometimes all individuals in the colony were sleeping. Number of individuals in the colony changed throughout the year: up to 70 individuals in spring and summer and down to 10 individuals in autumn and winter. Movements are closely related to oscillations in temperature and photoperiod. Echolocation calls sweep down from 25 kHz to 15 kHz, with durations of 5-8 milliseconds.	Colonies of Tropical Big-eared Brown Bats can be 6-12 individuals up to 70 individuals. They share roosts with Pallas’s Mastiff Bats (Molossus molossus) and Silver-tipped Myotis ( Myotis albescens ).	Classified as Data Deficient on The IUCN Red List. The Tropical Big-eared Brown Bat occurs in protected areas. Deforestation might be a threat, but additional research is needed to clarify it conservation status.	Barquez et al. (1999) | Bianconi & Pedro (2017) | Feijé, Rocha & Althoff (2015) | Fenton, Whitaker et al. (1999) | Gonzalez & Barquez (2016¢) | Handley & Gardner (2008) | Lopez-Gonzélez (2004) | Nora & Chaves (2006) | Simmons (2005) | Thomas (1916b) | Vizotto & Taddei (1973) | Welter (2009)	https://zenodo.org/record/6398170/files/figure.png	171. Tropical Big-eared Brown Bat Histiotus velatus French: Sérotine voilée / German: Tropische GrofRohrfledermaus / Spanish: Histiotus tropical Taxonomy. Plecotus velatus 1. Geoffroy Saint-Hilaire, 1824 , Curitiba, Parana , Brazil . Taxonomic status of Histiotus velatus is somewhat uncertain; in the past, more than one form seemed to exist in its wide distribution. Taxonomic revision is needed. Monotypic. Distribution. Peru , Bolivia , C & S Brazil , S Paraguay , and NE Argentina . Descriptive notes. Head-body 58-2-70-8 mm, tail 46-60 mm, ear 24-3 33-5 mm, hindfoot 6-11 mm, forearm 42-50 mm; weight 9-17 g. Tibia lengths are 18-8-20-2 mm. Dorsal hairs of the Tropical Big-eared Brown Bat are slightly bicolored, with blackish brown bases and lighter cinnamon-brown tips; ventral hairs have dark brown bases and grayish to dark chestnut tips. Ears are very large, triangular, brown to dark brown, and connected by well-developed membrane (c. 3 mm high) across forehead; tragus is long, broad, and rounded at tip. Membranes are naked and brown to dark brown. Calcar is well developed. Skull is robust, narrower than observed in congeners; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally and narrower than in congeners; caudal spine is well developed; and tympanic bullae are large and rounded. Habitat. Wide variety of habitats including moist forests, savannas in grasslands, evergreen forests, semideciduous forests, open and shrub savannas, plantations, and urban areas from sea level up to elevations of ¢. 2400 m . Food and Feeding. Tropical Big-eared Brown Bats are insectivorous. In south-eastern Brazil , a small group preyed on flying winged termites (Isoptera). Feces contained fragments of Lepidoptera , Hymenoptera , Coleoptera , and Araneae ( Pholcidae and Araneoidea). Breeding. In southern Brazil , pregnant Tropical Big-eared Brown Bats were captured in October-December, one female captured in October had two embryos, and young were observed in late October-November and early December. Available data suggest that parturition is synchronized and takes place once a year in spring. Activity patterns. Tropical Big-eared Brown Bats are nocturnal. Individuals were observed leaving roosts at sunset, and captures occurred during the first two hours after sunset. Roosts have been found in buildings, such as inside of churches and roofs. Study of a colony in southern Brazil showed that individuals were “asleep” ¢.75% of the time and “alert” ¢.17% of the time during the day; sometimes all individuals in the colony were sleeping. Number of individuals in the colony changed throughout the year: up to 70 individuals in spring and summer and down to 10 individuals in autumn and winter. Movements are closely related to oscillations in temperature and photoperiod. Echolocation calls sweep down from 25 kHz to 15 kHz, with durations of 5-8 milliseconds. Movements, Home range and Social organization. Colonies of Tropical Big-eared Brown Bats can be 6-12 individuals up to 70 individuals. They share roosts with Pallas’s Mastiff Bats (Molossus molossus) and Silver-tipped Myotis ( Myotis albescens ). Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Tropical Big-eared Brown Bat occurs in protected areas. Deforestation might be a threat, but additional research is needed to clarify it conservation status. Bibliography. Barquez et al. (1999), Bianconi & Pedro (2017), Feijé, Rocha & Althoff (2015), Fenton, Whitaker et al. (1999), Gonzalez & Barquez (2016¢), Handley & Gardner (2008), Lopez-Gonzélez (2004), Nora & Chaves (2006), Simmons (2005), Thomas (1916b), Vizotto & Taddei (1973), Welter (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus velatus	Eptesicus	Histiotus	velatus	I. Geoffroy	1824	1	Ann. Sci. Nat. Zool.	0.4347	Tropical Big-eared Brown Bat	 miotis Thomas, 1916.	Brazil, Parana, Curitiba.	E Brazil, Bolivia, Paraguay, NW Argentina	Not listed.	Data Deficient as Histiotus velatus 	Subgenus Histiotus .	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Histiotus velatus	23	Tropical Big-eared Brown Bat	Tropical Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	velatus	I. Geoffroy Saint-Hilaire	1824	1	Plecotus_velatus	Geoffroy Saint-Hilaire, I. (1824). Sur les Vespertilions du BrÃ©sil. Annales des sciences naturelles, 3, 446.	https://www.biodiversitylibrary.org/item/50402#page/452/mode/1up	MNHN 1997-1828		Curitiba, ParanÃ¡, Brazil.			velatus (I. Geoffroy Saint-Hilaire, 1824)|euryotis (Temminck, 1840) [nomen nudum]|miotis O. Thomas, 1916	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA	Peru|Bolivia|Brazil|Paraguay|Argentina	South America	Neotropic	DD	0	0	0	Histiotus_velatus	0	sciname match	Histiotus_velatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10203	Histiotus velatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Histiotus	velatus	(I. Geoffroy, 1824)		20000000	Histiotus velatus	Data Deficient		2016	2016-06-08 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of continuing problems with its taxonomy as well as absence of recent information on its extent of occurrence, status and ecological requirements. It needs taxonomic revision because it may be two species.	This species is insectivorous, but there is very little information about this bat. It is found in forests and buildings (Gonzalez pers. comm.).	The species could be threatened by deforestation.	There is no information about population trend, only that in Argentina it is scarce and in Brazil it is abundant.	Unknown	The species is found in east Brazil, Bolivia, and northwest Argentina (Simmons 2005), also in Paraguay (Lopez-Gonzalez 2004) and Peru (Handley and Gardner 2008). Presence in Uruguay is unconfirmed (Handley and Gardner 2008).		Terrestrial	Conservation actions for this species are unknown.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Histiotus		velatus	I. Geoffroy Saint-Hilaire	1824	0	Ann. Sci. Nat. Zool.	0.434722	Tropical Big-eared Brown Bat	 miotis Thomas, 1916.	Brazil, Parana, Curitiba.	E Brazil, Bolivia, Paraguay, NW Argentina	Not listed.	Data Deficient	None.	Histiotus velatus	1005560	23	Tropical Big-eared Brown Bat	Tropical Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	velatus	I. Geoffroy Saint-Hilaire	1824	1	Plecotus_velatus	Geoffroy Saint-Hilaire, I. (1824). Sur les Vespertilions du BrÃ©sil. Annales des sciences naturelles, 3, 446.	https://www.biodiversitylibrary.org/item/50402#page/452/mode/1up	MNHN 1997-1828		Curitiba, ParanÃ¡, Brazil.			velatus (I. Geoffroy Saint-Hilaire, 1824)|euryotis (Temminck, 1840) [nomen nudum]|miotis O. Thomas, 1916	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA				Peru|Bolivia|Brazil|Paraguay|Argentina	South America	Neotropic	DD	0	0	0	Histiotus_velatus	0	sciname match	Histiotus_velatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Histiotus_velatus	1005560	23	Tropical Big-eared Brown Bat	Tropical Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Histiotus	NA	velatus	I. Geoffroy Saint-Hilaire	1	Plecotus velatus	Geoffroy Saint-Hilaire, I. 1824. Sur les Vespertilions du BrÃ©sil. Annales des sciences naturelles 3:440-447.	https://www.biodiversitylibrary.org/page/5813767	MNHN-ZM-MO-1997-1828	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1997-1828	Curitiba, ParanÃ¡, Brazil.			NA	NA				Peru|Bolivia|Brazil|Paraguay|Argentina	South America	Neotropic	DD	0	0	0	Histiotus_velatus	0	sciname match	Histiotus_velatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Histiotus		velatus	I. Geoffroy Saint-Hilaire	1824	1	Ann. Sci. Nat. Zool.	0.434722	Tropical Big-eared Brown Bat	miotis Thomas, 1916.	Brazil, Parana, Curitiba	E Brazil, Bolivia, Paraguay, NW Argentina	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10203/22098684/' target='_blank'>Data Deficient</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Histiotus velatus; Histiotus velatus; Eptesicus velatus; Histiotus velatus; Histiotus velatus; Histiotus velatus; miotis; miotis; velatus; euryotis; miotis; Sérotine voilée; Tropische Grof Rohrfledermaus; Histiotus tropical; Tropical Big-eared Brown Bat; Tropical Leaf-eared Bat; Tropical Big-eared Brown Bat; Tropical Big-eared Brown Bat; H. velatus
