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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L457	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Eptesicus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus	Histiotus montanus		[MSW3] Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus; see Barquez et al. (1993) and Mares et al. (1995).; [HMW] Vespertilio montanus R. A. Philippi & Landbeck, 1861 , Cordillera, Santiago, Chile . Congeners H. alienus , H. laephotis , and H. magellanicus were previously considered subspecies of H. montanus , and its taxonomic status is confusing. It is frequently mistaken with H. laephotis and H. macrotus ; therefore, accurate data from literature are scarce. Three subspecies recognized.; [batnames2022] Subgenus Histiotus . Does not include colombiae ; see Rodríguez-Posada aet al. (2021). Does not include  laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus ; see Barquez et al. (1993).; [MDD2022] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [IUCN] Has taxonomic problems.; [batnames2023] Does not include colombiae ; see Rodríguez-Posada aet al. (2021). Does not include  laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus ; see Barquez et al. (1993).; [MDD2023] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2025_1.7] Does not include colombiae; see Rodríguez-Posada aet al. (2021). Does not include  laephotis; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus; see Barquez et al. (1993).						capucinus, colombiae, inambarus, magellanicus, segethii.	colombiae, inambarus, montanus, magellanicus, alienus	montanus, colombiae, inambarus	segethii	montanus, colombiae, inambarus		montanus, colombiae, inambarus	montanus - segethii	montanus, segethii, peruvianus, peruvianus, inambarus	Has taxonomic problems.	montanus, inambarus	montanus - segethii	montanus, segethii, peruvianus, peruvianus, inambarus	peruvianus, montanus, segethii, peruvianus, inambarus	inambarus, montanus	montanus - segethii	peruvianus (Le Conte, 1858)|montanus (R. A. Philippi & Landbeck, 1861)|segethii (W. C. H. Peters, 1865)|peruvianus (Fitzinger, 1872) [preoccupied]|inambarus H. E. Anthony, 1920		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Colombia – Chile, Argentina	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Histiotus montanus	Chile, Santiago Cordillera.	Philippi and Landbeck	1861	Arch. Naturgesch., p. 289.	Distribution: Ranging in the Andes from Venezuela to northwestern Argentina but south of the tropics occurring also in the lowlands south to Tierra del Fuego.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Venezuela, Colombia – Chile, Argentina, Uruguay	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Philippi and Landbeck	1861	Arch. Naturgesch., p. 289.		Chile, Argentina, Uruguay, W Bolivia, S Peru, Ecuador, Colombia, Venezuela, perhaps N Peru and S Brazil.	Chile, Santiago Cordillera.		PHILIPPI & LANDBECK	1861	Ears relatively short and rounded. Connecting band poorly developed. Skull relatively broad. Forearm length, 44 51 mm.	Distribution: Ranging in the Andes from Venezuela to northwestern Argentina but south of the tropics occurring also in the lowlands south to Tierra del Fuego.	Five subspecies are here recognized:	H. m. colombiae (western Venezuela to Ecuador), H. m. inambarus (southern Peru to Paraguay), H. m. montanus (Uruguay, northern and central Argentina, central Chile), H. m. magellanicus (southern Chile and southern Argentina), H. m. alienus (definitely known only from southeastern Brazil and often separated specifically).	122	species	H. montanus	PHILIPPI & LANDBECK	1861	Histiotus	genus	Histiotus montanus				Ears relatively short and rounded. Connecting band poorly developed. Skull relatively broad. Forearm length, 44 51 mm.	Five subspecies are here recognized:		1. H. montanus (PHILIPPI & LANDBECK 1861).	1	_H. m. inambarus_ Anthony, 1920 (synonyms: _peruvianus_ (Fitzinger, 1872), _peruvianus_ (Le Conte, 1858)); _H. m. montanus_ (Philippi & Landbeck, 1861) (synonyms: _segethii_ (Peters, 1865))			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Histiotus montanus	Histiotus		montanus	Philippi and Landbeck	y	1861		Arch. Naturgesch.			289		Small Big-eared Brown Bat	Chile, Santiago Cordillera.	N Chile, Argentina, Uruguay, W Bolivia, S Peru, Ecuador, Colombia, Venezuela, perhaps N Peru and S Brazil.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	segethii Peters, 1864; colombiae Thomas, 1916; inambarus Anthony, 1920.	Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus; see Barquez et al. (1993) and Mares et al. (1995).	4C3D87E8FFAF6A10FF4D93951662B97B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	842	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFAF6A10FF4D93951662B97B.xml	Histiotus montanus	Vespertilionidae	Histiotus	montanus		1861	Sérotine des montagnes @fr | Kleine Gro Rohrfledermaus @de | Histiotus montano @es	Vespertilio montanus R. A. Philippi & Landbeck, 1861 , Cordillera, Santiago, Chile . Congeners H. alienus , H. laephotis , and H. magellanicus were previously considered subspecies of H. montanus , and its taxonomic status is confusing. It is frequently mistaken with H. laephotis and H. macrotus ; therefore, accurate data from literature are scarce. Three subspecies recognized.	H.m.montanusR.A.Philippi&Landbeck,1861—S&SEBrazil,Uruguay,Argentina,andC&SChile. H.m.colombiaeThomas,1916—AndesofVenezuela,Colombia,andEcuador. H. m. inambarus Anthony, 1920 — Peru and N Chile ; probably in W Bolivia .	Head—body c.51-67 mm,tail 40-55 mm, ear 20-29-5 mm, hindfoot 8-12 mm, forearm 42-5-51-1 mm; weight 9-13 g. Dorsal hairs of the Small Big-eared Brown Bat are bicolored, with black or dark brown bases and whitish tips; ventral hairs are strongly bicolored, with dark brown bases and white tips; some specimens are paler, with generally grayish venters. Ears are large, rounded, but relatively small compared with congeners; tragus is well developed, reaching one-half the ear length. Membrane connecting ears is usually absent but sometimes poorly developed. Membranes, face, and ears are pale light brown. Skull is robust; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process of jugal; palateis slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded. I* is bicuspid; I’ is rudimentary and separated from C' by small gap; P* is robust and in contact with C' and M'; lowerincisors are tricuspid; and P, is one-half the height of P,. Chromosomal complement has 2n = 50 and FN = 48.	Dry environments to humid forests in steppes, shrub grasslands, savannas, ombrophilous forests, semideciduous forests, dense rainforests, and araucaria forests from sea level up to an elevation of 4117 m .	Small Big-eared Brown Bats are aerial insectivores. Fecal and stomach samples contained fragments of Lepidoptera and Coleoptera , including Curculionidae (genus Pantomorus ) and Tenebrionidae (genus Parepitragus).	Pregnant Small Big-eared Brown Bats were captured in July-August and November-December in Ecuador and Argentina . Lactating females were captured in July in Ecuador . It is speculated that a female’s reproductive cycle starts at one year of age, and one young is born to each female per pregnancy.	Small Big-eared Brown Bats commonly roost in caves but also loose rocks, holes and crevices, buildings, and hollow trees. Individuals seem to hibernate in the southern part of its distribution. Echolocation calls have downward FM component, followed by QCF component. Calls sweep down from ¢.46-53 kHz to ¢.25-29 kHz, with duration of c.3-5 milliseconds. Interpulse interval averages 144 milliseconds, intensity is ¢.38 dB, and peak frequencies are ¢.32-35 kHz.	Small Big-eared Brown Bats usually form small colonies of5-20 individuals. They will share roosts with Brazilian Free-tailed Bats ( Tadanrida brasiliensis ), Yellowish Myotis ( Myotis levis ), and Red Myotis ( M. ruber ).	Classified as Least Concern on The [UCN Red List. The Small Big-eared Brown Bat is widely distributed, presumably with large populations, and occurs in protected areas. In the southern part of its distribution,it seems to be particularly abundant.	Acosta (1950) | Barquez & Diaz (2016b) | Barquez, Carbajal et al. (2013) | Barquez, Mares & Braun (1999) | Berrocal (2016) | Bianconi & Pedro (2017) | Carvalho et al. (2013) | Garbino (2016) | Handley (1996) | Handley & Gardner (2008) | Miranda , Pulchério-Leite et al. (2006) | Ossa et al. (2015) | Rodriguez-San Pedro & Simonetti (2013a) | Ruiz (2015) | Simmons (2005) | Williams & Mares (1978)	https://zenodo.org/record/6398176/files/figure.png	174. Small Big-eared Brown Bat Histiotus montanus French: Sérotine des montagnes / German: Kleine GroRohrfledermaus / Spanish: Histiotus montano Taxonomy. Vespertilio montanus R. A. Philippi & Landbeck, 1861 , Cordillera, Santiago, Chile . Congeners H. alienus , H. laephotis , and H. magellanicus were previously considered subspecies of H. montanus , and its taxonomic status is confusing. It is frequently mistaken with H. laephotis and H. macrotus ; therefore, accurate data from literature are scarce. Three subspecies recognized. Subspecies and Distribution. H.m.montanusR.A.Philippi&Landbeck,1861—S&SEBrazil,Uruguay,Argentina,andC&SChile. H.m.colombiaeThomas,1916—AndesofVenezuela,Colombia,andEcuador. H. m. inambarus Anthony, 1920 — Peru and N Chile ; probably in W Bolivia . Descriptive notes. Head—body c.51-67 mm,tail 40-55 mm, ear 20-29-5 mm, hindfoot 8-12 mm, forearm 42-5-51-1 mm; weight 9-13 g. Dorsal hairs of the Small Big-eared Brown Bat are bicolored, with black or dark brown bases and whitish tips; ventral hairs are strongly bicolored, with dark brown bases and white tips; some specimens are paler, with generally grayish venters. Ears are large, rounded, but relatively small compared with congeners; tragus is well developed, reaching one-half the ear length. Membrane connecting ears is usually absent but sometimes poorly developed. Membranes, face, and ears are pale light brown. Skull is robust; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process of jugal; palateis slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded. I* is bicuspid; I’ is rudimentary and separated from C' by small gap; P* is robust and in contact with C' and M'; lowerincisors are tricuspid; and P, is one-half the height of P,. Chromosomal complement has 2n = 50 and FN = 48. Habitat. Dry environments to humid forests in steppes, shrub grasslands, savannas, ombrophilous forests, semideciduous forests, dense rainforests, and araucaria forests from sea level up to an elevation of 4117 m . Food and Feeding. Small Big-eared Brown Bats are aerial insectivores. Fecal and stomach samples contained fragments of Lepidoptera and Coleoptera , including Curculionidae (genus Pantomorus ) and Tenebrionidae (genus Parepitragus). Breeding. Pregnant Small Big-eared Brown Bats were captured in July-August and November-December in Ecuador and Argentina . Lactating females were captured in July in Ecuador . It is speculated that a female’s reproductive cycle starts at one year of age, and one young is born to each female per pregnancy. Activity patterns. Small Big-eared Brown Bats commonly roost in caves but also loose rocks, holes and crevices, buildings, and hollow trees. Individuals seem to hibernate in the southern part of its distribution. Echolocation calls have downward FM component, followed by QCF component. Calls sweep down from ¢.46-53 kHz to ¢.25-29 kHz, with duration of c.3-5 milliseconds. Interpulse interval averages 144 milliseconds, intensity is ¢.38 dB, and peak frequencies are ¢.32-35 kHz. Movements, Home range and Social organization. Small Big-eared Brown Bats usually form small colonies of5-20 individuals. They will share roosts with Brazilian Free-tailed Bats ( Tadanrida brasiliensis ), Yellowish Myotis ( Myotis levis ), and Red Myotis ( M. ruber ). Status and Conservation. Classified as Least Concern on The [UCN Red List. The Small Big-eared Brown Bat is widely distributed, presumably with large populations, and occurs in protected areas. In the southern part of its distribution,it seems to be particularly abundant. Bibliography. Acosta (1950), Barquez & Diaz (2016b), Barquez, Carbajal et al. (2013), Barquez, Mares & Braun (1999), Berrocal (2016), Bianconi & Pedro (2017), Carvalho et al. (2013), Garbino (2016), Handley (1996), Handley & Gardner (2008), Miranda , Pulchério-Leite et al. (2006), Ossa et al. (2015), Rodriguez-San Pedro & Simonetti (2013a), Ruiz (2015), Simmons (2005), Williams & Mares (1978).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus montanus	Eptesicus	Histiotus	montanus	Philippi and Landbeck	1861	1	Arch. Naturgesch.	p. 289	Small Big-eared Brown Bat	 segethii Peters, 1864; <b> colombiae </b> Thomas, 1916; <b> inambarus </b> Anthony, 1920.	Chile, Santiago Cordillera	N Chile, Argentina, Uruguay, W Bolivia, S Peru, Brazil perhaps N Peru	Not listed.	Least Concern as Histiotus montanus 	Subgenus Histiotus . Does not include colombiae ; see Rodríguez-Posada aet al. (2021). Does not include  laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus ; see Barquez et al. (1993).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Histiotus montanus	23	Small Big-eared Brown Bat	Common Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	montanus	R. A. Philippi & Landbeck	1861	1						Cordillera, Santiago, Chile.			montanus (R. A. Philippi & Landbeck, 1861)|segethii (W. Peters, 1864)|peruvianus (Le Conte, 1858) [nomen oblitum]|peruvianus (Fitzinger, 1872) [unavailable name]|inambarus Anthony, 1920	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA	Peru|Chile|Bolivia|Brazil|Uruguay|Argentina	South America	Neotropic	LC	0	0	0	Histiotus_montanus	0	sciname match	Histiotus_montanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10202	Histiotus montanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Histiotus	montanus	(Philippi &; Landbeck, 1861)	Has taxonomic problems.	20000000	Histiotus montanus	Least Concern		2016	2016-07-27 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Like other species in the genus, H. montanus is an aerial insectivorous bat (Handley and Gardner 2008). It is found roosting in houses, caves, and hollow trees, forming colonies with until 20 individuals. In the southern part of their range (Chile and Argentina), they may hibernate. Because of the many changes in the taxonomy of this species (some subspecies now treated as valid species: alienus , laephotis , magellanicus ), it is difficult to obtain a clear ecological characterization of the species.	There are no major threats to this species.	It seems abundant on the southern part of its distribution (Barquez, pers. comm.), but is rather uncommon at the northern part of the range.	Unknown	This species is found in central and southern Chile, Argentina, Bolivia, Peru, Ecuador, Colombia, and Venezuela (Simmons 2005, Handley and Gardner 2008).		Terrestrial	Because of the recent changes in its taxonomy, it is difficult to provide a definitive characterization of the species and to assess effective conservation actions. Because of its still wide distribution, their populations are protected over a number of natural areas.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Histiotus		montanus	Philippi and Landbeck	1861	0	Arch. Naturgesch.	p. 289	Small Big-eared Brown Bat	 segethii Peters, 1864; <b> colombiae </b> Thomas, 1916; <b> inambarus </b> Anthony, 1920.	Chile, Santiago Cordillera	N Chile, Argentina, Uruguay, W Bolivia, S Peru, Brazil perhaps N Peru	Not listed.	Least Concern	Does not include colombiae ; see Rodríguez-Posada aet al. (2021). Does not include  laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus ; see Barquez et al. (1993).	Histiotus montanus	1005559	23	Small Big-eared Brown Bat	Common Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	montanus	R. A. Philippi & Landbeck	1861	1						Cordillera, Santiago, Chile.			montanus (R. A. Philippi & Landbeck, 1861)|segethii (W. Peters, 1864)|peruvianus (Le Conte, 1858) [nomen oblitum]|peruvianus (Fitzinger, 1872) [unavailable name]|inambarus Anthony, 1920	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA				Peru|Chile|Bolivia|Brazil|Uruguay|Argentina	South America	Neotropic	LC	0	0	0	Histiotus_montanus	0	sciname match	Histiotus_montanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Histiotus_montanus	1005559	23	Small Big-eared Brown Bat	Common Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Histiotus	NA	montanus	R. A. Philippi & Landbeck	1	Vespertilio montanus	Philippi, R.A. and Landbeck, C.L. 1861. Neue Wirbelthiere von Chile. Archiv für Naturgeschichte 27:289-301.	https://www.biodiversitylibrary.org/page/7055816		holotype		Cordillera, Santiago, Chile.			NA	NA				Peru|Chile|Bolivia|Brazil|Uruguay|Argentina	South America	Neotropic	LC	0	0	0	Histiotus_montanus	0	sciname match	Histiotus_montanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Histiotus		montanus	Philippi & Landbeck	1861	1	Arch. Naturgesch.	p. 289	Small Big-eared Brown Bat	segethii Peters, 1864; colombiae Thomas, 1916; inambarus Anthony, 1920.	Chile, Santiago Cordillera	N Chile, Argentina, Uruguay, W Bolivia, S Peru, Brazil perhaps N Peru	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10202/22098875/' target='_blank'>Least Concern</a>	Does not include colombiae; see Rodríguez-Posada aet al. (2021). Does not include  laephotis; see Autino et al. (1999) and Barquez and Diaz (2001). Does not include magellanicus; see Barquez et al. (1993).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Histiotus montanus; Histiotus montanus; Eptesicus montanus; Histiotus montanus; Histiotus montanus; Histiotus montanus; montanus; colombiae; inambarus; segethii; montanus; colombiae; inambarus; colombiae; inambarus; segethii; montanus; segethii; peruvianus; peruvianus; inambarus; Sérotine des montagnes; Kleine Gro Rohrfledermaus; Histiotus montano; Small Big-eared Brown Bat; Common Leaf-eared Bat; Small Big-eared Brown Bat; Small Big-eared Brown Bat; H. montanus