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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L454	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Eptesicus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus	Histiotus macrotus		[MSW2] Includes laephotis; see Cabrera (1958:108).; [MSW3] Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001).; [HMW] Nycticeius macrotus Poeppig, 1835 , Antuco, Bio Bio, Chile . Histiotus laephotis has been previously considered a subspecies of H. macrotus . It is frequently mistaken with H. laephotis and H. montanus , therefore, accurate data from literature are scarce. Monotypic.; [batnames2022] Subgenus Histiotus . Does not include laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001).; [MDD2022] previously included H. colombiae; has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2023] Subgenus Histiotus . Does not include laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001).; [MDD2023] previously included H. colombiae; has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [MDD2025_2.0] previously included H. colombiae; [batnames2025_1.7] Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001).; [MDD2025_2.2] previously included H. colombiae				laephotis		chilensis, laephotis, poeppigii.	laephotis, macrotus		chilensis, poeppigii			macrotus 	macrotus - chilensis, poeppigii	macrotus, chilensis, poeppigii		macrotus 	macrotus - chilensis, poeppigii	macrotus, chilensis, poeppigii	macrotus, chilensis, poeppigii	macrotus	macrotus - chilensis, poeppigii	macrotus (Poeppig, 1835)|chilensis (Lesson, 1836)|macrotis (Gay, 1847) [incorrect subsequent spelling]|poeppigii (Fitzinger, 1872)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Chile, Peru	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Histiotus macrotus	Chile, Bio-Bio, Antuco.	Poeppig	1835	Reise Chile Peru Amaz., 1:451.	Distribution: Ranging in the Andes and on the Pacific slope from southern Peru to southern Patagonia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Peru – NW Argentina, Chile	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Poeppig	1835	Reise Chile Peru Amaz., 1:451.	Includes laephotis; see Cabrera (1958:108).	Chile, NW Argentina, S Bolivia, S Peru.	Chile, Bio-Bio, Antuco.		POEPPIG	1835	Ears relatively long but rounded. Connecting band well developed. Skull relatively broad. Forearm length, 46 51 mm.	Distribution: Ranging in the Andes and on the Pacific slope from southern Peru to southern Patagonia.	Two subspecies are recognized:	H. m. laephotis (southern Peru to northwestern Argentina), H. m. macrotus (Chile).	122	species	H. macrotus	POEPPIG	1835	Histiotus	genus	Histiotus macrotus				Ears relatively long but rounded. Connecting band well developed. Skull relatively broad. Forearm length, 46 51 mm.	Two subspecies are recognized:		2. H. macrotus (POEPPIG 1835).	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Histiotus macrotus	Histiotus		macrotus	Poeppig	y	1835		Reise Chile Peru Amaz.	1		451		Big-eared Brown Bat	Chile, Bio-BIo, Antuco.	Chile, Argentina.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	chilensis Lesson, 1836; poeppigii Fitzinger, 1872.	Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001).	4C3D87E8FFAF6A11FA579F6B1F2FB899	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	842	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFAF6A11FA579F6B1F2FB899.xml	Histiotus macrotus	Vespertilionidae	Histiotus	macrotus		1835	Sérotine a grandes oreilles @fr | Gewohnliche Gro Rohrfledermaus @de | Histiotus orejudo @es	Nycticeius macrotus Poeppig, 1835 , Antuco, Bio Bio, Chile . Histiotus laephotis has been previously considered a subspecies of H. macrotus . It is frequently mistaken with H. laephotis and H. montanus , therefore, accurate data from literature are scarce. Monotypic.	C Chile and NW, C & W Argentina . Records from Peru should be reviewed.	Head-body ¢.59-65 mm, tail 48-62 mm, ear 28-34 mm, hindfoot 9-11 mm, forearm 45-9-50-9 mm; weight 10-3-20 g. Dorsal hairs of the Common Big-eared Brown Bat are strongly bicolored, with black or dark brown bases and yellowish tips; ventral hairs have dark brown bases and yellowish white tips. Ears are extremely large; tragusis well developed. Membrane connecting ears is poorly developed. Membranes, face, and ears are brown to dark brown, notably darker than body. Skull is robust; rostrum is wide; sagittal and lambdoidalcrests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded.	Great variety of habitats ranging from deserts to montane rainforests, including Dry Chaco , Espinal, Yungas, Patagonian forests and steppes, temperate forests, moist montane Chacoan forests, desert and rocky canyons, and mature forests at elevations of 240-3600 m. Common Big-eared Brown Bats have been captured over streams, ponds, and irrigation canals.	Common Big-eared Brown Bats are insectivorous. Feces contained fragments of Lepidoptera (36-6% by frequency), Coleoptera (30-6%), Trichoptera (13.6%), Diptera (9-8%), and Acari (1-7 %).	In Rio Negro, Argentina , pregnant and lactating Common Big-eared Brown Bats were captured in December; pregnant females carried only one embryo.	Common Big-eared Brown Bats usually roost in mines, natural caves, buildings, and rocky crevices.	The Common Big-eared Brown Bat is usually solitary in central Chile , but in other parts ofits distribution, colonies from 20 to hundreds of individuals were found. It will share roosts with Brazilian Free-tailed Bats ( Tadarida brasiliensis ) and Myotis spp.	Classified as Least Concern on The IUCN Red List. The Common Big-eared Brown Batis widely distributed, presumably with large populations. It is relatively common in northern Chile but seems uncommon in Argentina .	Aragon & Aguirre (2014) | Barquez & Diaz (2016a) | Barquez et al. (1999) | Giménez (2010) | Giménez et al. (2015) | Handley & Gardner (2008) | Lopez-Gonzélez et al. (1998) | Mann (1978) | Mares et al. (1995) | Simmons (2005)	https://zenodo.org/record/6398180/files/figure.png	176. Common Big-eared Brown Bat Histiotus macrotus French: Sérotine a grandes oreilles / German: Gewohnliche GroRohrfledermaus / Spanish: Histiotus orejudo Taxonomy. Nycticeius macrotus Poeppig, 1835 , Antuco, Bio Bio, Chile . Histiotus laephotis has been previously considered a subspecies of H. macrotus . It is frequently mistaken with H. laephotis and H. montanus , therefore, accurate data from literature are scarce. Monotypic. Distribution. C Chile and NW, C & W Argentina . Records from Peru should be reviewed. Descriptive notes. Head-body ¢.59-65 mm, tail 48-62 mm, ear 28-34 mm, hindfoot 9-11 mm, forearm 45-9-50-9 mm; weight 10-3-20 g. Dorsal hairs of the Common Big-eared Brown Bat are strongly bicolored, with black or dark brown bases and yellowish tips; ventral hairs have dark brown bases and yellowish white tips. Ears are extremely large; tragusis well developed. Membrane connecting ears is poorly developed. Membranes, face, and ears are brown to dark brown, notably darker than body. Skull is robust; rostrum is wide; sagittal and lambdoidalcrests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded. Habitat. Great variety of habitats ranging from deserts to montane rainforests, including Dry Chaco , Espinal, Yungas, Patagonian forests and steppes, temperate forests, moist montane Chacoan forests, desert and rocky canyons, and mature forests at elevations of 240-3600 m. Common Big-eared Brown Bats have been captured over streams, ponds, and irrigation canals. Food and Feeding. Common Big-eared Brown Bats are insectivorous. Feces contained fragments of Lepidoptera (36-6% by frequency), Coleoptera (30-6%), Trichoptera (13.6%), Diptera (9-8%), and Acari (1-7 %). Breeding. In Rio Negro, Argentina , pregnant and lactating Common Big-eared Brown Bats were captured in December; pregnant females carried only one embryo. Activity patterns. Common Big-eared Brown Bats usually roost in mines, natural caves, buildings, and rocky crevices. Movements, Home range and Social organization. The Common Big-eared Brown Bat is usually solitary in central Chile , but in other parts ofits distribution, colonies from 20 to hundreds of individuals were found. It will share roosts with Brazilian Free-tailed Bats ( Tadarida brasiliensis ) and Myotis spp. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Common Big-eared Brown Batis widely distributed, presumably with large populations. It is relatively common in northern Chile but seems uncommon in Argentina . Bibliography. Aragon & Aguirre (2014), Barquez & Diaz (2016a), Barquez et al. (1999), Giménez (2010), Giménez et al. (2015), Handley & Gardner (2008), Lopez-Gonzélez et al. (1998), Mann (1978), Mares et al. (1995), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus macrotus	Eptesicus	Histiotus	macrotus	Poeppig	1835	1	Reise Chile Peru Amaz.	0.3549	Big-eared Brown Bat	 chilensis Lesson, 1836; poeppigii Fitzinger, 1872.	Chile, Bio-BIo, Antuco.	Chile, Argentina.	Not listed.	Least Concern as Histiotus macrotus 	Subgenus Histiotus . Does not include laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Histiotus macrotus	23	Common Big-eared Brown Bat	Greater Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	macrotus	Poeppig	1835	1						Antuco, Bío Bío, Chile.			macrotus (Poeppig, 1835)|chilensis (Lesson, 1836)|poeppigii (Fitzinger, 1872)	previously included H. colombiae; has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	Rodríguez-Posada, M. E., Morales-Martínez, D. M., Ramírez-Chaves, H. E., Martínez-Medina, D., & Calderón-Acevedo, C. A. (2021). A new species of Long-eared Brown Bat of the genus Histiotus (Chiroptera) and the revalidation of Histiotus colombiae. Caldasia, 43(2).	Argentina|Chile	South America	Neotropic	LC	0	0	0	Histiotus_macrotus	0	sciname match	Histiotus_macrotus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10201	Histiotus macrotus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Histiotus	macrotus	(Poeppig, 1835)		20000000	Histiotus macrotus	Least Concern		2016	2016-07-25 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its presumed large population, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Normally inhabits mines, but can be found in houses. In the vicinity of Santiago (central Chile), the species is found in rock crevices and under roofs of houses (Mann 1978). This species usually occurs at medium elevations, as compared to other members of the genus.	No major threats throughout its range.	It is relatively common in Chile, but seems uncommon in Argentina. According to Mann (1978), H. macrotus is usually solitary and relatively uncommon in central Chile, but farther north, however, especially near agricultural fields, this same author reported colonies numbering in the hundreds, if not thousands, roosting in abandoned mines in the Province of Chañaral.	Unknown	The species is known from Chile, Argentina, and Paraguay (Simmons 2005, Lopez-Gonzalez et al . 1998). A record from Peru is published (Aragon and Aguirre 2014) but the photograph does not correspond to that species (Barquez, pers. comm.).		Terrestrial	There is no enough information about the biology and status of the species as to suggest specific actions.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Histiotus		macrotus	Poeppig	1835	1	Reise Chile Peru Amaz.	0.354861	Big-eared Brown Bat	 chilensis Lesson, 1836; poeppigii Fitzinger, 1872.	Chile, Bio-BIo, Antuco.	Chile, Argentina.	Not listed.	Least Concern as Histiotus macrotus 	Subgenus Histiotus . Does not include laephotis ; see Autino et al. (1999) and Barquez and Diaz (2001).	Histiotus macrotus	1005557	23	Common Big-eared Brown Bat	Greater Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	macrotus	Poeppig	1835	1						Antuco, Bío Bío, Chile.			macrotus (Poeppig, 1835)|chilensis (Lesson, 1836)|poeppigii (Fitzinger, 1872)	previously included H. colombiae; has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	Rodríguez-Posada, M. E., Morales-Martínez, D. M., Ramírez-Chaves, H. E., Martínez-Medina, D., & Calderón-Acevedo, C. A. (2021). A new species of Long-eared Brown Bat of the genus Histiotus (Chiroptera) and the revalidation of Histiotus colombiae. Caldasia, 43(2).				Argentina|Chile	South America	Neotropic	LC	0	0	0	Histiotus_macrotus	0	sciname match	Histiotus_macrotus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Histiotus_macrotus	1005557	23	Common Big-eared Brown Bat	Greater Leaf-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Histiotus	NA	macrotus	Poeppig	1	Nycticeius macrotus	Poeppig, E.F. 1835. Reise in Chile, Peru und auf dem Amazonenstrome, während der Jahre 1827–1832. Erster Band. Friedrich Fleischer, Leipzig, 466 pp.	https://www.digitale-sammlungen.de/de/view/bsb10366733?page=1				Antuco, Bío Bío, Chile.			previously included H. colombiae	Rodríguez-Posada, M. E., Morales-Martínez, D. M., Ramírez-Chaves, H. E., Martínez-Medina, D., & Calderón-Acevedo, C. A. (2021). A new species of Long-eared Brown Bat of the genus Histiotus (Chiroptera) and the revalidation of Histiotus colombiae. Caldasia, 43(2).				Argentina|Chile	South America	Neotropic	LC	0	0	0	Histiotus_macrotus	0	sciname match	Histiotus_macrotus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Histiotus		macrotus	Poeppig	1835	1	Reise Chile Peru Amaz.	0.354861	Big-eared Brown Bat	chilensis Lesson, 1836; poeppigii Fitzinger, 1872.	Chile, Bio-BIo, Antuco.	Chile, Argentina.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10201/22098780/' target='_blank'>Least Concern</a>	Does not include laephotis; see Autino et al. (1999) and Barquez and Diaz (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Histiotus macrotus; Histiotus macrotus; Eptesicus macrotus; Histiotus macrotus; Histiotus macrotus; Histiotus macrotus; chilensis; poeppigii; chilensis; poeppigii; macrotus; chilensis; poeppigii; Sérotine a grandes oreilles; Gewohnliche Gro Rohrfledermaus; Histiotus orejudo; Common Big-eared Brown Bat; Greater Leaf-eared Bat; Big-eared Brown Bat; Big-eared Brown Bat; H. macrotus