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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L453	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Histiotus laephotis	Histiotus macrotus [synonym of]	N/A	Histiotus macrotus [synonym of]	Histiotus macrotus laephotis	Histiotus laephotis	Histiotus laephotis	Eptesicus laephotis	Histiotus laephotis	Histiotus laephotis	Histiotus laephotis	Histiotus laephotis	Histiotus laephotis	Histiotus laephotis	Histiotus laephotis		[MSW3] Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al. (1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001).; [HMW] Histiotus laephotis Thomas, 1916 , “Caiza,” Bolivia . Since its description, status of H. laephotis has been confusing. It has been previously considered a full species and later a subspecies of H. montanus and H. macrotus . It is now considered a distinct species by most authors based on morphological and ecological evidence. It is frequently mistaken with H. macrotus and H. montanus , therefore, accurate data in literature are scarce. Monotypic.; [batnames2022] Subgenus Histiotus . Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007); [MDD2022] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2023] Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007); [MDD2023] has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision; [batnames2025_1.7] Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007)														laephotis				laephotis 	laephotis 			laephotis O. Thomas, 1916		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Bolivia		N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Vespertilioninae	Vespertilionini	Histiotus laephotis	Histiotus		laephotis	Thomas		1916		Ann. Mag. Nat. Hist., ser. 8	17		275		Thomas's Big-eared Brown Bat	Bolivia, Caiza.	Argentina, S Bolivia, S Peru.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al. (1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001).	4C3D87E8FFAF6A10FA5394D718F1B3DB	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	842	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FFAF6A10FA5394D718F1B3DB.xml	Histiotus laephotis	Vespertilionidae	Histiotus	laephotis	Thomas	1916	Sérotine de Bolivie @fr | Thomas-Gro Rohrfledermaus @de | Histiotus de Thomas @es	Histiotus laephotis Thomas, 1916 , “Caiza,” Bolivia . Since its description, status of H. laephotis has been confusing. It has been previously considered a full species and later a subspecies of H. montanus and H. macrotus . It is now considered a distinct species by most authors based on morphological and ecological evidence. It is frequently mistaken with H. macrotus and H. montanus , therefore, accurate data in literature are scarce. Monotypic.	S Peru , Bolivia , N Chile , E slope of Andes in NW Argentina , W & C Paraguay , and S Brazil (Santa Catarina).	Head-body ¢.57-68 mm, tail 43-59-2 mm, ear 27-39-5 mm, hindfoot 7-12 mm, forearm 44-50-1 mm; weight 9-14 g. Dorsal hairs of Thomas’s Big-eared Brown Bat are yellowish to dark brown and slightly bicolored, with variable contrast between bases and tips. Ventral hairs are bicolored, with blackish bases and yellowish tips. Ears are pale brown or yellowish, very large, and connected by conspicuous membrane; tragus is wide and long, reaching one-half the ear length. Wings are thin and naked, pale gray or brown, and slightly transparent. Muzzle is inflated and has sparse hairs. Skull is robust; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded.	Forested areas at known elevations of 350-3686 m. Thomas's Big-eared Brown Bat has been captured in Atlantic rainforest, thorn forests, and pine forests.	No information.	A lactating Thomas’s Big-eared Brown Bat was caught in November in Argentina .	Roosts of Thomas's Big-eared Brown Bats were found in a wooded storage barn. Echolocation calls have QCF components. Calls sweep down from c.38 kHz to ¢.26 kHz, with duration of c.1-3 milliseconds. Interpulse interval averages 94 milliseconds, and peak frequency is ¢.30 kHz.	No information.	Classified as Least Concern on The [UCN Red List. Thomas's Big-eared Brown Bat is not abundant locally, and its habitatis in significant decline.	Acosta & Venegas (2006) | Autino et al. (1999) | Barquez & Diaz (2001, 2008a) | Barquez etal. (1999) | Handley & Gardner (2008) | Lopez-Gonzélez et al. (1998) | Miranda et al. (2007) | Ossa et al. (2015) | Simmons (2005) | Thomas (1916b)	https://zenodo.org/record/6398178/files/figure.png	175. Thomas’s Big-eared Brown Bat Histiotus laephotis French: Sérotine de Bolivie / German: Thomas-GroRohrfledermaus / Spanish: Histiotus de Thomas Taxonomy. Histiotus laephotis Thomas, 1916 , “Caiza,” Bolivia . Since its description, status of H. laephotis has been confusing. It has been previously considered a full species and later a subspecies of H. montanus and H. macrotus . It is now considered a distinct species by most authors based on morphological and ecological evidence. It is frequently mistaken with H. macrotus and H. montanus , therefore, accurate data in literature are scarce. Monotypic. Distribution. S Peru , Bolivia , N Chile , E slope of Andes in NW Argentina , W & C Paraguay , and S Brazil (Santa Catarina). Descriptive notes. Head-body ¢.57-68 mm, tail 43-59-2 mm, ear 27-39-5 mm, hindfoot 7-12 mm, forearm 44-50-1 mm; weight 9-14 g. Dorsal hairs of Thomas’s Big-eared Brown Bat are yellowish to dark brown and slightly bicolored, with variable contrast between bases and tips. Ventral hairs are bicolored, with blackish bases and yellowish tips. Ears are pale brown or yellowish, very large, and connected by conspicuous membrane; tragus is wide and long, reaching one-half the ear length. Wings are thin and naked, pale gray or brown, and slightly transparent. Muzzle is inflated and has sparse hairs. Skull is robust; sagittal and lambdoidal crests are practically absent; zygomatic arches have strongly developed postorbital process ofjugal; palate is slightly concave centrally; caudal spine is well developed; and tympanic bullae are large and rounded. Habitat. Forested areas at known elevations of 350-3686 m. Thomas's Big-eared Brown Bat has been captured in Atlantic rainforest, thorn forests, and pine forests. Food and Feeding. No information. Breeding. A lactating Thomas’s Big-eared Brown Bat was caught in November in Argentina . Activity patterns. Roosts of Thomas's Big-eared Brown Bats were found in a wooded storage barn. Echolocation calls have QCF components. Calls sweep down from c.38 kHz to ¢.26 kHz, with duration of c.1-3 milliseconds. Interpulse interval averages 94 milliseconds, and peak frequency is ¢.30 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The [UCN Red List. Thomas's Big-eared Brown Bat is not abundant locally, and its habitatis in significant decline. Bibliography. Acosta & Venegas (2006), Autino et al. (1999), Barquez & Diaz (2001, 2008a), Barquez etal. (1999), Handley & Gardner (2008), Lopez-Gonzélez et al. (1998), Miranda et al. (2007), Ossa et al. (2015), Simmons (2005), Thomas (1916b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Eptesicus laephotis	Eptesicus	Histiotus	laephotis	Thomas	1916	1	Ann. Mag. Nat. Hist.	ser. 8, 17: 275	Thomas' Big-eared Brown Bat	None.	Bolivia, Caiza.	Argentina, S Bolivia, S Peru, Brazil.	Not listed.	Least Concern as Histiotus laephotis 	Subgenus Histiotus . Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007)	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Histiotus laephotis	23	Thomas's Big-eared Brown Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	laephotis	O. Thomas	1916	0	Histiotus_laephotis	Thomas, O. (1916). Notes on bats of the genus Histiotus. Annals and Magazine of Natural History, ser. 8, 17, 275.	https://www.biodiversitylibrary.org/item/53306#page/299/mode/1up	BM 1897.2.25.1		"Caiza," Bolivia.			laephotis O. Thomas, 1916	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA	Peru|Bolivia|Chile|Argentina|Paraguay|Brazil	South America	Neotropic	LC	0	0	0	Histiotus_laephotis	0	sciname match	Histiotus_laephotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136502	Histiotus laephotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Histiotus	laephotis	Thomas, 1916		20000000	Histiotus laephotis	Least Concern		2019	2019-02-05 00:00:00 UTC	3.1	English	This species is listed as Least Concern because it is widely distributed, including in a variety of habitats. Although the core of its distribution is dependent upon a highly fragile habitat (southern montane forests, or Yungas), current population decline due to human population density and habitat transformation does not reach the threshold (&gt;30% over ten years) to qualify as threatened under criterion A2c.	This species is found in the interior of forests, but records from southern Peru and northern Chile (RodrÃ­guez-San Pedro et al . 2016) include arid formations. It is usually found using buildings and tree hollows as diurnal roosts. Barquez et al . (1999) recorded the use of an artificial diurnal roost by H. laephotis , as well as the cohabitation of this species with other bat species (Tadarida brasiliensis and Myotis sp). This type of behavior (sharing roost) tends to occur among all species in Histiotus (Miranda et al. 2007). As with other species in the genus, it is mostly insectivorous.	The species shows a wide distribution, but at the Yungas formations (montane forests in S Peru, S Bolivia and N Argentina) it may face some risks because of high rates of deforestation by changes in land use.	It is locally not abundant throughout its geographic distribution; usually rare. Few specimens and records are available.	Decreasing	The species is present in Argentina, south of Bolivia, and south of Peru (Simmons 2005, Handley and Gardner 2008). There are records from Paraguay (LÃ³pez-GonzÃ¡lez et al . 1998), and Chile (RodrÃ­guez-San Pedro et al . 2016). It has been recorded at an Araucaria pine forest in the State of Santa Catarina, Southern Brazil (Miranda et al . 2009).		Terrestrial	There is a need to reduce habitat loss and update the species distribution, focusing on the diversity of habitats.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Histiotus		laephotis	Thomas	1916	0	Ann. Mag. Nat. Hist.	ser. 8, 17: 275	Thomas' Big-eared Brown Bat	None.	Bolivia, Caiza.	Argentina, S Bolivia, S Peru, Brazil.	Not listed.	Least Concern	Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007)	Histiotus laephotis	1005556	23	Thomas's Big-eared Brown Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	VESPERTILIONINAE	EPTESICINI	Histiotus	NA	laephotis	O. Thomas	1916	0	Histiotus_laephotis	Thomas, O. (1916). Notes on bats of the genus Histiotus. Annals and Magazine of Natural History, ser. 8, 17, 275.	https://www.biodiversitylibrary.org/item/53306#page/299/mode/1up	BM 1897.2.25.1		"Caiza," Bolivia.			laephotis O. Thomas, 1916	has been included under Eptesicus, but the genus Histiotus is tentatively retained here pending a full revision	NA				Peru|Bolivia|Chile|Argentina|Paraguay|Brazil	South America	Neotropic	LC	0	0	0	Histiotus_laephotis	0	sciname match	Histiotus_laephotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Histiotus_laephotis	1005556	23	Thomas's Big-eared Brown Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Vespertilioninae	Nycticeiini	Histiotus	NA	laephotis	O. Thomas	0	Histiotus laephotis	Thomas, O. 1916-03-01. Notes on bats of the genus _Histiotus_. Annals and Magazine of Natural History (8)17(99):272-276.	https://www.biodiversitylibrary.org/page/15612445	BMNH:Mamm:1897.2.25.1	holotype	https://data.nhm.ac.uk/object/35775ebb-247d-47ed-8121-83c7e003bb44	"Caiza," Bolivia.			NA	NA				Peru|Bolivia|Chile|Argentina|Paraguay|Brazil	South America	Neotropic	LC	0	0	0	Histiotus_laephotis	0	sciname match	Histiotus_laephotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Histiotus		laephotis	Thomas	1916	0	Ann. Mag. Nat. Hist.	ser. 8, 17: 275	Thomas' Big-eared Brown Bat	None.	Bolivia, Caiza.	Argentina, S Bolivia, S Peru, Brazil.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136502/21974854/' target='_blank'>Least Concern</a>	Treated as a subspecies of montanus by Anderson (1997) and as a subspecies of macrotis by Koopman (1994) and Barquez et al.(1993, 1999), but apparently distinct; see Autino et al. (1999) and Barquez and Diaz (2001). Brazilian record from Miranda et al. (2007)		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Histiotus laephotis; Histiotus laephotis; Eptesicus laephotis; Histiotus laephotis; Histiotus laephotis; Histiotus laephotis; laephotis; Sérotine de Bolivie; Thomas-Gro Rohrfledermaus; Histiotus de Thomas; Thomas's Big-eared Brown Bat; Thomas's Big-eared Brown Bat; Thomas' Big-eared Brown Bat; H. laephotis
