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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L46	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Amorphochilus schnabli	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii	Amorphochilus schnablii		[HMW] Amorphochilus schnablii Peters, 1877 . “Tumbez un noérdlichen Peru , an der Grenze von Ecuador .” Restricted by A. Cabrera in 1958 to Tumbes , departamento Tumbes , Peru . This species is monotypic.; [batnames2022] For a revised distribution map see Guerra et al. (2020).; [batnames2023] For a revised distribution map see Guerra et al. (2020).; [batnames2025_1.7] For a revised distribution map see Guerra et al. (2020).						osgoodi.			osgoodi			schnablii	schnablii - osgoodi	schnablii, osgoodi		schnablii	schnablii - osgoodi	schnablii, osgoodi	schnablii, osgoodi, chnablii, schnabli	schnablii	schnablii - osgoodi	schnablii W. C. H. Peters, 1877|osgoodi J. A. Allen, 1914|schnabli Osgood, 1914 [incorrect subsequent spelling]|chnablii Tovar S., 1971 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		W Ecuador – N Chile	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Amorphochilus schnablii	Peru, Tumbes, Tumbes.	Peters	1877	Monatsb. Preuss. Akad. Wiss. Berlin, p. 185.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		W Ecuador – N Chile	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1877	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1877:185.		W Peru, W Ecuador, Puna Isl (Ecuador), N Chile.	Peru, Tumbes, Tumbes.		PETERS	1877	Size fairly small (forearm length, 34-38 mm).	Distribution: Same as for genus.	No subspecies are here recognized.		96	species	A. schnablii	PETERS	1877	Amorphochilus	genus	Amorphochilus schnablii				Size fairly small (forearm length, 34-38 mm).	No subspecies are here recognized.		1. A. schnablii PETERS 1877.	1	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Furipteridae			Amorphochilus schnablii	Amorphochilus		schnablii	Peters		1877		Monatsb. K. Preuss. Akad. Wiss. Berlin	1877		185		Smoky Bat	Peru, Tumbes, Tumbes.	W Peru, W Ecuador, Puna Isl (Ecuador), N Chile.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.	osgoodi J. A. Allen, 1914.		039187AC5C3F2A52F099001F399BF24E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Furipteridae_412.pdf.imf	hash://md5/ffa8ffd45c3e2a53f03609113f64ffc4	416	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/91/87/039187AC5C3F2A52F099001F399BF24E.xml	Amorphochilus schnablii	Furipteridae	Amorphochilus	schnablii	Peters	1877	Furie de Schnabl @fr | Felsenklsten-Stummeldaumen @de | Murciélagoahumado @es | Schnabl's Smoky Bat @en	Amorphochilus schnablii Peters, 1877 . “Tumbez un noérdlichen Peru , an der Grenze von Ecuador .” Restricted by A. Cabrera in 1958 to Tumbes , departamento Tumbes , Peru . This species is monotypic.	Known only from a narrow strip Wofthe Andes in C & S Ecuador (including Puna I), Peru , and N Chile .	Head-body 39-47 mm, tail 27-34 mm, ear 12-15 mm, hindfoot 6-10 mm, forearm 34-38 mm; weight 3-10 g. Tail extends nearly fourfifths the length of uropatagium andis entirely enclosed by that membrane. Height of braincase, including auditory bullae, is equal to distance from frontal angle to most posterior point of occipital region. Muzzleis relatively simple and well-furred; no fleshystructures occur under chin; palateis short, not extending much beyond last molar; and mesopterygoid fossa is longer than wide. The Smoky Bat has long, dark smokygrayfur, often with brownish tinge; hair tips are blackish; front and top of head have pale buffy suffusion; underparts are nearly the same color as the upperparts; ears are light brown; and wing and tail membranes arelittle darker than ears. Snout is prominently rimmedand blunt; lips are conspicuouslyfleshy; and chin excrescences and lowerlips havefleshy wart-like structures. On average, third metacarpal is 34 mm , tibia is 15-5 mm, total skull length is 12 mm , braincase breadthis 6 mm , and uppertooth rowlength (with canine) is 5 mm . Dental formula is I 2/3, Cl1/1,P2/3.M 3/3 (x2) =36.	South American west coast deserts from sea level up to elevations of 2680 m . Two Smoky Bats were found on Hacienda Limon, Cajamarca , Peru , in a small arid valley above the Rio Maranon in the Amazon Basin. It is found in arid regions and cultivated areas; it is a cave-dweller and known to roost in culverts, caves, sugar mills, irrigation tunnels, and abandoned urban-rural buildings near crop areas (banana, rice; etc.).	Smoky Bats eat moths and butterflies (Lepidoptera). Adult moth scales were found in stomachs offive specimens from central Ecuador .	In central Ecuador in middle November, a large colony included males that did not show testes in the scrotal position, and several females were pregnant (eight of ten) with single fetuses but not lactating. This high percentage of pregnant females suggested a relatively synchronized seasonal breeding period, undoubtedly related to the rainy season (January-May); similarly, a breeding colony was found in the rainy months (December-March) in southern Peru . In a single reproductively inactive male, the small, paired testes were attached to the posterior body wall and inferior pole of the kidney by a suspensoryligament. The Smoky Bat has an ampullary gland. One specimen appeared to contain a tiny baculum, capping the urethra at tip ofglans.	Echolocation calls of the SmokyBat are the FMtype, with an average duration of4-4 milliseconds, important for foraging in dense vegetation and taking small insects while flying. Maximum averagefrequencyat 75-8 kHz.	In Guayas , Ecuador , a large monospecific colony(c.300 individuals) of Smoky Bats included males and females but not juveniles. This bat species has been collected along with Long-snouted Bat ( Platalina genovensium ) and Small Big-eared Brown Bat ( Histiotus montanus).	Classified as Vulnerable on The IUCNRed List. The Smoky Bat is considered an endangered species in Ecuador because known populations are small, restricted to areas smaller than 20 km ?, and known fromless than five localities. In Peru , it is considered a vulnerable species. In Chile , it has not been evaluated, but it is thought to be beneficial to agroforestry.	Abuja (1999) | Aragon & Aguirre (2014) | Cabrera (1958) | Gardner (2008h) | Ibanez (1986) | Krutzsch (2000) | Miller (1907) | Morgan & Czaplewski (1999) | Nowak (1994) | Rodriguez-San Pedro et al. (2016) | Tirira et al. (2012) | Ugarte-Nunez (2014)	https://zenodo.org/record/5733448/files/figure.png	1. Smoky Bat Amorphochilus schnablii French: Furie de Schnabl / German: Felsenklsten-Stummeldaumen / Spanish: Murciélago ahumado Other common names: Schnabl's Smoky Bat Taxonomy. Amorphochilus schnablii Peters, 1877 . “Tumbez un noérdlichen Peru , an der Grenze von Ecuador .” Restricted by A. Cabrera in 1958 to Tumbes , departamento Tumbes , Peru . This species is monotypic. Distribution. Known only from a narrow strip Wofthe Andes in C & S Ecuador (including Puna I), Peru , and N Chile . Descriptive notes. Head-body 39-47 mm, tail 27-34 mm, ear 12-15 mm, hindfoot 6-10 mm, forearm 34-38 mm; weight 3-10 g. Tail extends nearly fourfifths the length of uropatagium andis entirely enclosed by that membrane. Height of braincase, including auditory bullae, is equal to distance from frontal angle to most posterior point of occipital region. Muzzleis relatively simple and well-furred; no fleshystructures occur under chin; palateis short, not extending much beyond last molar; and mesopterygoid fossa is longer than wide. The Smoky Bat has long, dark smokygrayfur, often with brownish tinge; hair tips are blackish; front and top of head have pale buffy suffusion; underparts are nearly the same color as the upperparts; ears are light brown; and wing and tail membranes arelittle darker than ears. Snout is prominently rimmedand blunt; lips are conspicuouslyfleshy; and chin excrescences and lowerlips havefleshy wart-like structures. On average, third metacarpal is 34 mm , tibia is 15-5 mm, total skull length is 12 mm , braincase breadthis 6 mm , and uppertooth rowlength (with canine) is 5 mm . Dental formula is I 2/3, Cl1/1,P2/3.M 3/3 (x2) =36. Habitat. South American west coast deserts from sea level up to elevations of 2680 m . Two Smoky Bats were found on Hacienda Limon, Cajamarca , Peru , in a small arid valley above the Rio Maranon in the Amazon Basin. It is found in arid regions and cultivated areas; it is a cave-dweller and known to roost in culverts, caves, sugar mills, irrigation tunnels, and abandoned urban-rural buildings near crop areas (banana, rice; etc.). Food and Feeding. Smoky Bats eat moths and butterflies (Lepidoptera). Adult moth scales were found in stomachs offive specimens from central Ecuador . Breeding. In central Ecuador in middle November, a large colony included males that did not show testes in the scrotal position, and several females were pregnant (eight of ten) with single fetuses but not lactating. This high percentage of pregnant females suggested a relatively synchronized seasonal breeding period, undoubtedly related to the rainy season (January-May); similarly, a breeding colony was found in the rainy months (December-March) in southern Peru . In a single reproductively inactive male, the small, paired testes were attached to the posterior body wall and inferior pole of the kidney by a suspensoryligament. The Smoky Bat has an ampullary gland. One specimen appeared to contain a tiny baculum, capping the urethra at tip ofglans. Activity patterns. Echolocation calls of the SmokyBat are the FMtype, with an average duration of4-4 milliseconds, important for foraging in dense vegetation and taking small insects while flying. Maximum averagefrequencyat 75-8 kHz. Movements, Home range and Social organization. In Guayas , Ecuador , a large monospecific colony(c.300 individuals) of Smoky Bats included males and females but not juveniles. This bat species has been collected along with Long-snouted Bat ( Platalina genovensium ) and Small Big-eared Brown Bat ( Histiotus montanus). Status and Conservation. Classified as Vulnerable on The IUCNRed List. The Smoky Bat is considered an endangered species in Ecuador because known populations are small, restricted to areas smaller than 20 km ?, and known fromless than five localities. In Peru , it is considered a vulnerable species. In Chile , it has not been evaluated, but it is thought to be beneficial to agroforestry. Bibliography. Abuja (1999), Aragon & Aguirre (2014), Cabrera (1958), Gardner (2008h), Ibanez (1986), Krutzsch (2000), Miller (1907), Morgan & Czaplewski (1999), Nowak (1994), Rodriguez-San Pedro et al. (2016), Tirira et al. (2012), Ugarte-Nunez (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Furipteridae	Amorphochilus schnablii	Amorphochilus		schnablii	Peters	1877	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1880:05:00	Smoky Bat	 osgoodi J.A. Allen, 1914	Peru, Tumbes, Tumbes	W Peru, W Ecuador, Puna Isl (Ecuador), N Chile	Not listed.	Vulnerable	For a revised distribution map see Guerra et al. (2020).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Amorphochilus schnablii	23	Smoky Bat	Schnabl's Smoky Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	FURIPTERIDAE	NA	NA	Amorphochilus	NA	schnablii	W. Peters	1877	0						"Tumbez im nördlichen Peru, an der Grenze von Ecuador." Restricted by A. Cabrera in 1958 to Tumbes, departamento Tumbes, Peru.			schnablii W. Peters, 1877|osgoodi J. A. Allen, 1914	NA	NA	Ecuador|Peru|Chile	South America	Neotropic	VU	0	0	0	Amorphochilus_schnablii	0	sciname match	Amorphochilus_schnablii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	1154	Amorphochilus schnablii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	FURIPTERIDAE	Amorphochilus	schnablii	Peters, 1877		20000000	Amorphochilus schnablii	Vulnerable	A2c	2015	2014-07-14 00:00:00 UTC	3.1	English	This species is listed as Vulnerable because of a population decline, estimated to be more than 30% over the past three generations (where generation length is six years; Pacifici et al. 2013), which is suspected from an observed shrinkage in distribution and information on habitat destruction and degradation.	It is found in arid regions, cultivated areas, culverts, caves and abandoned urban-rural buildings near crop areas (banana, rice, etc.; Ibáñez 1985, Aragón A and Aguirre Q 2014). Adult moths have been found in the stomachs of A. schnablii (Ibáñez 1985, Ortiz de la Puente 1951). Pregnant females were reported from Ecuador (mid November) and Peru (mid February), each one with a single foetus, suggesting that births and lactation are synchronized with the brief rainy season (January to May; Ibáñez 1985, L. Huamani and R. Cadenillas pers. comm.).	The major threat that this species faces is the destruction of its refuges because of increased cultivation. Use in witchcraft activities is also a threat. Population reduction was observed in a large refuge in northern Peru, where the population decreased by 25% in only seven months. This population decrease occurred due to man-made fire inside the refuge (L. Huamaní and R. Cadenillas pers. comm.).	It is found in small groups of one to five, up to 300 individuals (Ibáñez 1985, L. Huamaní pers. comm.). It might be more common that previously suspected, as recent acoustic ;records suggest it is present (even abundant) along inter-Andean valleys in southwestern Peru (J. Ugarte pers. comm.).	Stable	This species is known to occur west of the Andes in southern Ecuador, including Puná Island, Peru, and northern Chile, with the exception of one record from the department of Cajamarca in highlands of northern Peru (Gardner 2008).	There is occasional use of individuals for folkloric (witchcraft) activities.	Terrestrial	The most important conservation actions should be the protection of the refuges where these species occur and the education of local people through workshops on the importance of this species in the ecosystem.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Furipteridae	Amorphochilus		schnablii	Peters	1877	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1880:05:00	Smoky Bat	 osgoodi J.A. Allen, 1914	Peru, Tumbes, Tumbes	W Peru, W Ecuador, Puna Isl (Ecuador), N Chile	Not listed.	Vulnerable	For a revised distribution map see Guerra et al. (2020).	Amorphochilus schnablii	1004848	23	Smoky Bat	Schnabl's Smoky Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Furipteridae	NA	NA	Amorphochilus	NA	schnablii	W. Peters	1877	0						"Tumbez im nördlichen Peru, an der Grenze von Ecuador." Restricted by A. Cabrera in 1958 to Tumbes, departamento Tumbes, Peru.			schnablii W. Peters, 1877|osgoodi J. A. Allen, 1914	NA	NA				Ecuador|Peru|Chile	South America	Neotropic	VU	0	0	0	Amorphochilus_schnablii	0	sciname match	Amorphochilus_schnablii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Amorphochilus_schnablii	1004848	23	Smoky Bat	Schnabl's Smoky Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Furipteridae	NA	NA	Amorphochilus	NA	schnablii	W. C. H. Peters	0	Amorphochilus Schnablii	Peters, W.C.H. 1877. Ãœber eine neue Gattung von Flederthieren, _Amorphochilus_, aus Peru und über eine neue _Crocidura_ aus Liberia. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1877:184-188.	https://www.biodiversitylibrary.org/page/35723464				"Tumbez im nördlichen Peru, an der Grenze von Ecuador." Restricted by A. Cabrera in 1958 to Tumbes, departamento Tumbes, Peru.			NA	NA				Ecuador|Peru|Chile	South America	Neotropic	VU	0	0	0	Amorphochilus_schnablii	0	sciname match	Amorphochilus_schnablii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Furipteridae	Amorphochilus		schnablii	Peters	1877	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	1880:05:00	Smoky Bat	osgoodi J.A. Allen, 1914	Peru, Tumbes, Tumbes	W Peru, W Ecuador, Puna Isl (Ecuador), N Chile	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/1154/22070889/' target='_blank'>Vulnerable</a>	For a revised distribution map see Guerra et al. (2020).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Amorphochilus schnablii; Amorphochilus schnabli; schnablii; osgoodi; Furie de Schnabl; Felsenklsten-Stummeldaumen; Murciélago ahumado; Schnabl's Smoky Bat; Smoky Bat; Schnabl's Smoky Bat; Smoky Bat; Smoky Bat; Amorphochilus schnablii osgoodi; A. schnablii