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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L441	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Doryrhina stenotis	Hipposideros stenotis	Doryrhina stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis	Hipposideros stenotis		[MSW3] cyclops species group.; [HMW] Hipposideros stenotis Thomas, 1913 , “Mary River,” Northern Territory , Australia . This species is monotypic.; [batnames2022] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group.; [MDD2022] moved from Hipposideros to Doryrhina; [batnames2023] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group.; [MDD2023] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [MDD2025_2.0] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [batnames2025_1.7] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group.; [MDD2025_2.2] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being														stenotis				stenotis	stenotis			stenotis O. Thomas, 1913		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Narrow-eared leaf-nosed	NW, N Australia bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros stenotis	Australia, Northern Territory, Mary River.	Thomas	1913	Ann. Mag. Nat. Hist., ser. 8, 12:206.	Distribution: Confined to the dryer portions of tropical Australia from northern Western Australia to northwestern Queensland.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Narrow-eared leaf-nosed	NW, N Australia bat	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1913	Ann. Mag. Nat. Hist., ser. 8, 12:206.		Northern Territory, N Western Australia and N Queensland (Australia). A New Guinea record is probably erroneous, see Hill (1963b:87).	Australia, Northern Territory, Mary River.		THOMAS	1913	Posterior lateral leaflet extending anteriorly be neath the anterior noseleaf. Premaxillae narrow, not enclosing the anterior palatal foramina, mak ing a narrow spatulate junction with the maxillae. Median process of intermediate noseleaf well de veloped and club-shaped. Skull with a deep frontal depression. Sphenoidal depression absent. Posterior ridge of last upper molar little reduced. Width of each cochlea at least six times their dis tance apart. Interorbital region relatively wide. Median process of posterior noseleaf poorly de veloped. Rostrum relatively short, its eminences moderately inflated. Antorbital foramen relative ly large. Size relatively small (forearm length, 42-45 mm).	Distribution: Confined to the dryer portions of tropical Australia from northern Western Australia to northwestern Queensland.	No subspecies.		64	species	H. stenotis	THOMAS	1913	Hipposideros	genus	Hipposideros stenotis				Posterior lateral leaflet extending anteriorly be neath the anterior noseleaf. Premaxillae narrow, not enclosing the anterior palatal foramina, mak ing a narrow spatulate junction with the maxillae. Median process of intermediate noseleaf well de veloped and club-shaped. Skull with a deep frontal depression. Sphenoidal depression absent. Posterior ridge of last upper molar little reduced. Width of each cochlea at least six times their dis tance apart. Interorbital region relatively wide. Median process of posterior noseleaf poorly de veloped. Rostrum relatively short, its eminences moderately inflated. Antorbital foramen relative ly large. Size relatively small (forearm length, 42-45 mm).	No subspecies.		39. H. stenotis THOMAS 1913 [cyclops group],	39	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros stenotis	Hipposideros		stenotis	Thomas		1913		Ann. Mag. Nat. Hist., ser. 8	12		206		Narrow-eared Leaf-nosed Bat	Australia, Northern Territory, Mary River.	Northern Territory, N Western Australia and N Queensland (Australia). A New Guinea record is probably erroneous, see Hill (1963b:87).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		cyclops species group.	03BD87A2C672A201F8ADF264FEB245CC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	231	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C672A201F8ADF264FEB245CC.xml	Doryrhina stenotis	Hipposideridae	Doryrhina	stenotis		1913	Phyllorhine à oreilles fines @fr | Nördliche Rundblattnase @de | Doryrina nortena @es | Narrow-eared Leaf-nosed Bat @en | Narrow-eared Roundleaf Bat @en	Hipposideros stenotis Thomas, 1913 , “Mary River,” Northern Territory , Australia . This species is monotypic.	Endemic to Australia , including NE Western Australia , N Northern Territory , and NW Queensland (Mt Isa); also on several offshore Is such as Boongaree, Koolan, and Bathurst Is.	Head-body 40-46 mm, tail 26-27 mm, ear 19-21 mm, forearm 42—46 mm ; weight 4-6-6-4 g. The Northern Leaf-nosed Bat is very similar to Semon’s Leaf-nosed Bat { D. semoni ), but considerably smaller. Both species present complex noseleafwith two supplementary leaflets: anterior leaflet is short and broad, and posterior leaflet extends beneath anterior leaf onto upper lip. The Northern Leaf-nosed Bat also presents processes in intermediate and posterior leaves, but differs by having median process of posterior leaf less prominent. Ears are long with pointed tips, and rostral eminences are moderately inflated. Fur is brownish dorsally, paler ventrally.	The Northern Leaf-nosed Bat occurs at low densities and occupies a wide variety of habitats. It has been recorded in areas with sandstone cliffs, escarpments, gorges and waterholes, tall open forest, floodplains, and open meadows. Elevational range is unknown.	The Northern Leaf-nosed Bat forages both in woodland and on open hilly plains, flying slowly with frequent changes of direction and great ability to maneuver. It mainly hunts flying insects close to the vegetation.	Between October and January, females give birth to a single young. Northern Leaf-nosed Bats form maternity colonies.	The Northern Leaf-nosed Bat can roost in sandstone and limestone caves, abandoned mines, crevices and boulder caves. It is thought that this species prefers low humidity, roosting near to cave entrances. This species emits echolocation calls at 102-106 kHz.	Northern Leaf-nosed Bats have been observed roosting alone, in pairs, or in small maternity colonies of 6-12 individuals.	Classified as Least Concern on The IUCN Red List (as Hipposideros stenotis ). Population size and trends are unknown; population is probably not greatly fragmented and is not declining markedly. No major threats are known, although destruction and disturbance of roosts are increasing. Habitat loss and degradation might also be significant, as well as reported predation by feral cats.	Churchill (2008) | Duncan et al. (1999) | Flannery & Colgan (1993) | Hourigan (2011) | Menkhorst & Knight (2001) | Milne & Hall (2008) | Schulz & Menkhorst (1986) | Thomson & McKenzie (2008) | Woinarski et al. (2014)	https://zenodo.org/record/3749059/files/figure.png	11. Northern Leaf-nosed Bat Doryrhina stenotis French: Phyllorhine à oreilles fines / German: Nördliche Rundblattnase / Spanish: Doryrina nortena Other common names: Narrow-eared Leaf-nosed Bat , Narrow-eared Roundleaf Bat Taxonomy. Hipposideros stenotis Thomas, 1913 , “Mary River,” Northern Territory , Australia . This species is monotypic. Distribution. Endemic to Australia , including NE Western Australia , N Northern Territory , and NW Queensland (Mt Isa); also on several offshore Is such as Boongaree, Koolan, and Bathurst Is. Descriptive notes. Head-body 40-46 mm, tail 26-27 mm, ear 19-21 mm, forearm 42—46 mm ; weight 4-6-6-4 g. The Northern Leaf-nosed Bat is very similar to Semon’s Leaf-nosed Bat { D. semoni ), but considerably smaller. Both species present complex noseleafwith two supplementary leaflets: anterior leaflet is short and broad, and posterior leaflet extends beneath anterior leaf onto upper lip. The Northern Leaf-nosed Bat also presents processes in intermediate and posterior leaves, but differs by having median process of posterior leaf less prominent. Ears are long with pointed tips, and rostral eminences are moderately inflated. Fur is brownish dorsally, paler ventrally. Habitat. The Northern Leaf-nosed Bat occurs at low densities and occupies a wide variety of habitats. It has been recorded in areas with sandstone cliffs, escarpments, gorges and waterholes, tall open forest, floodplains, and open meadows. Elevational range is unknown. Food and Feeding. The Northern Leaf-nosed Bat forages both in woodland and on open hilly plains, flying slowly with frequent changes of direction and great ability to maneuver. It mainly hunts flying insects close to the vegetation. Breeding. Between October and January, females give birth to a single young. Northern Leaf-nosed Bats form maternity colonies. Activity patterns. The Northern Leaf-nosed Bat can roost in sandstone and limestone caves, abandoned mines, crevices and boulder caves. It is thought that this species prefers low humidity, roosting near to cave entrances. This species emits echolocation calls at 102-106 kHz. Movements, Home range and Social organization. Northern Leaf-nosed Bats have been observed roosting alone, in pairs, or in small maternity colonies of 6-12 individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List (as Hipposideros stenotis ). Population size and trends are unknown; population is probably not greatly fragmented and is not declining markedly. No major threats are known, although destruction and disturbance of roosts are increasing. Habitat loss and degradation might also be significant, as well as reported predation by feral cats. Bibliography. Churchill (2008), Duncan et al. (1999), Flannery & Colgan (1993), Hourigan (2011), Menkhorst & Knight (2001), Milne & Hall (2008), Schulz & Menkhorst (1986), Thomson & McKenzie (2008), Woinarski et al. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros stenotis	Hipposideros		stenotis	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 12: 206	Narrow-eared Leaf-nosed Bat	None.	Australia, Northern Territory, Mary River.	Northern Territory, N Western Australia and N Queensland (Australia). A New Guinea record is probably erroneous, see Hill (1963b:87).	Not listed.	Vulnerable	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Doryrhina stenotis	23	Northern Leaf-nosed Bat	Narrow-eared Leaf-nosed Bat|Narrow-eared Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Doryrhina	NA	stenotis	O. Thomas	1913	1	Hipposideros_stenotis	Thomas, O. (1913). On new mammals obtained by the Utakwa Expedition to Dutch New Guinea. The Annals and magazine of natural history; zoology, botany, and geology, ser. 8, 12, 206.	https://www.biodiversitylibrary.org/item/71910#page/218/mode/1up	BM 1897.4.12.7		"Mary River," Northern Territory, Australia.			stenotis (O. Thomas, 1913)	moved from Hipposideros to Doryrhina	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.	Australia	Oceania	Australasia/Oceania	VU	0	0	0	Doryrhina_stenotis	0	sciname match	Hipposideros_stenotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10163	Hipposideros stenotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	stenotis	Thomas, 1913		20000000	Hipposideros stenotis	Vulnerable	C1+2a(i)	2021	2020-11-28 00:00:00 UTC	3.1	English	Hipposideros stenotis is assessed as Vulnerable under criteria C1+2a(i). The species occurs at low density and the overall population is presumed to be at least 10,000 individuals (ca 5,000 mature individuals) with the largest subpopulations having ca 500 mature individuals. An estimated decline of 5–10% over 15 years (three generations; GL = 5 yrs; Pacifici et al. 2013) is estimated from the loss of several known colonies and a reduction in reporting rates (just five new records in a recent 10-year period in the Northern Territory (Milne and Pavey 2011).	Hipposideros stenotis is found in rugged rocky areas, such as sandstone cliffs, escarpments and boulder country, gorges, and waterholes bordered by paperbark trees. It roosts in boulder piles, crevices, road culverts, old mines, shallow caves, and semi-shaded sites along cliff lines, and is usually found singly or in small groups. They forage close to the ground in a range of vegetation types, including eucalypt open forests and woodlands and grasslands, but typically where these are close to escarpments and other rocky areas. The diet comprises small insects, including beetles and moths (Milne et al. 2006, Churchill 2008, Milne and Hall 2008, Milne and Pavey 2011, Woinarski et al. 2014). There appears to be some geographic variation in echolocation call characteristic frequency (McKenzie and Bullen 2012).	Key threats have not been identified that would likely contribute to a significant decline of this species in the short-term, but habitat degradation through high fire frequency, which occurs in a significant part of its habitat, is the most likely factor to have a negative effect on area of occupancy and local survival rates.	The species is rare and occurs at low density across its range. The overall population is presumed to be at least 10,000 individuals (ca 5,000 mature individuals) with the largest subpopulations having ca 500 mature individuals. An estimated decline of 5-10% over 15 years (three generations, GL = 5 yrs, Pacifici et al. ; 2013) is inferred from the loss of several known colonies and a reduction in reporting rates (just five new records in a recent 10-year period in the Northern Territory (Milne and Pavey 2011). Further, surveys of Kakadu National Park, the region where most records have occurred, only encountered H. stenotis in one locality (Milne and Hall 2008).	Unknown	Hipposideros stenotis occurs in a series of subpopulations across the Kimberley region of Western Australia, the Top End and the Gulf Coastal bioregion of the Northern Territory, and the Mount Isa Inlier bioregion of north-western Queensland. Within this extensive range it is very patchily distributed (Milne et al. 2006, Milne and Hall 2008, Milne and Pavey 2011). It has been recorded from several offshore islands in the Kimberley region, including Bathurst, Bigge, Boongaree, Coronation, Gibbings, Irvine, Jungulu, Koolan, Lachlan, Middle Osborn, Sir Graham Moore, St Andrew, Storr, unnamed (west of Storr), Uwins, Wargul and Wulalum Islands (McKenzie et al. 1995, McKenzie and Bullen 2012). Interestingly, it is not known from any islands in the Northern Territory (D. Milne pers. comm.).		Terrestrial	The species is present in several protected areas where fire management is a priority, but it is not part of a targeted management effort. There has been some management of abandoned mines in the Kakadu area, including consideration of their use by bats. Much of its biology relevant to effective management is poorly known. There is a requirement for further study on broad habitat preference through modelling and surveys, roost site preference and breeding requirements, an estimate of population size, seasonal and spatial patterning of foraging habitat use, a requirement to emphasise the need for targeted acoustic detection on environmental surveys within its range, a need for investigation of population level genetic similarity across its range, and monitoring of known occurrences to confirm suspicions of a decline (Woinarski et al. 2014).	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		stenotis	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 12: 206	Narrow-eared Leaf-nosed Bat	None.	Australia, Northern Territory, Mary River.	Northern Territory, N Western Australia and N Queensland (Australia). A New Guinea record is probably erroneous, see Hill (1963b:87).	Not listed.	Vulnerable	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group.	Hipposideros stenotis	1004572	23	Northern Leaf-nosed Bat	Narrow-eared Leaf-nosed Bat|Narrow-eared Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	stenotis	O. Thomas	1913	0	Hipposideros_stenotis	Thomas, O. (1913). On new mammals obtained by the Utakwa Expedition to Dutch New Guinea. The Annals and magazine of natural history; zoology, botany, and geology, ser. 8, 12, 206.	https://www.biodiversitylibrary.org/item/71910#page/218/mode/1up	BM 1897.4.12.7		"Mary River," Northern Territory, Australia.			stenotis (O. Thomas, 1913)	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Australia	Oceania	Australasia/Oceania	VU	0	0	0	Doryrhina_stenotis	0	sciname match	Hipposideros_stenotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_stenotis	1004572	23	Northern Roundleaf Bat	Narrow-eared Leaf-nosed Bat|Narrow-eared Roundleaf Bat|Northern Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	stenotis	O. Thomas	0	Hipposideros stenotis	Thomas, O. 1913-08-01. On new mammals obtained by the Utakwa Expedition to Dutch New Guinea. Annals and Magazine of Natural History (8)12(68):205-212.	https://www.biodiversitylibrary.org/page/22099693	BMNH:Mamm:1897.4.12.7	holotype	https://data.nhm.ac.uk/object/2318a985-b66b-4741-aa2c-2ffec72b4886	"Mary River," Northern Territory, Australia.			has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Australia	Oceania (Continent)	Australasia	VU	0	0	0	Doryrhina_stenotis	0	sciname match	Hipposideros_stenotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		stenotis	Thomas	1913	0	Ann. Mag. Nat. Hist.	ser. 8, 12: 206	Narrow-eared Leaf-nosed Bat	None.	Australia, Northern Territory, Mary River.	Northern Territory, N Western Australia and N Queensland (Australia). A New Guinea record is probably erroneous, see Hill (1963b:87).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10163/22099463/' target='_blank'>Vulnerable</a>	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros sumbae; Hipposideros sumbae; Hipposideros sumbae; Hipposideros sumbae; Hipposideros sumbae; Hipposideros sumbae; sumbae; rotiensis; sumbawae; sumbae; rotiensis; sumbawae; rotiensis; sumbawae; sumbae; rotiensis; sumbawae; Phyllorhine de Sumba; Sumba-Rundblattnase; Hiposidérido de Samba; Sumba Roundleaf Bat; Sumban Leaf-nosed Bat; Sumba Roundleaf Bat; Sumban Leaf-nosed Bat; Sumban Leaf-nosed Bat; H. sumbae