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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L438	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni	Hipposideros sorenseni		[MSW3] larvatus species group.; [HMW] Hipposideros sorenseni Kitchener & Maryanto, 1993 , “ Gua Kramat (s holy cave), Pangandaran, W[est]. Java (c. 7° 41’S , 108° 40’E ).” Hipposideros sorenseni is in the larvatus species group. This species was initially allocated to H. larvatus . Monotypic.; [batnames2022]  larvatus species group.; [batnames2023]  larvatus species group.; [batnames2025_1.7] larvatusspecies group.														sorenseni				sorenseni 	sorenseni 			sorenseni D. J. Kitchener & Maryanto, 1993						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros sorenseni	Hipposideros		sorenseni	Kitchener and Maryanto		1993		Rec. West. Aust. Mus.	16		142		Sorensen's Leaf-nosed Bat	Indonesia, W Java, Pangandaran, Gua Karmat (= holy cave)(c. 7°41’S, 108°40’E).	C and W Java (Indonesia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		larvatus species group.	03BD87A2C67AA209F8A7F031FA57453A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	239	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67AA209F8A7F031FA57453A.xml	Hipposideros sorenseni	Hipposideridae	Hipposideros	sorenseni	Kitchener & Maryanto	1993	Phyllorhine de Sorensen @fr | Sorensen-Rundblattnase @de | Hiposidérido de Sorensen @es | Sorensen's Roundleaf Bat @en	Hipposideros sorenseni Kitchener & Maryanto, 1993 , “ Gua Kramat (s holy cave), Pangandaran, W[est]. Java (c. 7° 41’S , 108° 40’E ).” Hipposideros sorenseni is in the larvatus species group. This species was initially allocated to H. larvatus . Monotypic.	Known only from the type locality in WGJava.	Head-body 50-2-60 mm, tail 30-1-36-2 mm, ear 20-2-22-8 mm, forearm 55-4-60-2 mm. Greatest lengths of skulls are 21-1-22-1 mm, tibia 21-4-23-3 mm. Ears of Sorensen’s Leaf-nosed Bat are large and triangular, slightly concave before reaching tip. Anterior noseleaf has small depression in center and three supplementary lateral leaflets. Nasal septum is slighdy inflated. Upper border of posterior leaf is semicircular and there are three septa forming four noticeable cells on frontal surface, delimited by three vertical septa. Pelage is brownish reddish, with whitish at base of each hair; fur is grayish in ventral part. Baculum is large and robust. Both males and females present frontal sexual sac.	Sorensen’s Leaf-nosed Bat has been reported in caves as well as over ricefields and other agricultural land. Altitudinal range is thought to be up to 1000 m .	Sorensen’s Leaf-nosed Bat probably forages over different types of plantation, feeding predominandy on insects.	Sorensen’s Leaf-nosed Bat forms breeding colonies.	This species is a cave-dweller.	Sorensen’s Leaf-nosed Bats seem to be gregarious, forming colonies of several individuals.	Classified as Vulnerable on The IUCN Red List because this species is only found in a small area. Sorensen’s Leaf-nosed Bat has been and is protected in its whole area of occurrence. Although it is abundant within its small range, the increasing disturbance of its roosts might be threatening its survival.	Kitchener & Maryanto (1993) | Maryanto et al. (2008) | Simmons (2005)	https://zenodo.org/record/3749105/files/figure.png	34. Sorensen’s Leaf-nosed Bat Hipposideros sorenseni French: Phyllorhine de Sorensen / German: Sorensen-Rundblattnase I Spanish: Hiposidérido de Sorensen Other common names: Sorensen's Roundleaf Bat Taxonomy. Hipposideros sorenseni Kitchener & Maryanto, 1993 , “ Gua Kramat (s holy cave), Pangandaran, W[est]. Java (c. 7° 41’S , 108° 40’E ).” Hipposideros sorenseni is in the larvatus species group. This species was initially allocated to H. larvatus . Monotypic. Distribution. Known only from the type locality in WGJava. Descriptive notes. Head-body 50-2-60 mm, tail 30-1-36-2 mm, ear 20-2-22-8 mm, forearm 55-4-60-2 mm. Greatest lengths of skulls are 21-1-22-1 mm, tibia 21-4-23-3 mm. Ears of Sorensen’s Leaf-nosed Bat are large and triangular, slightly concave before reaching tip. Anterior noseleaf has small depression in center and three supplementary lateral leaflets. Nasal septum is slighdy inflated. Upper border of posterior leaf is semicircular and there are three septa forming four noticeable cells on frontal surface, delimited by three vertical septa. Pelage is brownish reddish, with whitish at base of each hair; fur is grayish in ventral part. Baculum is large and robust. Both males and females present frontal sexual sac. Habitat. Sorensen’s Leaf-nosed Bat has been reported in caves as well as over ricefields and other agricultural land. Altitudinal range is thought to be up to 1000 m . Food and Feeding. Sorensen’s Leaf-nosed Bat probably forages over different types of plantation, feeding predominandy on insects. Breeding. Sorensen’s Leaf-nosed Bat forms breeding colonies. Activity patterns. This species is a cave-dweller. Movements, Home range and Social organization. Sorensen’s Leaf-nosed Bats seem to be gregarious, forming colonies of several individuals. Status and Conservation. Classified as Vulnerable on The IUCN Red List because this species is only found in a small area. Sorensen’s Leaf-nosed Bat has been and is protected in its whole area of occurrence. Although it is abundant within its small range, the increasing disturbance of its roosts might be threatening its survival. Bibliography. Kitchener & Maryanto (1993), Maryanto et al. (2008), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros sorenseni	Hipposideros		sorenseni	Kitchener & Maryanto	1993	0	Rec. West. Aust. Mus.	0.7653	Sorensen's Leaf-nosed Bat	None.	Indonesia, W Java, Pangandaran, Gua Karmat (= holy cave)(c. 7Â°41âS, 108Â°40âE).	C and W Java (Indonesia).	Not listed.	Endangered	 larvatus species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros sorenseni	23	Sorensen's Leaf-nosed Bat	Sorensen's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	sorenseni	Kitchener & Maryanto	1993	0	Hipposideros_sorenseni	Kitchener, D. H., & Maryanto, I. (1993). Taxonomic reappraisal of the Hipposideros larvatus species complex (Chiroptera Hipposideridae) in the Greater and Lesser Sunda Islands, Indonesia. Records of the Western Australian Museum, 16, 142.	https://www.biodiversitylibrary.org/item/239286#page/158/mode/1up	MZB 11333		"Gua Kramat (â‰¡ holy cave), Pangandaran, W[est]. Java (c. 7Â° 41'S, 108Â° 40'E)."	-7.683	108.67	sorenseni Kitchener & Maryanto, 1993	NA	NA	Indonesia	Asia	Indomalaya	EN	0	0	0	Hipposideros_sorenseni	0	sciname match	Hipposideros_sorenseni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10161	Hipposideros sorenseni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	sorenseni	Kitchener &; Maryanto, 1993		20000000	Hipposideros sorenseni	Endangered	B1ab(iii)	2021	2021-01-01 00:00:00 UTC	3.1	English	Hipposideros sorenseni is assessed as Endangered under criterion B1ab(iii) as <a style="mso-comment-reference:SW_1;mso-comment-date:20201225T1240">the species is endemic to the island of Java, Indonesia and the entire global population is found in one cave in the type-locality of </a>Gua Kramat in Pangandaran Regency, southern border of West Java and six caves from the Gombomg Karst Area Central Java province; combined they are considered to be one population occurring in fewer than five threat-based locations. Its e<a style="mso-comment-reference:DW_3;mso-comment-date:20201230T2024">xtent of occurrence (EOO) is 2,240 kmÂ²</a> and the karst area is used farming-irrigation, mining, forestry, tourism, and to supply water. The caves are subject to ongoing disturbance from mining, tourism and religious developments.	Hipposideros sorenseni roosts inside caves (Wijayanti et al. 2012, Wijayanti and Maryanto 2017) and can form relatively large colonies. Within a cave, their roost sites tend to be separate from other species (e.g., Rousettus amplexicaudatus, Eonycteris spelaea, Rhinolophus borneensis , and other hipposiderid bats (H. ater, H. diadema and H. bicolor )) that also roost in the cave (Wijayanti and Maryanto 2017). The cave roosts are located in landscapes dominated by agriculture (mixed gardens, rice paddies, and plantations), shrublands, development, and more distant forests (Wijayanti et al. 2012). The species has been documented to feed on at least 17 families in invertebrates and have a preference for Coleoptera, Orthoptera, and Neuroptera (Wijayanti et al. 2012).	The species is at risk from a diverse set of ongoing management and land use actions in the species limited range. There is extensive agriculture and water use in the region, loss of remnant forests, and mining within the species range (Ansori and Puswanto 2011, Wijayanti et al. 2012, Wijayanti and Maryanto 2017), and given the current understanding of the species limited use of caves (one cave as type locality and six other caves as confirmed roosts; Wijayanti et al. 2012, Wijayanti and Maryanto 2017), cave tourism and cave development (Ansori 2011) are major threats that could result in significant harm to the species if a catastrophic even happens at a known roost.	The current status and trends of the speciesâ€™ global population is unknown as there is no active monitoring. The species can congregate in relatively large colonies and can be relatively common locally (I. Maryanto pers. comm.). There are at least 112 caves in the Gombong Karst Area and the species was documented in six of the 12 caves surveyed (Wijayanti et al. 2012).	Unknown	This species is endemic to Java Island, Indonesia and the entire global population is known from the type locality in Gua Kramat, Pangandaran, West Java and from at least six caves in Gombong Karst Area, Central Java, Indonesia (Kitchener and Maryanto 1993, Wijayanti et al. 2012, Wijayanti and Maryanto 2017). <a style="">Its extent of occurrence (EOO) is ca 2,240 kmÂ². ;</a>Despite targeted surveys of the surrounding caves, the species has not been found elsewhere on Java (I. Maryanto pers. comm.). Its elevational range is below 1,000 m asl.	The species is not known to be used.	Terrestrial	This species has been recorded from Pangandaran Nature Reserve, but there are no species-specific conservation actions in place for the species. A comprehensive species conservation action plan is needed. Proactive campaigns to protect and restore the known cave roosts, including community awareness programs, are warranted to secure the species future. Additional research is needed to determine the species broader distribution in the region and alternative roosts. Information on the species population status and trend, distribution, ecology, life history, and threats is needed to inform the species conservation.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		sorenseni	Kitchener & Maryanto	1993	0	Rec. West. Aust. Mus.	0.765278	Sorensen's Leaf-nosed Bat	None.	Indonesia, W Java, Pangandaran, Gua Karmat (= holy cave)(c. 7Â°41â€™S, 108Â°40â€™E).	C and W Java (Indonesia).	Not listed.	Endangered	 larvatus species group.	Hipposideros sorenseni	1004634	23	Sorensen's Leaf-nosed Bat	Sorensen's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	sorenseni	Kitchener & Maryanto	1993	0	Hipposideros_sorenseni	Kitchener, D. H., & Maryanto, I. (1993). Taxonomic reappraisal of the Hipposideros larvatus species complex (Chiroptera Hipposideridae) in the Greater and Lesser Sunda Islands, Indonesia. Records of the Western Australian Museum, 16, 142.	https://www.biodiversitylibrary.org/item/239286#page/158/mode/1up	MZB 11333		"Gua Kramat (â‰¡ holy cave), Pangandaran, W[est]. Java (c. 7Â° 41'S, 108Â° 40'E)."	-7.68333	108.6667	sorenseni Kitchener & Maryanto, 1993	NA	NA				Indonesia	Asia	Indomalaya	EN	0	0	0	Hipposideros_sorenseni	0	sciname match	Hipposideros_sorenseni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_sorenseni	1004634	23	Sorensen's Roundleaf Bat	Sorensen's Roundleaf Bat|Sorensen's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	sorenseni	D. J. Kitchener & Maryanto	0	Hipposideros sorenseni	Kitchener, D.J. and Maryanto, I. 1993. Taxonomic reappraisal of the _Hipposideros larvatus_ species complex (Chiroptera, Hipposideridae) in the Greater and Lesser Sunda Islands, Indonesia. Records of the Western Australian Museum 16(2):119-173.	https://www.biodiversitylibrary.org/page/54768571	MZB 11333	holotype		"Gua Kramat (â‰¡ holy cave), Pangandaran, W[est]. Java (c. 7Â° 41'S, 108Â° 40'E)."	-7.68333	108.6667	NA	NA				Indonesia	Asia	Indomalaya	EN	0	0	0	Hipposideros_sorenseni	0	sciname match	Hipposideros_sorenseni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		sorenseni	Kitchener & Maryanto	1993	0	Rec. West. Aust. Mus.	0.765278	Sorensen's Leaf-nosed Bat	None.	Indonesia, W Java, Pangandaran, Gua Karmat (= holy cave)(c. 7Â°41â€™S, 108Â°40â€™E).	C and W Java (Indonesia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10161/22099115/' target='_blank'>Endangered</a>	larvatusspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros sorenseni; Hipposideros sorenseni; Hipposideros sorenseni; Hipposideros sorenseni; Hipposideros sorenseni; Hipposideros sorenseni; sorenseni; Phyllorhine de Sorensen; Sorensen-Rundblattnase; Hiposidérido de Sorensen; Sorensen's Roundleaf Bat; Sorensen's Leaf-nosed Bat; Sorensen's Roundleaf Bat; Sorensen's Leaf-nosed Bat; Sorensen's Leaf-nosed Bat; H. sorenseni
