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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L437	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Doryrhina semoni	Hipposideros semoni	Doryrhina semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni	Hipposideros semoni		[MSW3] cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [HMW] Hipposideros semoni Matschie, 1903 , Cooktown, Queensland , Australia . This species is monotypic.; [batnames2022] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2022] moved from Hipposideros to Doryrhina; [IUCN] The type locality of Hipposideros semoni is near Cooktown in Queensland, Australia (Mahoney and Walton 1988). Both the identity and taxonomic affinity of the handful of specimens attributed to H. semoni from across Papua New Guinea are currently being investigated, but so far specimens either side of the Torres Strait appear to be the same species (K.N. Armstrong unpublished data).; [batnames2023] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2023] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [MDD2025_2.0] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being; [batnames2025_1.7] Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. See Flannery (1995a) and Bonaccorso (1998).; [MDD2025_2.2] has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being														semoni	The type locality of Hipposideros semoni is near Cooktown in Queensland, Australia (Mahoney and Walton 1988). Both the identity and taxonomic affinity of the handful of specimens attributed to H. semoni from across Papua New Guinea are currently being investigated, but so far specimens either side of the Torres Strait appear to be the same species (K.N. Armstrong unpublished data).			semoni 	semoni 			semoni Matschie, 1903		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Semon's leaf-nosed bat	E New Guinea, N Queensland	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros semoni	Australia, Queensland, Cooktown.	Matschie	1903	Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.), 8:774 (Heft 6:132).	Distribution: Known only from eastern New Guinea and northeastern Queens land (Australia).		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Semon's leaf-nosed bat	E New Guinea, NE Queensland	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Matschie	1903	Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.), 8:774 (Heft 6:132).		N Queensland (Australia), E New Guinea.	Australia, Queensland, Cooktown.		MATSCHIE	1903	Posterior lateral leaflet extending anteriorly beneath the anterior noseleaf. Premaxillae narrow, not enclosing the anterior palatal foramina, mak ing a narrow spatulate junction with the maxillae. Median process of intermediate noseleaf well de veloped and club-shaped. Skull with a shallow frontal depression. Sphenoidal depression absent. Posterior ridge of last upper molar little reduced. Width of each cochlea at least six times their dis tance apart. Interorbital region relatively wide. Median process of posterior noseleaf well-devel oped. Rostrum relatively short, its eminences greatly inflated, Antorbital foramen relatively small. Size relatively small (forearm length, 42-49 mm).	Distribution: Known only from eastern New Guinea and northeastern Queens land (Australia).	No subspecies.		64	species	H. semoni	MATSCHIE	1903	Hipposideros	genus	Hipposideros semoni				Posterior lateral leaflet extending anteriorly beneath the anterior noseleaf. Premaxillae narrow, not enclosing the anterior palatal foramina, mak ing a narrow spatulate junction with the maxillae. Median process of intermediate noseleaf well de veloped and club-shaped. Skull with a shallow frontal depression. Sphenoidal depression absent. Posterior ridge of last upper molar little reduced. Width of each cochlea at least six times their dis tance apart. Interorbital region relatively wide. Median process of posterior noseleaf well-devel oped. Rostrum relatively short, its eminences greatly inflated, Antorbital foramen relatively small. Size relatively small (forearm length, 42-49 mm).	No subspecies.		38. H. semoni MATSCHIE 1903 [cyclops group].	38	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros semoni	Hipposideros		semoni	Matschie		1903		Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.)	8		774 (Heft 6:132)		Semon's Leaf-nosed Bat	Australia, Queensland, Cooktown.	N Queensland (Australia), E New Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	03BD87A2C672A200F89DFECCF4D84ED7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	231	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C672A200F89DFECCF4D84ED7.xml	Doryrhina semoni	Hipposideridae	Doryrhina	semoni		1903	Phyllorhine de Semon @fr | Semon-Rundblattnase @de | Doryrina de Semon @es | Semon's Roundleaf Bat @en	Hipposideros semoni Matschie, 1903 , Cooktown, Queensland , Australia . This species is monotypic.	E New Guinea and NE Australia , where it has been reported in several locations of coastal Queensland from the E side of Cape York Peninsula to S of Cooktown; S limit of its distribution is not clear, as there are acoustic reports from other distant areas, such as in the Kroombit Tops National Park.	Head-body 40-50 mm, tail 22-28 mm, ear 18-20 mm, hindfoot 7-12 mm, forearm 38-424 mm; weight 6—10 g . As in the case of Wollaston’s Leafnosed Bat (Z). wollastoni ) and the Fly River Leaf-nosed Bat ( D. muscinus ), Semon’s Leafnosed Bat presents two supplementary leaflets: anterior leaflet is short and broad and posterior leaflet extends beneath anterior leaf onto upper lip. It also presents processes in intermediate and posterior leaves. Median process of posterior leaf is large and club-shaped. Its ears are notably long and well developed with pointed tips, and its rostrum is markedly inflated. Its dark grayish pelage is long and presents a curled appearance, being paler on ventral area.	Semon’s Leaf-nosed Bat is found in a wide variety of habitats, from rainforest to savanna forest and dry open areas, at elevations from the sea level up to 1400 m .	Semon’s Leaf-nosed Bat usually feeds on insects, especially moths; it also takes beetles, as well as spiders. It hunts prey from the ground, low vegetation or surfaces such as rocks or tree trunks. It has a slow flight, with considerable ability to maneuver, and usually flies within 1—2 m of the ground, foraging within dense vegetation and scrub.	Females are probably mature in their first year, and give birth probably in early November. Each female produces a single young, but it is thought that in a few cases there may be twins. Generation length is estimated at 6-7 years.	Semon’s Leaf-nosed Bat has been observed roosting in many different places such as in caves, mines, abandoned buildings, road culverts, fissures, and tree hollows. Some authors have suggested that this species is more tree-dweller than cave-dweller, inhabiting caves probably during the drier periods. Males and females have been suggested to have different echolocation call frequencies, being c.75 kHz and 95 kHz, respectively.	Semon’s Leaf-nosed Bats have been found roosting alone or in small aggregations. Movement patterns between roosts or habitats have not yet been studied. It may use vegetation strips in rainforests, following rivers and watercourses to move within large dense forest areas. Migration has not been reported.	Classified as Least Concern on The IUCNRed List (as Hipposideros semoni ). Current population size and trends are unknown, but it is probably not greatly fragmented. Habitat loss and fragmentation might be threats for this species, although no major risks have been yet identified. Predation by feral cats has been reported and inappropriate fire management could also become a threat for this species in Australia . In fact, a Recovery Plan has been established for Semon’s Leaf-nosed Bats, as it is considered a threatened species under Australian legislation. In Papua New Guinea , it is probably widely distributed, as much intact habitat remains. However, it has been suggested that it may occur at low densities, as the species has not been captured or recorded in recent surveys. In addition, its echolocation call might be confused with those of other bat species, which could easily give an erroneous idea of its true range. More research is needed to study and assess its ecology and population status.	Armstrong & Aplin (2017f) | Bonaccorso (1998) | Churchill (1998) | Coles (1993) | Flannery & Colgan (1993) | Hall (2008c) | Oliveira & Schulz (1997) | Vestjens & Hall (1977)	https://zenodo.org/record/3749057/files/figure.png	10. Semon’s Leaf-nosed Bat Doryrhina semoni French: Phyllorhine de Semon / German: Semon-Rundblattnase / Spanish: Doryrina de Semon Other common names: Semon's Roundleaf Bat Taxonomy. Hipposideros semoni Matschie, 1903 , Cooktown, Queensland , Australia . This species is monotypic. Distribution. E New Guinea and NE Australia , where it has been reported in several locations of coastal Queensland from the E side of Cape York Peninsula to S of Cooktown; S limit of its distribution is not clear, as there are acoustic reports from other distant areas, such as in the Kroombit Tops National Park. Descriptive notes. Head-body 40-50 mm, tail 22-28 mm, ear 18-20 mm, hindfoot 7-12 mm, forearm 38-424 mm; weight 6—10 g . As in the case of Wollaston’s Leafnosed Bat (Z). wollastoni ) and the Fly River Leaf-nosed Bat ( D. muscinus ), Semon’s Leafnosed Bat presents two supplementary leaflets: anterior leaflet is short and broad and posterior leaflet extends beneath anterior leaf onto upper lip. It also presents processes in intermediate and posterior leaves. Median process of posterior leaf is large and club-shaped. Its ears are notably long and well developed with pointed tips, and its rostrum is markedly inflated. Its dark grayish pelage is long and presents a curled appearance, being paler on ventral area. Habitat. Semon’s Leaf-nosed Bat is found in a wide variety of habitats, from rainforest to savanna forest and dry open areas, at elevations from the sea level up to 1400 m . Food and Feeding. Semon’s Leaf-nosed Bat usually feeds on insects, especially moths; it also takes beetles, as well as spiders. It hunts prey from the ground, low vegetation or surfaces such as rocks or tree trunks. It has a slow flight, with considerable ability to maneuver, and usually flies within 1—2 m of the ground, foraging within dense vegetation and scrub. Breeding. Females are probably mature in their first year, and give birth probably in early November. Each female produces a single young, but it is thought that in a few cases there may be twins. Generation length is estimated at 6-7 years. Activity patterns. Semon’s Leaf-nosed Bat has been observed roosting in many different places such as in caves, mines, abandoned buildings, road culverts, fissures, and tree hollows. Some authors have suggested that this species is more tree-dweller than cave-dweller, inhabiting caves probably during the drier periods. Males and females have been suggested to have different echolocation call frequencies, being c.75 kHz and 95 kHz, respectively. Movements, Home range and Social organization. Semon’s Leaf-nosed Bats have been found roosting alone or in small aggregations. Movement patterns between roosts or habitats have not yet been studied. It may use vegetation strips in rainforests, following rivers and watercourses to move within large dense forest areas. Migration has not been reported. Status and Conservation. Classified as Least Concern on The IUCNRed List (as Hipposideros semoni ). Current population size and trends are unknown, but it is probably not greatly fragmented. Habitat loss and fragmentation might be threats for this species, although no major risks have been yet identified. Predation by feral cats has been reported and inappropriate fire management could also become a threat for this species in Australia . In fact, a Recovery Plan has been established for Semon’s Leaf-nosed Bats, as it is considered a threatened species under Australian legislation. In Papua New Guinea , it is probably widely distributed, as much intact habitat remains. However, it has been suggested that it may occur at low densities, as the species has not been captured or recorded in recent surveys. In addition, its echolocation call might be confused with those of other bat species, which could easily give an erroneous idea of its true range. More research is needed to study and assess its ecology and population status. Bibliography. Armstrong & Aplin (2017f), Bonaccorso (1998), Churchill (1998), Coles (1993), Flannery & Colgan (1993), Hall (2008c), Oliveira & Schulz (1997), Vestjens & Hall (1977).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros semoni	Hipposideros		semoni	Matschie	1903	0	Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.)	8: 774 (Heft 6:132)	Semon's Leaf-nosed Bat	None.	Australia, Queensland, Cooktown.	N Queensland (Australia), E New Guinea.	Not listed.	Least Concern	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Doryrhina semoni	23	Semon's Leaf-nosed Bat	Semon's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Doryrhina	NA	semoni	Matschie	1903	1	Hipposideros_semoni	Matschie, P. (1903). Die Chiropteren, Insectivoren und Muriden der SemonÂ´schen Forschungsreise. Zoologische Forschungsreisen in Australien und dem Malayischen Archipel. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft Jena, 8, 774.	https://www.biodiversitylibrary.org/item/121460#page/832/mode/1up	ZMB 22225		Cooktown, Queensland, Australia.			semoni (Matschie, 1903)	moved from Hipposideros to Doryrhina	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.	Papua New Guinea|Australia	Oceania	Australasia/Oceania	LC	0	0	0	Doryrhina_semoni	0	sciname match	Hipposideros_semoni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10160	Hipposideros semoni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	semoni	Matschie, 1903	The type locality of Hipposideros semoni is near Cooktown in Queensland, Australia (Mahoney and Walton 1988). Both the identity and taxonomic affinity of the handful of specimens attributed to H. semoni from across Papua New Guinea are currently being investigated, but so far specimens either side of the Torres Strait appear to be the same species (K.N. Armstrong unpublished data).	200000000	Hipposideros semoni	Least Concern		2021	2016-02-14 00:00:00 UTC	3.1	English	<p>Hipposideros semoni is recorded from both Papua New Guinea and northern Queensland, Australia. It has been listed as Endangered since 2001 under national environmental legislation in Australia because of its apparent rarity and past reductions in habitat. More recent views of the Australian population have matched it against Vulnerable (B1ab(iii); Reardon et al. 2010) and Near Threatened (Woinarski et al. 2014). Lack of information was an obstacle to making a clear decision in the latter assessmentâ€”area of occupancy is not well known but is possibly ca 2,000 kmÂ², the number of known locations is thought to be &lt; 10, total Australian population size was judged to be &gt; 10 000 mature individuals, though with the largest subpopulation probably &lt; 1,000 individualsâ€”and population size was not thought to be declining at a rate &gt;30% over a 20 year (3 generations) period. At a global scale, its presence in Papua New Guinea means that it does not currently meet the criteria for listing as threatened. A suggestion that specimens collected from Papua New Guinea are instead attributable to the morphologically-similar H. wollastoni is true for some records listed on GBIF (South Australian Museum accessions M10483â€“10488 are actually H. corynophyllus and H. wollastoni ; T.B. Reardon and K.N. Armstrong unpublished obs.), but not all (e.g. Smithsonian Institution USNM553716 has diagnostic external morphology of H. semoni ; K.N. Armstrong unpublished obs.). There is also a recent possible acoustic record from Gulf Province (K.N. Armstrong and K.P. Aplin unpublished data). Thus, while rare and seldom encountered, there is sufficient evidence that H. semoni does occur over a wide area in Papua New Guinea. The species is therefore listed as Least Concern (LC), but it needs to be reassessed following the completion of taxonomic studies. ;</p>	<p>This species is found in rainforest, savanna forest, and dry open areas. It roosts in caves and overhangs, mines, old buildings, culverts under roads, and tree hollows (Bonaccorso 1998; Hall 2008). In Australia, it apparently uses the thin strips of rainforest vegetation in riparian zones of major rivers to move out from larger patches of occupied rainforest habitat (K.N. Armstrong unpublished obs.). It is not known to form larger roosting aggregations like other hipposiderids species, instead roosting singly or in small groups of up to ten individuals (Hall 2008). Females give birth to single young around November (Churchill 2008). Females are larger and produce echolocation calls with a lower frequency compared to males. Generation length is around 6â€“7 years (Woinarski et al. 2014).</p>	<p>Major threats to this species have not been observed to operate but several are considered plausible (Woinarski et al . 2014). Disturbance and loss of roost sites is thought to be a threat but of relatively low risk, with broader scale effects from inappropriate burning regimes and fragmentation and modification of habitat from pastoralism and other land uses more likely to have an effect on the area of occupancy and range of Hipposideros semoni in Australia. Predation by feral cats is also regarded as a threat, albeit a minor one (Woinarski et al . 2014). In Queensland, there was a gradual reduction in vegetation clearing that peaked in 2009, but changes in policy and legislation since that time have once more led to an increase (Maron et al .2015). Such a trend reversal, coupled with inappropriate fire management of native bushland has the potential to result in further losses of habitat for this species. There are no recognised threats in Papua New Guinea.</p>	<p>H. semoni is generally considered to be uncommon and occurring at low density because it is encountered rarely, but it can be captured or recorded if targeted in rainforest habitats in Australia (Hall 2008; Reardon et al. 2010; K.N. Armstrong unpublished data). There have been no attempts at census in Australia, and estimates need to consider that animals have been observed seasonally in drier, open woodlands on Cape York Peninsula as well as rainforest patches (Hall 2008). There is opinion that numbers are greater than previous estimates of &lt;2,500 on Cape York Peninsula, and that it is relatively secure in this region (Reardon et al. 2010). For populations further south, there are contrasting views on whether a decline has occurred in the past, and factors that may have caused such a decline have not been demonstrated, but habitat modelling suggests that a decline in the south is likely over the next 50 years (Dennis 2012; Woinarski 2014). </p> <p>There is no information on population size or connectedness in Papua New Guinea, though numerous surveys in the past five years across several provinces have only produced a single putative acoustic record (Armstrong and Aplin 2011, 2014; Armstrong et al. 2015; K.N. Armstrong and K.P. Aplin unpublished data). </p>	Unknown	<p>The presence of H. semoni in Papua New Guinea is represented by museum specimens from near Vanimo in Sandaun (West Sepik) Province, localities in Morobe Province, and several localities around Port Moresby. Given the intactness of much of the habitat in Papua New Guinea, it might have a broad distribution, though occur at low density given that numerous surveys in recent years in several provinces have not produced captures. Acoustic detection in Papua New Guinea, the most efficient way to assess area of occupancy, is hindered by a lack of information on the similarity of its call to other closely related New Guinea Hipposideros (K.N. Armstrong and K.P. Aplin unpublished data). In Australia, the majority of recent records are from the eastern side of Cape York Peninsula, Queenslandâ€”Iron Range, Kulla, Oyala Thumotang and Cape Melville National Parks, near Cooktown and south to Townsville (Reardon et al. 2010; Woinarski et al. 2014). There is also a single report of a disjunct occurrence ca 700 km further south in Kroombit Tops National Park (south of Gladstone, Queensland; Schulz and de Oliveira 1995), and an unconfirmed and likely erroneous acoustic record as far south as Maryborough (de Oliveira and Pavey 1995; probably from Rhinolophus megaphyllus ). It ranges from sea level such as at Cape Melville National Park, Queensland to over 1,400 m Asl in Morobe Province, Papua New Guinea.</p>		Terrestrial	<p>A Recovery Plan has been developed for Hipposideros semoni in Australia (Thomson et al. 2001). It is found in several protected areas in Queensland, and is listed under national environmental legislation in Australia as a threatened species. Taxonomic studies need to be completed, and further studies are needed into the distribution (particularly within New Guinea), abundance, natural history, and threats to this species.</span></p>	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		semoni	Matschie	1903	0	Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.)	8: 774 (Heft 6:132)	Semon's Leaf-nosed Bat	None.	Australia, Queensland, Cooktown.	N Queensland (Australia), E New Guinea.	Not listed.	Least Concern	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclops species group. See Flannery (1995a) and Bonaccorso (1998).	Hipposideros semoni	1004571	23	Semon's Leaf-nosed Bat	Semon's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	semoni	Matschie	1903	0	Hipposideros_semoni	Matschie, P. (1903). Die Chiropteren, Insectivoren und Muriden der SemonÂ´schen Forschungsreise. Zoologische Forschungsreisen in Australien und dem Malayischen Archipel. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft Jena, 8, 774.	https://www.biodiversitylibrary.org/item/121460#page/832/mode/1up	ZMB 22225		Cooktown, Queensland, Australia.			semoni (Matschie, 1903)	has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Papua New Guinea|Australia	Oceania	Australasia/Oceania	LC	0	0	0	Doryrhina_semoni	0	sciname match	Hipposideros_semoni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_semoni	1004571	23	Semon's Roundleaf Bat	Semon's Roundleaf Bat|Semon's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	semoni	Matschie	0	Hipposideros semoni	Matschie, P. 1903. Die Chiropteren, Insectivoren und Muriden der Semon'schen Porschungsreise. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft Jena 8:771-778.	https://www.biodiversitylibrary.org/page/39292575	ZMB 22225	holotype		Cooktown, Queensland, Australia.			has been included under Doryrhina based on suspected close relationship to Doryrhina cyclops, but this relationship has not been examined using molecular data at this point and it is retained under Hipposideros here for the time being	Foley, N. M., Goodman, S. M., Whelan, C. V., Puechmaille, S. J., & Teeling, E. (2017). Towards navigating the Minotaur's labyrinth: cryptic diversity and taxonomic revision within the speciose genus Hipposideros (Hipposideridae). Acta Chiropterologica, 19(1), 1-18.|Wilson D.E. & Mittermeier R.A. 2019. Handbook of the mammals of the world. Vol. 9. Bats. Lynx Edicions, Barcelona.				Papua New Guinea|Australia	Oceania (Continent)	Australasia	LC	0	0	0	Doryrhina_semoni	0	sciname match	Hipposideros_semoni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		semoni	Matschie	1903	0	Denks. Med. Nat. Ges. Jena (Semon Zool. Forsch. Austr.)	8: 774 (Heft 6:132)	Semon's Leaf-nosed Bat	None.	Australia, Queensland, Cooktown.	N Queensland (Australia), E New Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10160/209537564/' target='_blank'>Least Concern</a>	Transferred to Doryrhina by Tuneu-Corral (2019) and the Mammal Diversity Database (2021). See our comments in the generic account. cyclopsspecies group. See Flannery (1995a) and Bonaccorso (1998).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros semoni; Doryrhina semoni; Hipposideros semoni; Doryrhina semoni; Hipposideros semoni; Hipposideros semoni; semoni; Phyllorhine de Semon; Semon-Rundblattnase; Doryrina de Semon; Semon's Roundleaf Bat; Semon's Leaf-nosed Bat; Semon's Roundleaf Bat; Semon's Leaf-nosed Bat; Semon's Leaf-nosed Bat; H. semoni
