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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L436 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A N/A N/A Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares Hipposideros scutinares [MSW3] pratti species group.; [HMW] Hipposideros scutinares M. F. Robinson et al., 2003 , “Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province , Khammouan Limestone NBCA, Lao PDR, 17°58’N , 104°49’E .” Hipposideros scutinares is in the pratti species group. Monotypic.; [batnames2022] swinhoei species group.; [batnames2023] swinhoei species group.; [batnames2025_1.7] swinhoii species group. scutinares scutinares scutinares scutinares M. F. Robinson, P. D. Jenkins, C. M. Francis, & Fulford, 2003 N/A NA Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Hipposideridae Hipposideros scutinares Hipposideros scutinares Robinson, Jenkins, Francis, and Fulford 2003 Acta. Chiropt. 5 33 Shield-nosed Leaf-nosed Bat Laos, Khammouan Limestone NBCA, Bolikhamsai Province, along the upper Nam Hinboun, Ban Khankeo, 17˚58’N, 104˚49’E. Laos, Vietnam. IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001). pratti species group. 03BD87A2C67BA20AF882F09AFA924472 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Hipposideridae_210.pdf.imf hash://md5/ff84ffdac676a204fff8ff9affef4346 240 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67BA20AF882F09AFA924472.xml Hipposideros scutinares Hipposideridae Hipposideros scutinares 2003 Phyllorhine à blason @fr | Nasenschild-Rundblattnase @de | Hiposidérido de escudo @es | Shield-nosed Roundleaf Bat @en Hipposideros scutinares M. F. Robinson et al., 2003 , “Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province , Khammouan Limestone NBCA, Lao PDR, 17°58’N , 104°49’E .” Hipposideros scutinares is in the pratti species group. Monotypic. Endemic to C Laos and adjacent C Vietnam . Tail 50-4—54-3 mm, ear 27*6-29* 4 mm , hindfoot 15—18* 8 mm , forearm 77*9-82* 7 mm ; weight 42 g . Ears of the Shield-nosed Leaf-nosed Bat are well developed, haired in one-third of their length and with a slightly concave shape below tip. On dorsum, fur is brown or orange-reddish at base, followed by a brown-yellowish color in central portion of each hair, with brown tips; in ventral part, fur is brown-reddish at base with brown-yellow tips. In middle of anterior noseleaf there is a small emargination. There are two supplementary lateral leaflets. Lateral borders of anterior and posterior leaves are joined. To rear of posterior noseleaf there is a protuberance on each side of frontal sac; shape of this protuberance reminds supplementary lateral leaflet and is more noticeable in males. Rostrum eminences are slighdy inflated. The Shield-nosed Leaf-nosed Bat has been observed roosting in caves in limestone areas, karst, and cliffs. It has also been reported in evergreen forest, and deciduous forest with bushes and bamboo. It occurs in primary forests as well as in more degraded habitats including agricultural land, such as ricefields, reaching altitudes of 160 m . The Shield-nosed Leaf-nosed Bat is thought to forage in primary forests, as well as agricultural lands and other disturbed habitats. Its diet is probably based on insects. Breeding colonies are located only in caves. No other information is known as yet about the breeding biology of the Shield-nosed Leaf-nosed Bat. The Shield-nosed Leaf-nosed Bat is a cave-dwelling species. Echolocation call frequency of this species is c.63*6 kHz. No information. Classified as Vulnerable on The IUCNRed List due to its small geographic range. Reported only in eight areas, six in Laos and two from adjacent Vietnam . The population of the Shield-nosed Leaf-nosed Bat is probably decreasing, its population size being estimated at 10,000 individuals or less. Habitat loss could be the most important threat to this species. Francis & Bates (2008b) | Robinson et al. (2003) | Thomas et al. (2013) https://zenodo.org/record/3749218/files/figure.png 38. Shield-nosed Leaf-nosed Bat Hipposideros scutinares French: Phyllorhine à blason I German: Nasenschild-Rundblattnase / Spanish: Hiposidérido de escudo Other common names: Shield-nosed Roundleaf Bat Taxonomy. Hipposideros scutinares M. F. Robinson et al., 2003 , “Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province , Khammouan Limestone NBCA, Lao PDR, 17°58’N , 104°49’E .” Hipposideros scutinares is in the pratti species group. Monotypic. Distribution. Endemic to C Laos and adjacent C Vietnam . Descriptive notes. Tail 50-4—54-3 mm, ear 27*6-29* 4 mm , hindfoot 15—18* 8 mm , forearm 77*9-82* 7 mm ; weight 42 g . Ears of the Shield-nosed Leaf-nosed Bat are well developed, haired in one-third of their length and with a slightly concave shape below tip. On dorsum, fur is brown or orange-reddish at base, followed by a brown-yellowish color in central portion of each hair, with brown tips; in ventral part, fur is brown-reddish at base with brown-yellow tips. In middle of anterior noseleaf there is a small emargination. There are two supplementary lateral leaflets. Lateral borders of anterior and posterior leaves are joined. To rear of posterior noseleaf there is a protuberance on each side of frontal sac; shape of this protuberance reminds supplementary lateral leaflet and is more noticeable in males. Rostrum eminences are slighdy inflated. Habitat. The Shield-nosed Leaf-nosed Bat has been observed roosting in caves in limestone areas, karst, and cliffs. It has also been reported in evergreen forest, and deciduous forest with bushes and bamboo. It occurs in primary forests as well as in more degraded habitats including agricultural land, such as ricefields, reaching altitudes of 160 m . Food and Feeding. The Shield-nosed Leaf-nosed Bat is thought to forage in primary forests, as well as agricultural lands and other disturbed habitats. Its diet is probably based on insects. Breeding. Breeding colonies are located only in caves. No other information is known as yet about the breeding biology of the Shield-nosed Leaf-nosed Bat. Activity patterns. The Shield-nosed Leaf-nosed Bat is a cave-dwelling species. Echolocation call frequency of this species is c.63*6 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCNRed List due to its small geographic range. Reported only in eight areas, six in Laos and two from adjacent Vietnam . The population of the Shield-nosed Leaf-nosed Bat is probably decreasing, its population size being estimated at 10,000 individuals or less. Habitat loss could be the most important threat to this species. Bibliography. Francis & Bates (2008b), Robinson et al. (2003), Thomas et al. (2013). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Hipposideridae Hipposideros scutinares Hipposideros scutinares Robinson, Jenkins, Francis & Fulford 2003 0 Acta Chiropterol. 5:33 Shield-nosed Leaf-nosed Bat None. Laos, Khammouan Limestone NBCA, Bolikhamsai Province, along the upper Nam Hinboun, Ban Khankeo, 17Ë58âN, 104Ë49âE. Laos, Vietnam. Not listed. Vulnerable swinhoei species group. Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Hipposideros scutinares 23 Shield-nosed Leaf-nosed Bat Shield-nosed Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA HIPPOSIDERIDAE NA NA Hipposideros NA scutinares M. F. Robinson, Jenkins, Francis, & Fulford 2003 0 Hipposideros_scutinares Robinson, M. F., Jenkins, P. D., Francis, C. M., & Fulford, A. J. C. (2003). A new species of the Hipposideros pratti group (Chiroptera, Hipposideridae) from Lao PDR and Vietnam. Acta Chiropterologica, 5(1), 33. https://bioone.org/journals/acta-chiropterologica/volume-5/issue-1/001.005.0103/A-New-Species-of-the-Hipposideros-pratti-Group-Chiroptera-Hipposideridae/10.3161/001.005.0103.full SMF 88.023 "Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province, Khammouan Limestone NBCA, Lao PDR, 17°58'N, 104°49'E." 17.967 104.82 scutinares M. F. Robinson, Jenkins, Francis, & Fulford, 2003 NA NA Laos|Vietnam Asia Indomalaya VU 0 0 0 Hipposideros_scutinares 0 sciname match Hipposideros_scutinares 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 136586 Hipposideros scutinares ANIMALIA CHORDATA MAMMALIA CHIROPTERA HIPPOSIDERIDAE Hipposideros scutinares Robinson, Jenkins, Francis &; Fulford, 2003 20000000 Hipposideros scutinares Vulnerable C1 2019 2018-08-31 00:00:00 UTC 3.1 English The species is confirmed as Vulnerable as it is only known from a few locations, based on current surveys the population is estimated to be between 8,000 and 10,000 individuals (future surveys may find more individuals), and the population is projected to decline due to levels of exploitation (hunting) at a rate of 10% over the next 15 years (three generations; Pacifici et al. 2013). It was found roosting in caves in limestone areas (Robinson et al. 2003). A pregnant female was found foraging at a disturbed forest patch in buffer zone of a protected area in central Vietnam in mid-March 2018 (Furey and Tu unpublished data). This species has been threatened by cave disturbance and deforestation in the region in recent times (Thomas et al. 2013, V.T. Tu pers. comm). In Viet Nam, illegal hunting for local consumption is also identified as an additional threat to this species (V.T. Tu pers. comm). Surveys around the area, though not specifically for this species, have not captured any individuals (P. Bates pers. comm.). It is likely to have a population size between 8,000 to 10,000 individuals (C. Francis pers. comm.) and is projected to be declining from cave disturbance and deforestation in the region and illegal hunting for local consumption in Vietnam. Decreasing This species is currently known from a few localities associated with limestone karst areas in central Lao PDR and central Viet Nam (Robinson et al. 2003, Kruskop 2013, Furey and Tu unpublished data). The record from Tham Pasat-thia (Louang-namtha Province) listed by Thomas et al. (2013) has been rejected as upon re-examination these specimens are turned to be represent H. lylei . The Tham Nam (Houaphan Province) data listed by Thomas et al. (2013) is regarded ambiguous on zoogeographical grounds. Terrestrial This species occurs in protected areas in Viet Nam. Further studies are required to determine the population status of this species (C. Francis pers. comm.). Indomalayan FALSE FALSE Global Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Hipposideridae Hipposideros scutinares Robinson, Jenkins, Francis & Fulford 2003 0 Acta Chiropterol. 5:33 Shield-nosed Leaf-nosed Bat None. Laos, Khammouan Limestone NBCA, Bolikhamsai Province, along the upper Nam Hinboun, Ban Khankeo, 17Ëš58’N, 104Ëš49’E. Laos, Vietnam. Not listed. Vulnerable swinhoei species group. Hipposideros scutinares 1004633 23 Shield-nosed Leaf-nosed Bat Shield-nosed Roundleaf Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA PTEROPODIFORMES NA NA RHINOLOPHOIDEA Hipposideridae NA NA Hipposideros NA scutinares M. F. Robinson, Jenkins, Francis, & Fulford 2003 0 Hipposideros_scutinares Robinson, M. F., Jenkins, P. D., Francis, C. M., & Fulford, A. J. C. (2003). A new species of the Hipposideros pratti group (Chiroptera, Hipposideridae) from Lao PDR and Vietnam. Acta Chiropterologica, 5(1), 33. https://bioone.org/journals/acta-chiropterologica/volume-5/issue-1/001.005.0103/A-New-Species-of-the-Hipposideros-pratti-Group-Chiroptera-Hipposideridae/10.3161/001.005.0103.full SMF 88.023 "Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province, Khammouan Limestone NBCA, Lao PDR, 17°58'N, 104°49'E." 17.96667 104.8167 scutinares M. F. Robinson, Jenkins, Francis, & Fulford, 2003 NA NA Laos|Vietnam Asia Indomalaya VU 0 0 0 Hipposideros_scutinares 0 sciname match Hipposideros_scutinares 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Hipposideros_scutinares 1004633 23 Shield-nosed Roundleaf Bat Shield-nosed Roundleaf Bat|Shield-nosed Leaf-nosed Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yinpterochiroptera NA NA Rhinolophoidea Hipposideridae NA NA Hipposideros NA scutinares M. F. Robinson, P. D. Jenkins, C. M. Francis, & Fulford 0 Hipposideros scutinares Robinson, M.F., Jenkins, P.D., Francis, C.M. and Fulford, A.J.C. 2003-06. A new species of the _Hipposideros pratti_ group (Chiroptera, Hipposideridae) from Lao PDR and Vietnam. Acta Chiropterologica 5(1):31-48. https://doi.org/10.3161/001.005.0103 SMF:MAMM:88023 holotype "Ban Khankeo (formerly known as Ban Phahôm), along the upper Nam Hinboun, Bolikhamsai Province, Khammouan Limestone NBCA, Lao PDR, 17°58'N, 104°49'E." 17.96667 104.8167 NA NA Laos|Vietnam Asia Indomalaya VU 0 0 0 Hipposideros_scutinares 0 sciname match Hipposideros_scutinares 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Hipposideridae Hipposideros scutinares Robinson, Jenkins, Francis & Fulford 2003 0 Acta Chiropterol. 5:33 Shield-nosed Leaf-nosed Bat None. Laos, Khammouan Limestone NBCA, Bolikhamsai Province, along the upper Nam Hinboun, Ban Khankeo, 17Ëš58’N, 104Ëš49’E. Laos, Vietnam. Not Listed Vulnerable swinhoii species group. Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Hipposideros scutinares; Hipposideros scutinares; Hipposideros scutinares; Hipposideros scutinares; Hipposideros scutinares; Hipposideros scutinares; scutinares; Phyllorhine à blason; Nasenschild-Rundblattnase; Hiposidérido de escudo; Shield-nosed Roundleaf Bat; Shield-nosed Leaf-nosed Bat; Shield-nosed Roundleaf Bat; Shield-nosed Leaf-nosed Bat; Shield-nosed Leaf-nosed Bat; H. scutinares