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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L434	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis	Hipposideros rotalis		[MSW3] bicolor species group.; [HMW] Hipposideros rotalis Francis, Kock & Habersetzer, 1999 , “Ban Keng Bit, 18°15’ N , 104°34’ E , Nam (River) Kading, Bolikhamxai Prov[ince]., Laos .” Hipposideros rotalis was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic.; [batnames2022]  bicolor species group.; [batnames2023]  bicolor species group.; [batnames2025_1.7] bicolorspecies group.														rotalis				rotalis 	rotalis 			rotalis C. M. Francis, Kock, & Habersetzer, 1999						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros rotalis	Hipposideros		rotalis	Francis, Kock, and Habersetzer		1999		Senkenbergiana Biol.	79		266		Laotian Leaf-nosed Bat	Laos, Bolikhamxai Prov., Nam (River) Kading, Ban Keng Bit, 18°15'N, 104°34'E.	Laos.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		bicolor species group.	03BD87A2C669A21BF872FECBF5B74BD3	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	258	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C669A21BF872FECBF5B74BD3.xml	Hipposideros rotalis	Hipposideridae	Hipposideros	rotalis	Francis, Kock & Habersetzer	1999	Phyllorhine du Laos @fr | Laos-Rundblattnase @de | Hiposidérido de Laos @es | Laotian Roundleaf Bat @en	Hipposideros rotalis Francis, Kock & Habersetzer, 1999 , “Ban Keng Bit, 18°15’ N , 104°34’ E , Nam (River) Kading, Bolikhamxai Prov[ince]., Laos .” Hipposideros rotalis was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic.	Endemic to C Laos .	Tail 31-2-39 mm, ear 24-3-26-7 mm, forearm 45-49 mm; weight 11 g . Ears of the Laotian Leaf-nosed Bat are broad and concave below tip. Anterior noseleaf is broad and its anterior margin is rounded. It has just one supplementary lateral leaflet. Intemarial disk is large. In posterior noseleaf, three vertical septa form four noticeable cells on frontal surface. Fur is brownish on dorsum, paler on ventral part; base of each hair is white, intermediate portion is brown, and tip lighter.	Primary evergreen forests and dry evergreen-dipterocarp mixed forests.	The Laotian Leaf-nosed Bat probably forages in forest habitats. Its diet is based on insects.	No information.	The Laotian Leaf-nosed Bat has been reported roosting in limestone caves. Call frequency of the F segment is c.69-71 kHz.	The Laotian Leaf-nosed Bat has usually been observed roosting in small groups.	Classified as Least Concern on The IUCNRed List. The Laotian Leaf-nosed Bat is only known from some registers. Population is thought to be relatively large, occupying a range of over 20,000 km 2 and is thought that it may tolerate a degree of disturbance. Very tittle is known about the ecology and status of this species, but to date no major threats have been detected.	Francis (2008b) | Guillén-Servent & Francis (2006) | Murray et al. (2012) | Pader et al. (2017) | Thom as et al. (2013) | Thong Vu Dinh, Dietz et al. (2012)		88. Laotian Leaf-nosed Bat Hipposideros rotalis French: Phyllorhine du Laos / German: Laos-Rundblattnase I Spanish: Hiposidérido de Laos Other common names: Laotian Roundleaf Bat Taxonomy. Hipposideros rotalis Francis, Kock & Habersetzer, 1999 , “Ban Keng Bit, 18°15’ N , 104°34’ E , Nam (River) Kading, Bolikhamxai Prov[ince]., Laos .” Hipposideros rotalis was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic. Distribution. Endemic to C Laos . Descriptive notes. Tail 31-2-39 mm, ear 24-3-26-7 mm, forearm 45-49 mm; weight 11 g . Ears of the Laotian Leaf-nosed Bat are broad and concave below tip. Anterior noseleaf is broad and its anterior margin is rounded. It has just one supplementary lateral leaflet. Intemarial disk is large. In posterior noseleaf, three vertical septa form four noticeable cells on frontal surface. Fur is brownish on dorsum, paler on ventral part; base of each hair is white, intermediate portion is brown, and tip lighter. Habitat. Primary evergreen forests and dry evergreen-dipterocarp mixed forests. Food and Feeding. The Laotian Leaf-nosed Bat probably forages in forest habitats. Its diet is based on insects. Breeding. No information. Activity patterns. The Laotian Leaf-nosed Bat has been reported roosting in limestone caves. Call frequency of the F segment is c.69-71 kHz. Movements, Home range and Social organization. The Laotian Leaf-nosed Bat has usually been observed roosting in small groups. Status and Conservation. Classified as Least Concern on The IUCNRed List. The Laotian Leaf-nosed Bat is only known from some registers. Population is thought to be relatively large, occupying a range of over 20,000 km 2 and is thought that it may tolerate a degree of disturbance. Very tittle is known about the ecology and status of this species, but to date no major threats have been detected. Bibliography. Francis (2008b), Guillén-Servent & Francis (2006), Murray eta/. (2012), Pader eta/. (2017),Thom as eta/. (2013), Thong Vu Dinh, Dietz eta/. (2012).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros rotalis	Hipposideros		rotalis	Francis, Kock & Habersetzer	1999	0	Senkenbergiana Biol.	83:26:00	Laotian Leaf-nosed Bat	None.	Laos, Bolikhamxai Prov., Nam (River) Kading, Ban Keng Bit, 18Â°15'N, 104Â°34'E.	Laos.	Not listed.	Least Concern	 bicolor species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros rotalis	23	Laotian Leaf-nosed Bat	Laotian Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	rotalis	Francis, Kock, & Habersetzer	1999	0						"Ban Keng Bit, 18Â°15' N, 104Â°34' E, Nam (River) Kading, Bolikhamxai Prov[ince]., Laos."	18.25	104.57	rotalis Francis, Kock, & Habersetzer, 1999	NA	NA	Laos	Asia	Indomalaya	LC	0	0	0	Hipposideros_rotalis	0	sciname match	Hipposideros_rotalis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136477	Hipposideros rotalis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	rotalis	Francis, Kock &; Habersetzer, 1999		20000000	Hipposideros rotalis	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	This species is confirmed as Least Concern; although there is not much information on the population status or ecology of this species it is unlikely to meet any of the criteria for Vulnerable or higher listing. Its extent of occurrence is likely larger than 20,000 kmÂ²; there are no data on population size, but there is no reason to believe it is less than 10,000 individuals. Although the population is likely declining due to loss of habitat, it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Little is known of the ecology or roosting sites of this species. Individuals were captured in dry forests, in some cases with some degree of disturbance, generally in areas with extensive limestone karst nearby. However, as no roosts have been found, it is unknown whether it is dependent on caves.	The threats to this species are not known, though it has likely been subject to significant habitat loss due to clearing of forests for agriculture, and at least some of the habitat around the type locality.	This species is so far reported from only 5 sites, with only 1-3 individuals captured at each site. However, as no roosts have been found, this does not provide reliable information on population size or density.	Unknown	This species has been reported from 5 localities in central Lao PDR (Thomas et al. 2013). The type locality is Ban Keng Bit (18Â°15'N, 104Â°34'E), Nam (River) Kading, Bolikhamxai province. It is quite likely that the species may be found in central Vietnam, given the presence of similar habitat there.		Terrestrial	Several of the records are from protected areas although there is habitat loss within these (C. Francis pers. comm.).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		rotalis	Francis, Kock & Habersetzer	1999	0	Senkenbergiana Biol.	83:26:00	Laotian Leaf-nosed Bat	None.	Laos, Bolikhamxai Prov., Nam (River) Kading, Ban Keng Bit, 18Â°15'N, 104Â°34'E.	Laos.	Not listed.	Least Concern	 bicolor species group.	Hipposideros rotalis	1004631	23	Laotian Leaf-nosed Bat	Laotian Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	rotalis	Francis, Kock, & Habersetzer	1999	0						"Ban Keng Bit, 18Â°15' N, 104Â°34' E, Nam (River) Kading, Bolikhamxai Prov[ince]., Laos."	18.25	104.5667	rotalis Francis, Kock, & Habersetzer, 1999	NA	NA				Laos	Asia	Indomalaya	LC	0	0	0	Hipposideros_rotalis	0	sciname match	Hipposideros_rotalis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_rotalis	1004631	23	Laotian Roundleaf Bat	Laotian Roundleaf Bat|Laotian Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	rotalis	C. M. Francis, Kock, & Habersetzer	0	Hipposideros rotalis	Francis, C.M., Kock, D. and Habersetzer, J. 1999-11-30. Sibling species of _Hipposideros ridleyi_ (Mammalia, Chiroptera, Hipposideridae). Senckenbergiana biologica 79(2):255-270.		SMF:MAMM:85786	holotype		"Ban Keng Bit, 18Â°15' N, 104Â°34' E, Nam (River) Kading, Bolikhamxai Prov[ince]., Laos."	18.25	104.5667	NA	NA				Laos	Asia	Indomalaya	LC	0	0	0	Hipposideros_rotalis	0	sciname match	Hipposideros_rotalis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		rotalis	Francis, Kock & Habersetzer	1999	0	Senkenbergiana Biol.	83:26:00	Laotian Leaf-nosed Bat	None.	Laos, Bolikhamxai Prov., Nam (River) Kading, Ban Keng Bit, 18Â°15'N, 104Â°34'E.	Laos.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136477/21985931/' target='_blank'>Least Concern</a>	bicolorspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros rotalis; Hipposideros rotalis; Hipposideros rotalis; Hipposideros rotalis; Hipposideros rotalis; Hipposideros rotalis; rotalis; Phyllorhine du Laos; Laos-Rundblattnase; Hiposidérido de Laos; Laotian Roundleaf Bat; Laotian Leaf-nosed Bat; Laotian Roundleaf Bat; Laotian Leaf-nosed Bat; Laotian Leaf-nosed Bat; H. rotalis
