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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L430	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Hipposideros bicolor [synonym of]	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona	Hipposideros pomona		[MSW2] Formerly included in H. bicolor but see Hill et al. (1986).; [MSW3] bicolor species group. Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet and Hill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b).; [HMW] Hipposideros pomona K Andersen, 1918 , “Haleri, N[orth]. Coorg [= Kodagu],” Karnataka State , south-western India . Hipposideros pomona was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic.; [batnames2022]  bicolor species group. Includes  megalotis , which was formerly the sole member of the genus Paracoelops ; see Thong et al. (2012). Does not include gentilis ; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).; [MDD2022] previously included H. gentilis; [IUCN] This species belongs to bicolor species group. This species was listed under Hipposideros bicolor (Temminck, 1834), but is now considered distinct (Hill et al. 1986, Srinivasulu and Srinivasulu 2012). Based on morphology and bacular structure, Srinivasulu and Srinivasulu (2018) upgraded the taxon gentilis Andersen, 1918 to specific rank, and suggested that H. pomona Andersen, 1918 sensu stricto is restricted to southern India, and H. gentilis sensu lato occurs from northeast India to Southeast Asia.; [batnames2023]  bicolor species group. Includes  megalotis , which was formerly the sole member of the genus Paracoelops ; see Thong et al. (2012). Does not include gentilis ; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).; [MDD2023] previously included H. gentilis; [MDD2025_2.0] previously included H. gentilis; [batnames2025_1.7] bicolor species group. Includes  megalotis, which was formerly the sole member of the genus Paracoelops; see Thong et al. (2012). Does not include gentilis; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).; [MDD2025_2.2] previously included H. gentilis						gentilis, sinensis.	pomona, gentilis, sinensis	pomona, gentiles, sinensis				pomona	pomona - megalotis, maybe macrobullatus	pomona	This species belongs to bicolor species group. This species was listed under Hipposideros bicolor (Temminck, 1834), but is now considered distinct (Hill et al. 1986, Srinivasulu and Srinivasulu 2012). Based on morphology and bacular structure, Srinivasulu and Srinivasulu (2018) upgraded the taxon gentilis Andersen, 1918 to specific rank, and suggested that H. pomona Andersen, 1918 sensu stricto is restricted to southern India, and H. gentilis sensu lato occurs from northeast India to Southeast Asia.	pomona	pomona - megalotis	pomona	pomona	pomona	pomona - megalotis	pomona Andersen, 1918						N/A					Distribution: Ranging from India, east to southern China and south to the Malay pen insula.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		India – Thailand, S China; ref. 4.118	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Andersen	1918	Ann. Mag. Nat. Hist., ser. 9, 2:380, 381.	Formerly included in H. bicolor but see Hill et al. (1986).	India to S China and W Malaysia.	India, Mysore, Coorg, Haleri.		ANDERSEN	1918	Ears relatively large. Periotics relatively large, the width of each definitely greater than their distance apart. Size fairly small (forearm length, 37-44 mm).	Distribution: Ranging from India, east to southern China and south to the Malay pen insula.	Three subspecies are recognized (HILL & al. 1986):	H. p. pomona (southern India), H. p. gentilis (northern In dia to Vietnam and Malaya), H. p. sinensis (southern China to northern Thailand).	60	species	H. pomona	ANDERSEN	1918	Hipposideros	genus	Hipposideros pomona				Ears relatively large. Periotics relatively large, the width of each definitely greater than their distance apart. Size fairly small (forearm length, 37- 44 mm).	Three subspecies are recognized (HILL & al. 1986):		3. H. pomona ANDERSEN 1918 [bicolor group].	3	_H. p. pomona_ Andersen, 1918			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros pomona	Hipposideros		pomona	K. Andersen		1918		Ann. Mag. Nat. Hist., ser. 9	2		380, 381		Pomona Leaf-nosed Bat	India, Mysore, N Coorg, Haleri (a few miles N of Mercara, Coorg Dist., Karnataka).	Bangladesh and India to Burma, Thailand, Laos, Cambodia, Vietnam, S China and W Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	gentilis K. Andersen, 1918; sinensis K. Andersen, 1918.	bicolor species group. Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet and Hill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997) and Hendrichsen et al. (2001b).	03BD87A2C668A21AF898FEA4F2CA4B5E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	257	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C668A21AF898FEA4F2CA4B5E.xml	Hipposideros pomona	Hipposideridae	Hipposideros	pomona	K Andersen	1918	Phyllorhine pomone @fr | Pomona-Rundblattnase @de | Hiposidérido pomona @es | Pomona Roundleaf Bat @en	Hipposideros pomona K Andersen, 1918 , “Haleri, N[orth]. Coorg [= Kodagu],” Karnataka State , south-western India . Hipposideros pomona was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic.	S India ( Andhra Pradesh , Karnataka , Kerala , and Tamil Nadu ).			The Pomona Leaf-nosed Bat feeds in gaps of the forest understory.	No information.	The Pomona Leaf-nosed Bat roosts in caves, rock crevices, or hollow trees. Call frequency is 123-128 kHz.	No information.	Classified as Least Concern on IUCN Red List. As the revised species is now restricted to southern India , its population status needs to be re-assessed. The Pomona Leaf-nosed Bat is likely to qualify to be listed under threatened category and may require conservation measures.	Bates, Bumrungsri, Csorba, Molur & Srinivasulu (2008a) | Corbet & Hill (1992) | Douangboubpha, Bumrungsri, Soisook, Satasook et al. (2010) | Francis (2008a) | Simmons (2005) | Wordley et al. (2014)	https://zenodo.org/record/3749207/files/figure.png	85. Pomona Leaf-nosed Bat Hipposideros pomona French: Phyllorhine pomone I German: Pomona-Rundblattnase / Spanish: Hiposidérido pomona Other common names: Pomona Roundleaf Bat Taxonomy. Hipposideros pomona K Andersen, 1918 , “Haleri, N[orth]. Coorg [= Kodagu],” Karnataka State , south-western India . Hipposideros pomona was formerly included in the bicolor species group but is now placed in the new ater species group. Monotypic. Distribution. S India ( Andhra Pradesh , Karnataka , Kerala , and Tamil Nadu ). Descriptive notes. Ear 18—19 mm , forearm 38—39 mm . Pelage of the Pomona Leaf-nosed Bat is generally dark brown, with hairs paler at base; ventral side is slightly paler. Noseleaf is wide and rounded, with no lateral supplementary leaflet. Intemarial septum is parallel-sided. Rostrum is narrow and poorly inflated. Baculum is very small, with straight shaft and simple blunt base and tip. Habitat No information. Food and Feeding. The Pomona Leaf-nosed Bat feeds in gaps of the forest understory. Breeding. No information. Activity patterns. The Pomona Leaf-nosed Bat roosts in caves, rock crevices, or hollow trees. Call frequency is 123-128 kHz. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on IUCN Red List. As the revised species is now restricted to southern India , its population status needs to be re-assessed. The Pomona Leaf-nosed Bat is likely to qualify to be listed under threatened category and may require conservation measures. Bibliography. Bates, Bumrungsri, Csorba, Molur & Srinivasulu (2008a), Corbet & Hill (1992), Douangboubpha, Bumrungsri, Soisook, Satasook eta/. (2010), Francis (2008a), Simmons (2005), Wordley eta/. (2014).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros pomona	Hipposideros		pomona	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 380, 381	Pomona Leaf-nosed Bat	 megalotis Dorst, 1947.	India, Mysore, N Coorg, Haleri (a few miles N of Mercara, Coorg Dist., Karnataka).	S India.	Not listed.	Endangered	 bicolor species group. Includes  megalotis , which was formerly the sole member of the genus Paracoelops ; see Thong et al. (2012). Does not include gentilis ; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros pomona	23	Pomona Leaf-nosed Bat	Pomona Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	pomona	K. Andersen	1918	0	Hipposideros_pomona	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History, ser. 9, 2, 380.	https://www.biodiversitylibrary.org/item/71919#page/394/mode/1up	ZSI 21529		"Haleri, N[orth]. Coorg [= Kodagu]," Karnataka State, south-western India.			pomona K. Andersen, 1918	previously included H. gentilis	Srinivasulu, B., & Srinivasulu, C. (2018). In plain sight: Bacular and noseleaf morphology supports distinct specific status of Roundleaf Bats Hipposideros pomona Andersen, 1918 and Hipposideros gentilis Andersen, 1918 (Chiroptera: Hipposideridae). Journal of Threatened Taxa, 10(8), 12018-12026.	India	Asia	Indomalaya	EN	0	0	0	Hipposideros_pomona	0	sciname match	Hipposideros_pomona	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	200000000	Hipposideros pomona	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	pomona	K. Andersen, 1918	This species belongs to bicolor species group. This species was listed under Hipposideros bicolor (Temminck, 1834), but is now considered distinct (Hill et al. 1986, Srinivasulu and Srinivasulu 2012). Based on morphology and bacular structure, Srinivasulu and Srinivasulu (2018) upgraded the taxon gentilis Andersen, 1918 to specific rank, and suggested that H. pomona Andersen, 1918 sensu stricto is restricted to southern India, and H. gentilis sensu lato occurs from northeast India to Southeast Asia.	200000000	Hipposideros pomona	Endangered	B2ab(ii,iii,iv,v)	2020	2020-07-05 00:00:00 UTC	3.1	English	Hipposideros pomona is assessed as ;Endangered under criterion B (EN B2ab(ii,iii,iv,v)). The species is known from only 8-10 roost locations where they spend over half of their lives. Based on the limited roosts, its area of occupancy (AOO) is calculated ranging from 32 to 40 km². The species has a restricted and fragmented distribution, and its population is inferred to be declining due to a continuing loss and degradation of its subterranean roosts and foraging habitat due to human encroachment.	<p>This species is known from fewer than 10 records restricted to parts of southern Karnataka, Kerala, and parts of Tamil Nadu. Little is known about the habitat or ecology of this species except that it roosts in small colonies of a few individuals in caves and crevices in subterranean habitats (Molur et al. 2002, Bates and Harrison 1997). This species has been observed to roost in old disused wells and small caves (A. Madhavan pers. com. 2016). There is a continuing decline in area and quality of habitat due to disturbance and loss of subterranean roosts. ; ;</p>	<p>There is limited information on the most important threats to this species. The type locality of this taxon from southern Karnataka was recently visited and no individuals of this species could be found. The subterranean roost and foraging habitats of this species is under threat due to human encroachment and habitat change.</p>	<p>There is very little data on the population status of this species. It roosts in small colonies of a few individuals in caves and crevices in subterranean habitats (Molur et al. 2002, Bates and Harrison 1997) and there is an inferred population decline due to a continuing decline in area and quality of its subterranean habitats.</p>	Decreasing	This species is endemic to India and is known from fewer than 10 records from southern Karnataka, Kerala, and Tamil Nadu (Srinivasulu and Srinivasulu 2018). Records of this taxon from Andhra Pradesh, India is erroneous based on misidentified H. fulvus. ;This species has been taken from 100 m asl to around 1,200 m asl.	Nothing ;is known ;about the use and trade of this species.	Terrestrial	<p>This taxon is restricted in distribution and is very uncommon in its range. Although the species occurs in protected areas, there are no conservation actions in place for this species. Further studies are needed to understand its population status and trends, threats, and habitat requirements are needed to formulate effective ;conservation action plans.</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		pomona	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 380, 381	Pomona Leaf-nosed Bat	 megalotis Dorst, 1947.	India, Mysore, N Coorg, Haleri (a few miles N of Mercara, Coorg Dist., Karnataka).	S India.	Not listed.	Endangered	 bicolor species group. Includes  megalotis , which was formerly the sole member of the genus Paracoelops ; see Thong et al. (2012). Does not include gentilis ; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).	Hipposideros pomona	1004628	23	Pomona Leaf-nosed Bat	Pomona Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	pomona	K. Andersen	1918	0	Hipposideros_pomona	Andersen, K. (1918). Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History, ser. 9, 2, 380.	https://www.biodiversitylibrary.org/item/71919#page/394/mode/1up	ZSI 21529		"Haleri, N[orth]. Coorg [= Kodagu]," Karnataka State, south-western India.			pomona K. Andersen, 1918	previously included H. gentilis	Srinivasulu, B., & Srinivasulu, C. (2018). In plain sight: Bacular and noseleaf morphology supports distinct specific status of Roundleaf Bats Hipposideros pomona Andersen, 1918 and Hipposideros gentilis Andersen, 1918 (Chiroptera: Hipposideridae). Journal of Threatened Taxa, 10(8), 12018-12026.				India	Asia	Indomalaya	EN	0	0	0	Hipposideros_pomona	0	sciname match	Hipposideros_pomona	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_pomona	1004628	23	Pomona Roundleaf Bat	Pomona Roundleaf Bat|Pomona Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	pomona	Andersen	0	Hipposideros pomona	Andersen, K.C. 1918-10-01. Diagnoses of new bats of the families Rhinolophidae and Megadermatidae. Annals and Magazine of Natural History (9)2(10):374-384.	https://www.biodiversitylibrary.org/page/22103263	BMNH:Mamm:1918.8.3.4, ZSI 21529	holotype	https://data.nhm.ac.uk/object/66928609-c5f0-42a5-961a-ff3416af5efe	"Haleri, N[orth]. Coorg [= Kodagu]," Karnataka State, south-western India.			previously included H. gentilis	Srinivasulu, B., & Srinivasulu, C. (2018). In plain sight: Bacular and noseleaf morphology supports distinct specific status of Roundleaf Bats Hipposideros pomona Andersen, 1918 and Hipposideros gentilis Andersen, 1918 (Chiroptera: Hipposideridae). Journal of Threatened Taxa, 10(8), 12018-12026.				India	Asia	Indomalaya	EN	0	0	0	Hipposideros_pomona	0	sciname match	Hipposideros_pomona	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		pomona	K. Andersen	1918	0	Ann. Mag. Nat. Hist.	ser. 9, 2: 380, 381	Pomona Leaf-nosed Bat	megalotis Dorst, 1947.	India, Mysore, N Coorg, Haleri (a few miles N of Mercara, Coorg Dist., Karnataka).	S India.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/180990825/180990948/' target='_blank'>Endangered</a>	bicolor species group. Includes  megalotis, which was formerly the sole member of the genus Paracoelops; see Thong et al. (2012). Does not include gentilis; see Srinvasulu and Srinvasulu (2018). Formerly included in bicolor but see Hill et al (1986). May be conspecific with macrobullatus; see Corbet andHill (1992). Some specimens from peninsular India previously referred to this species were subsequently removed to form the type series of hypophyllus. Reviewed in part by Bates and Harrison (1997), Hendrichsen et al. (2001b), and Douangboubpha et al. (2010).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros pomona; Hipposideros pomona; Hipposideros pomona; Hipposideros pomona; Hipposideros pomona; Hipposideros pomona; pomona; gentiles; sinensis; megalotis; maybe macrobullatus; pomona; Phyllorhine pomone; Pomona-Rundblattnase; Hiposidérido pomona; Pomona Roundleaf Bat; Pomona Leaf-nosed Bat; Pomona Roundleaf Bat; Pomona Leaf-nosed Bat; Pomona Leaf-nosed Bat; H. pomona