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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L420	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae	Hipposideros marisae		[MSW3] bicolor species group.; [HMW] Hipposideros marisae Aellen, 1954 , White Leopard Rock, Duékoué, Ivory Coast . Hipposideros marisae was previously included in the bicolor species group, but is now placed in the speoris species group. Monotypic.; [batnames2022]  bicolor species group.; [batnames2023]  bicolor species group.; [batnames2025_1.7] bicolorspecies group.														marisae				marisae 	marisae 			marisae Aellen, 1954		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Aellen's leaf-nosed bat	Ivory Coast, Guinea	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros marisae	Ivory Coast, Duekoue, White Leopard Rock.	Aellen	1954	Rev. Suisse Zool., 61:474.	Distribution: Confined to a small area in western Africa, including parts of Guinea, Liberia, and Ivory Coast.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Aellen's leaf-nosed bat	Ivory Coast – Guinea	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Aellen	1954	Rev. Suisse Zool., 61:474.		Ivory Coast, Liberia, Guinea.	Ivory Coast, Duekoue, White Leopard Rock.		AELLEN	1954	In ternarial septum expanded to form an ellipsoidal structure. A single, very small lateral leaflet pres ent. Size relatively small (forearm length, 40-41 mm).	Distribution: Confined to a small area in western Africa, including parts of Guinea, Liberia, and Ivory Coast.	No subspecies.		62	species	H. marisae	AELLEN	1954	Hipposideros	genus	Hipposideros marisae				In ternarial septum expanded to form an ellipsoidal structure. A single, very small lateral leaflet pres ent. Size relatively small (forearm length, 40-41 mm).	No subspecies.		19. H. marisae AELLEN 1954 [bicolor group].	19	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros marisae	Hipposideros		marisae	Aellen		1954		Rev. Suisse Zool.	61		474		Aellen's Leaf-nosed Bat	Côte d’Ivoire, Duékoué, White Leopard Rock.	Côte d’Ivoire, Liberia, Guinea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		bicolor species group.	03BD87A2C664A216FF55EFC7F2B844F7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	245	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C664A216FF55EFC7F2B844F7.xml	Hipposideros marisae	Hipposideridae	Hipposideros	marisae	Aellen	1954	Phyllorhine d'Aellen @fr | Aellen-Rundblattnase @de | Hiposidérido de Aellen @es | Aellen's Roundleaf Bat @en	Hipposideros marisae Aellen, 1954 , White Leopard Rock, Duékoué, Ivory Coast . Hipposideros marisae was previously included in the bicolor species group, but is now placed in the speoris species group. Monotypic.		Head—body 41—47 mm , tail 19-21 mm, ear 15-17 mm, hindfoot 6-8 mm, forearm 38—42 mm ; weight 5 g . There are no obvious sexual differences in pelage or size. Aellen’s Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with large but simple noseleaf that has one lateral leaflet. A frontal sac is present in both sexes posterior to noseleaf. Intemarial septum is swollen and partially covering the nares. Ears are separate, and relatively large. Fur is fine and short, gray-brown dorsally and slightly lighter ventrally.	Tropical forest in lowland and hilly situations.	Exact diet of Aellen’s Leaf-nosed Bat is not known, but it is insectivorous.	No information.	Aellen’s Leaf-nosed Bat rests up in caves and old mine adits. Echolocation call includes a F component at 146 kHz.	Aellen’s Leaf-nosed Bat roosts singly or in small groups, typically consisting of 2-6 individuals. It may share its roost with other species of Hipposideros .	Classified as Vulnerable on The IUCNRed List due to its small, fragmented distribution, which is under threat of deforestation. Disturbance ofits cave roosting sites is another threat.	Fahr (2013h) | Monadjem, Richards & Denys (2016) | Rosevear (1965) | Wolton et al. (1982)	https://zenodo.org/record/3749133/files/figure.png	49. Aellen’s Leaf-nosed Bat Hipposideros marisae French: Phyllorhine d'Aellen / German: Aellen-Rundblattnase / Spanish: Hiposidérido de Aellen Other common names: Aellen's Roundleaf Bat Taxonomy. Hipposideros marisae Aellen, 1954 , White Leopard Rock, Duékoué, Ivory Coast . Hipposideros marisae was previously included in the bicolor species group, but is now placed in the speoris species group. Monotypic. Distribution. Very patchily in W Africa with records in Guinea , Sierra Leone , Liberia , and Ivory Coast Descriptive notes. Head—body 41—47 mm , tail 19-21 mm, ear 15-17 mm, hindfoot 6-8 mm, forearm 38—42 mm ; weight 5 g . There are no obvious sexual differences in pelage or size. Aellen’s Leaf-nosed Bat has short, rounded wings. Muzzle is relatively short with large but simple noseleaf that has one lateral leaflet. A frontal sac is present in both sexes posterior to noseleaf. Intemarial septum is swollen and partially covering the nares. Ears are separate, and relatively large. Fur is fine and short, gray-brown dorsally and slightly lighter ventrally. Habitat. Tropical forest in lowland and hilly situations. Food and Feeding. Exact diet of Aellen’s Leaf-nosed Bat is not known, but it is insectivorous. Breeding. No information. Activity patterns. Aellen’s Leaf-nosed Bat rests up in caves and old mine adits. Echolocation call includes a F component at 146 kHz. Movements, Home range and Social organization. Aellen’s Leaf-nosed Bat roosts singly or in small groups, typically consisting of 2-6 individuals. It may share its roost with other species of Hipposideros . Status and Conservation. Classified as Vulnerable on The IUCNRed List due to its small, fragmented distribution, which is under threat of deforestation. Disturbance ofits cave roosting sites is another threat. Bibliography. Fahr (2013h), Monadjem, Richards & Denys (2016), Rosevear (1965), Wolton et al. (1982).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros marisae	Hipposideros		marisae	Aellen	1954	0	Rev. Suisse Zool.	68:54:00	Aellen's Leaf-nosed Bat	None.	CÃ´te dâIvoire, DuÃ©kouÃ©, White Leopard Rock.	CÃ´te dâIvoire, Liberia, Guinea.	Not listed.	Vulnerable	 bicolor species group.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros marisae	23	Aellen's Leaf-nosed Bat	Aellen's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	marisae	Aellen	1954	0	Hipposideros_marisae	Aellen, V. (1954). Description d'un nouvel Hipposideros (Chiroptera) de la CÃ´te d'Ivoire. Revue suisse de zoologie, 61, 474.	https://www.biodiversitylibrary.org/item/126660#page/496/mode/1up	MHNG 924.090		White Leopard Rock, DuÃ©kouÃ©, Ivory Coast.			marisae Aellen, 1954	NA	NA	Guinea|Sierra Leone|Liberia|CÃ´te d'Ivoire	Africa	Afrotropic	VU	0	0	0	Hipposideros_marisae	0	sciname match	Hipposideros_marisae	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	10149	Hipposideros marisae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	marisae	Aellen, 1954		20000000	Hipposideros marisae	Vulnerable	A4c	2020	2019-06-21 00:00:00 UTC	3.1	English	Hipposideros marisae ;is listed as ;Vulnerable under criterion A4c as its global population is suspected to have declined by &gt;30% over the past 10 years (two generations) and is expected to continue declining at the same rate over the next 5 years (one generation; generation length = 5 years, Pacifici et al. 2013). The species' extent of occurrence (EOO), area of occupancy (AOO), and habitat is projected to continue to decline ;due to continuing threats from forest loss and degradation and cave/mine loss, degradation, and disturbance.	The natural history of this species is not well known. In general,it may be found in both drier and wetter types of lowland and sub-montane rainforest (Fahr 2013). Roost sites include natural caves, boulder caves, overhanging cliffs, and disused mine adits (Monadjem et al. 2013, 2016).	The species is threatened by general deforestation, mining operations on Mount Nimba, the disturbance of roosting caves, and the loss of old mine adits to ongoing mining operations. Due to its small size it is unlikely to harvested for food. Disturbance at roost sites are also likely to be significant threats to this species (Fahr 2013). This species also occurs in areas of high population density, so persecution and human-bat conflict likely to be a significant threat.	This is generally considered to be a very rare species, occurring in low numbers. For example, at Mount Nimba, an old mine adit was occupied by five adult Hipposideros marisae compared with about 200 H. cf. ruber . In a neighbouring adit there were a further two H. marisae and over 50 ;H. ;cf. ;ruber (A. Monadjem, 2016). The global population is suspected to have declined in the past and will continue to decline due to loss and degradation of the species habitat, especially the loss of cave and mine roosts.	Decreasing	This West African species has been recorded from seven localities in Guinea, Liberia (central and western uplands), and CÃ´te d'Ivoire. It also occurs in the Gola Forest in eastern Sierra Leone. It may also occur in Ghana, however, there are no records from this country (Grubb et al. 1998). It is a lowland species generally recorded between sea level and 650 m asl. However, it has been recorded roosting in old mine adits at 720 m and 920 m asl adjacent to Mount Nimba (Monadjem et al. 2013, Monadjem 2016).		Terrestrial	This species is endemic to Africa, and found within the Mount Nimba Strict Nature Reserve World Heritage Site in Guinea as well as the East Nimba Nature Reserve in Liberia, and Gola Forest in Sierra Leone. There is a need to protect important roost sites and areas of suitable forest habitat. Further studies are needed into the distribution, population size and trend, ecology and significant threats to this species. Given the extensive mining, logging and agricultural expansion and potential cave disturbance in the area it is also recommended to engage key stakeholders and work with local communities and government authorities given they have an endemic rare species in the area. Specific research is needed to understand disturbance and if protection or gating of important maternity roosts is viable.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		marisae	Aellen	1954	0	Rev. Suisse Zool.	68:54:00	Aellen's Leaf-nosed Bat	None.	CÃ´te dâ€™Ivoire, DuÃ©kouÃ©, White Leopard Rock.	CÃ´te dâ€™Ivoire, Liberia, Guinea.	Not listed.	Vulnerable	 bicolor species group.	Hipposideros marisae	1004619	23	Aellen's Leaf-nosed Bat	Aellen's Roundleaf Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	marisae	Aellen	1954	0	Hipposideros_marisae	Aellen, V. (1954). Description d'un nouvel Hipposideros (Chiroptera) de la CÃ´te d'Ivoire. Revue suisse de zoologie, 61, 474.	https://www.biodiversitylibrary.org/item/126660#page/496/mode/1up	MHNG 924.090		White Leopard Rock, DuÃ©kouÃ©, Ivory Coast.			marisae Aellen, 1954	NA	NA				Guinea|Sierra Leone|Liberia|CÃ´te d'Ivoire	Africa	Afrotropic	VU	0	0	0	Hipposideros_marisae	0	sciname match	Hipposideros_marisae	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_marisae	1004619	23	Aellen's Roundleaf Bat	Aellen's Roundleaf Bat|Aellen's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	marisae	Aellen	0	Hipposideros marisae	Aellen, V. 1954. Description d'un nouvel _Hipposideros_ (Chiroptera) de la CÃ´te d'Ivoire. Revue suisse de Zoologie 61:473-483.	https://www.biodiversitylibrary.org/page/40778042	MHNG 924.90	holotype		White Leopard Rock, DuÃ©kouÃ©, Ivory Coast.			NA	NA				Guinea|Sierra Leone|Liberia|Cote d'Ivoire	Africa	Afrotropic	VU	0	0	0	Hipposideros_marisae	0	sciname match	Hipposideros_marisae	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		marisae	Aellen	1954	0	Rev. Suisse Zool.	68:54:00	Aellen's Leaf-nosed Bat	None.	CÃ´te dâ€™Ivoire, DuÃ©kouÃ©, White Leopard Rock.	CÃ´te dâ€™Ivoire, Liberia, Guinea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/10149/22101390/' target='_blank'>Vulnerable</a>	bicolorspecies group.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros marisae; Hipposideros marisae; Hipposideros marisae; Hipposideros marisae; Hipposideros marisae; Hipposideros marisae; marisae; Phyllorhine d'Aellen; Aellen-Rundblattnase; Hiposidérido de Aellen; Aellen's Roundleaf Bat; Aellen's Leaf-nosed Bat; Aellen's Roundleaf Bat; Aellen's Leaf-nosed Bat; Aellen's Leaf-nosed Bat; H. marisae
