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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L413	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	N/A	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus	Hipposideros larvatus		[MSW3] larvatus species group. Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. See also Hill (1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993).; [HMW] Rhinolophus larvatus Horsfield, 1823 , Java , Indonesia . Hipposideros larvatus is in the larvatus species group. Clarification is needed of differentiating characters and geographic limits with H. grandis , particularly the taxon leptophyllus. Monotypic.; [batnames2022]  larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018).Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993).; [MDD2022] includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; [IUCN] Excludes ;Hipposideros ;<span class="SpellE">grandis</span> , now considered separate species (Thabah ;et al. 2006, ;Kruskop</span> ;2013).; [batnames2023]  larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018; Yuzefovich et al., 2022). Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. Does not include poutensis ; see Yuzefovich et al. (2022). See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993). Includes khasiana (Thabah et al., 2006) as a nomen nudum as no specific holotype was designated.; [MDD2023] previously included H. poutensis and includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; the species limits between H. grandis, H. larvatus, and H. poutensis is currently unresolved and represents a species complex; [MDD2025_2.0] previously included H. poutensis and includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; the species limits between H. grandis, H. larvatus, and H. poutensis is currently unresolved and represents a species complex; [batnames2025_1.7] larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018; Yuzefovich et al., 2022). Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. Does not include poutensis; see Yuzefovich et al. (2022). See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993). Includes khasiana (Thabah et al., 2006) as a nomen nudum as no specific holotype was designated.; [MDD2025_2.2] previously included H. poutensis and includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; the species limits between H. grandis, H. larvatus, and H. poutensis is currently unresolved and represents a species complex						alongensis, barbensis, deformis, grandis, insignis, leptophyllus, neglectus, poutensis, sumbae, vulgaris.	leptophyllus, grandis, alongensis, poutensis, neglectus, barbensis, larvatus, sumbae	larvatus, barbensis, leptophyllus, neglectus, poutensis	deformis, insignis, vulgaris			lanosa, barbensis, leptophyllus, neglectus, poutensis	lanosa - deformis, insignis, vulgaris	larvatus, deformis, insignis, vulgaris, leptophyllus, barbensis, poutensis, neglectus, khasiana	Excludes ;Hipposideros ;<span class="SpellE">grandis</span> , now considered separate species (Thabah ;et al. 2006, ;Kruskop</span> ;2013).	larvatus, barbensis, leptophyllus, neglectus	larvatus - deformis, khasiana, insignis, vulgaris	larvatus, deformis, insignis, vulgaris, leptophyllus, barbensis, neglectus, khasiana	deformis, insignis, larvatus, vulgaris, leptophyllus, barbensis, neglectus, khasiana	barbensis, larvatus, leptophyllus, neglectus	larvatus - deformis, insignis, khasiana, vulgaris	deformis (Horsfield, 1823)|insignis (Horsfield, 1823)|larvatus (Horsfield, 1823)|vulgaris (Horsfield, 1823)|leptophyllus (Dobson, 1874)|barbensis G. S. Miller, 1900|neglectus Sody, 1936|khasiana Thabah, Rossiter, Kingston, Zhang Shuyi, S. Parsons, Khin Mya Mya, Zubaid, & G. Jones, 2006 [nomen nudum]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Horsfield's leaf-nosed bat	Burma – Java, Sumba, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Hipposideros larvatus	Indonesia, Java.	Horsfield	1823	Zool. Res. Java, 6, pl. 9.	Distribution: Ranging from north western India and southern China through the Malay peninsula and Sumatra to Borneo, Java, and the Lesser Sundas.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Horsfield's leaf-nosed bat	N India, S China – Java, Sumba, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Horsfield	1823	Zool. Res. Java, 6, pl. 9.		Bangladesh to Vietnam; Yunnan, Kwangsi and Hainan (China); and through W Malaysia to Sumatra, Java, Borneo, and Sumba (Indonesia), and adjacent small islands.	Indonesia, Java.		HORSFIELD	1823	Anterior noseleaf with a definite median emar gination. Rostrum rounded, supraorbital ridges barely developed. Vomer projecting behind pal ate. Upper canines not grooved and lacking a pos terior cusp. Size medium (forearm length, 53-67 mm).	Distribution: Ranging from north western India and southern China through the Malay peninsula and Sumatra to Borneo, Java, and the Lesser Sundas.	Eight subspecies are cur rently recognized, though there is considerable uncertainty concerning their ranges and validity:	H. I. leptophyllus (northern India), H. I. grandis (Burma, Thailand, southern Vietnam). H. I. alongensis (northern Vietnam), H. I. poutensis (southeastern China), H. I. neglectus (Malay peninsula, Sumatra, Borneo, and most neigh boring small islands), H. I. barbensis (several small islands in the South China Sea), H. I. larvatus (Java, Bali), H. I. sumbae (Sumba island in the lesser Sundas).	66	species	H. larvatus	HORSFIELD	1823	Hipposideros	genus	Hipposideros larvatus				Anterior noseleaf with a definite median emar gination. Rostrum rounded, supraorbital ridges barely developed. Vomer projecting behind pal ate. Upper canines not grooved and lacking a pos terior cusp. Size medium (forearm length, 53-67 mm).	Eight subspecies are cur rently recognized, though there is considerable uncertainty concerning their ranges and validity:		45. H. larvatus (HORSFIELD 1823) [speoris group].	45	_H. l. barbensis_ Miller, 1900; _H. l. larvatus_ (Horsfield, 1823) (synonyms: _deformis_ (Horsfield, 1823), _insignis_ (Horsfield, 1823), _vulgaris_ (Horsfield, 1823)); _H. l. leptophyllus_ (Dobson, 1874); _H. l. neglectus_ Sody, 1936			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Hipposideros larvatus	Hipposideros		larvatus	Horsfield	y	1823		Zool. Res. Java	6		Rhinolophus larvatus, pl. and 10 unno. pp		Intermediate Leaf-nosed Bat	Indonesia, Java.	N and E India and Bangladesh; Yunnan, Kwangsi and Hainan (China); Burma, Thailand, Cambodia, Laos, and Vietnam; W Malaysia to Sumatra, Java, Borneo, and adjacent small islands including Kangean Isls (Indonesia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	deformis Horsfield, 1823; insignis Horsfield, 1823; vulgaris Horsfield, 1823; barbensis Miller, 1900; leptophyllus Dobson, 1874; neglectus Sody, 1936; poutensis Allen, 1906.	larvatus species group. Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. See also Hill (1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993).	03BD87A2C67AA208FF57F253F4564418	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	239	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C67AA208FF57F253F4564418.xml	N/A	Hipposideridae	Hipposideros	larvatus		1823	Phyllorhine de Horsfield @fr | Horsfield-Rundblattnase @de | Hiposidérido de Horsfield @es | Horsfield’s Roundleaf Bat @en | @en | ntermediate Leaf-nosed Bat @en	Rhinolophus larvatus Horsfield, 1823 , Java , Indonesia . Hipposideros larvatus is in the larvatus species group. Clarification is needed of differentiating characters and geographic limits with H. grandis , particularly the taxon leptophyllus. Monotypic.	NE India , NE Bangladesh , S China (including Hainan I), mainland SE Asia, Sumatra (including Simeulue, Nias, and Mentawai Is), Borneo, Java, Kangean, Bali, and many offshore Is.	Head-body 52-75 mm, tail 32-35 mm, ear 21-25 mm, hindfoot 7—12 mm , forearm 50—67 mm ; weight 15—23 g . Greatest skull lengths are 20-4— 22-8 mm, tibia 18-4—24-8 mm. Horsfield’s Leafnosed Bat is very similar to the Grand Leaf-nosed Bat (77. grandis ) but smaller. Ears are broad with pointed tip. There are three supplementary leaflets on each side of noseleaf. Posterior noseleaf is as broad as width of anterior leaf. Thickened and swollen structure behind posterior leaf is well defined in males. Pelage is dark gray-brown to orange-reddish brown. In skull, rostral chambers are well inflated. Sagittal crest is well developed. C1 and C1 are large. Chromosomal complement has 2n = 32 and FN = 60.	Horsfield’s Leaf-nosed Bat forages in the forest understory, and at edges and gaps in the vegetation.	Horsfield’s Leaf-nosed Bat feeds on insects in the forest understory.	As in many other Hipposideros , females were usually found pregnant in March—May and lactating in April-July. Young were sighted attached to females at the roost in April-June.	Horsfield’s Leaf-nosed Bat can be found roosting in limestone or sandstone caves or man-made underground tunnels. Echolocation calls typically have F components terminating with a FM tail. The range of the frequency of the F part can be variable between populations (89-100 kHz).	Horsfield’s Leaf-nosed Bat forms small to very large colonies of up to 800 individuals, in caves. It is often found in the same cave with other species; for example, in Thailand it is found together with the Diadem Leaf-nosed Bat (77. diadema ), Pendlebury’s Leaf-nosed Bat (77. pendleburyi ), and the Great Himalayan Leaf-nosed Bat (77. armigef). Local movements of this species have been observed in several places in Myanmar and Thailand . Colonies are usually dramatically reduced, or totally missing from the known roosts from December to April, the bats returning in May orJune; this may reflect movements to the breeding colony.	Classified as Least Concern on 77ze IUCN Red List, but some populations could be at risk locally due to cave-bat hunting by local people. Horsfield’s Leaf-nosed Bat is a common species in limestone habitats, and populations appear to be generally stable. It is recorded in several protected areas.	Bates, Bumrungsri, Suyanto, Molur & Srinivasulu (2008) | Corbet & Hill (1992) | Francis (2008a) | Kitchener & Maryanto (1993) | Lekagul & McNeely (1988) | Simmons (2005)	https://zenodo.org/record/6470409/files/figure.png	32. Horsfield’s Leaf-nosed Bat Hipposideros larvatus French: Phyllorhine de Horsfield / German: Horsfield-Rundblattnase I Spanish: Hiposidérido de Horsfield Other common names: Horsfield’s Roundleaf Bat , Intermediate Leaf-nosed Bat Taxonomy. Rhinolophus larvatus Horsfield, 1823 , Java , Indonesia . Hipposideros larvatus is in the larvatus species group. Clarification is needed of differentiating characters and geographic limits with H. grandis , particularly the taxon leptophyllus. Monotypic. Distribution. NE India , NE Bangladesh , S China (including Hainan I), mainland SE Asia, Sumatra (including Simeulue, Nias, and Mentawai Is), Borneo, Java, Kangean, Bali, and many offshore Is. Descriptive notes. Head-body 52-75 mm, tail 32-35 mm, ear 21-25 mm, hindfoot 7—12 mm , forearm 50—67 mm ; weight 15—23 g . Greatest skull lengths are 20-4— 22-8 mm, tibia 18-4—24-8 mm. Horsfield’s Leafnosed Bat is very similar to the Grand Leaf-nosed Bat (77. grandis ) but smaller. Ears are broad with pointed tip. There are three supplementary leaflets on each side of noseleaf. Posterior noseleaf is as broad as width of anterior leaf. Thickened and swollen structure behind posterior leaf is well defined in males. Pelage is dark gray-brown to orange-reddish brown. In skull, rostral chambers are well inflated. Sagittal crest is well developed. C1 and C1 are large. Chromosomal complement has 2n = 32 and FN = 60. Habitat. Horsfield’s Leaf-nosed Bat forages in the forest understory, and at edges and gaps in the vegetation. Food and Feeding. Horsfield’s Leaf-nosed Bat feeds on insects in the forest understory. Breeding. As in many other Hipposideros , females were usually found pregnant in March—May and lactating in April-July. Young were sighted attached to females at the roost in April-June. Activity patterns. Horsfield’s Leaf-nosed Bat can be found roosting in limestone or sandstone caves or man-made underground tunnels. Echolocation calls typically have F components terminating with a FM tail. The range of the frequency of the F part can be variable between populations (89-100 kHz). Movements, Home range and Social organization. Horsfield’s Leaf-nosed Bat forms small to very large colonies of up to 800 individuals, in caves. It is often found in the same cave with other species; for example, in Thailand it is found together with the Diadem Leaf-nosed Bat (77. diadema ), Pendlebury’s Leaf-nosed Bat (77. pendleburyi ), and the Great Himalayan Leaf-nosed Bat (77. armigef). Local movements of this species have been observed in several places in Myanmar and Thailand . Colonies are usually dramatically reduced, or totally missing from the known roosts from December to April, the bats returning in May orJune; this may reflect movements to the breeding colony. Status and Conservation. Classified as Least Concern on 77ze IUCN Red List, but some populations could be at risk locally due to cave-bat hunting by local people. Horsfield’s Leaf-nosed Bat is a common species in limestone habitats, and populations appear to be generally stable. It is recorded in several protected areas. Bibliography. Bates, Bumrungsri, Suyanto, Molur & Srinivasulu (2008), Corbet & Hill (1992), Francis (2008a), Kitchener & Maryanto (1993), Lekagul & McNeely (1988), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Hipposideros larvatus	Hipposideros		larvatus	Horsfield	1823	1	Zool. Res. Java	6: Rhinolophus larvatus , pl. and 10 unno. pp	Intermediate Leaf-nosed Bat	 deformis Horsfield, 1823; insignis Horsfield, 1823; vulgaris Horsfield, 1823; <b> barbensis </b> Miller, 1900; <b> leptophyllus </b> Dobson, 1874; <b> neglectus </b> Sody, 1936; <b>poutensis</b> Allen, 1906.	Indonesia, Java.	N and E India and Bangladesh; Southernmost China; Burma, Thailand, Cambodia, Laos, and Vietnam; W Malaysia to Sumatra, Java, Borneo, and adjacent small islands including Kangean Isls (Indonesia).	Not listed.	Least Concern	 larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018).Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Hipposideros larvatus	23	Horsfield's Leaf-nosed Bat	Horsfield's Roundleaf Bat|Intermediate Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Hipposideros	NA	larvatus	Horsfield	1823	1	Rhinolophus_larvatus	Horsfield, T. (1823). Zoological researches in Java, and the neighboring islands. Printed for Kingsbury, Parbury, & Allen, 1824, London, unpaginated.	https://www.biodiversitylibrary.org/page/31111887#page/71/mode/1up	BM 1879.11.21.93		Java, Indonesia.			larvatus (Horsfield, 1823)|deformis (Horsfield, 1823)|insignis (Horsfield, 1823)|vulgaris (Horsfield, 1823)|leptophyllus (Dobson, 1874)|barbensis G. S. Miller, 1900|poutensis J. A. Allen, 1906|neglectus Sody, 1936|khasiana Thabah, Rossiter, Kingston, Zhang Shuyi, Parsons, Mya Mya Khin, Akbar, & G. Jones, 2006 [nomen nudum]	includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form	Thabah, A., Rossiter, S. J., Kingston, T., Zhang, S., Parsons, S., Mya, K. M., ... & Jones, G. (2006). Genetic divergence and echolocation call frequency in cryptic species of Hipposideros larvatus sl.(Chiroptera: Hipposideridae) from the Indo-Malayan region. Biological Journal of the Linnean Society, 88(1), 119-130.|Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.	India|Bangladesh|China|Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_larvatus	0	sciname match	Hipposideros_larvatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Hipposideros larvatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Hipposideros	larvatus	(Horsfield, 1823)	Excludes ;Hipposideros ;<span class="SpellE">grandis</span> , now considered separate species (Thabah ;et al. 2006, ;Kruskop</span> ;2013).	20000000	Hipposideros larvatus	Least Concern		2020	2020-07-05 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In South Asia, this species roosts in caves, mineshafts, pagodas, and buildings (Molur et al. 2002, Thabah et al. 2006, Saikia et al. 2018). In Myanmar, it is found in a variety of habitats from highly disturbed agricultural land to secondary forest, in dry zones and humid areas (P. Bates pers. comm., K.M. Swe pers. comm., Bates and Harrison 1997). It is often associated with limestone caves in Myanmar and Indonesia (A. Suyanto pers. comm.). On Peninsular Malaysia, the species has been recorded from primary lowland tropical moist forest. It is also found roosting in human habitations (P. Bates pers. comm.).	There are no major threats to this species as a whole. There currently appear to be no major threats to this species in South Asia. However, disturbance to roosting sites such as caves by humans may pose a threat to this species as observed in Meghalaya (U. Saikia pers. comm.). In neighbouring Myanmar, mining activities in limestone caves for manufacture of cement is a threat to some populations of this species (K.M. Swe pers. comm., Molur et al. 2002). It may be affected by hunting in some parts of its range (C.M. Francis pers. comm.).	This is a common bat throughout much of its range (P. Soisook pers. comm.). In South Asia, this species is distributed across three northeastern states of India, but the sightings are sparse. Only two colonies of less than 100 individuals have been recorded in India (Molur et al. 2002, Thabah et al. 2006, Saikia et al. 2018). In northern Myanmar, it is one of the more common bats and lives in large colonies. However, there is a decline in their numbers due to mining and it is estimated to have declined by 10% over the last 10-15 years (K.M. Swe pers. comm.).	Unknown	This species is widely distributed, ranging from northeastern India, throughout much of southern China, including Hainan Island), and mainland Southeast Asia, into several islands within insular Southeast Asia. In South Asia, this species is found in Bangladesh (Chittagong and Sylhet divisions) and India (Arunachal Pradesh, Assam and Meghalaya) (Molur et al. 2002). In China, it has been recorded from Hainan Island, Guangdong, Guangxi, and Guizhou (Smith and Xie 2008). On mainland Southeast Asia, it is distributed widely across Myanmar (Pipat Soisook pers. comm.), ranging into Thailand, Cambodia, Lao PDR, Viet Nam and Peninsular Malaysia. In insular Southeast Asia, it has been recorded from numerous islands including the Sumatra (Mentawi Islands, Nias Island, Simeulue Island), Java, Bali, the Kangean Islands (all to Indonesia) and the southern part of the island of Borneo (Kalimantan [Indonesia] and possibly Sarawak [Malaysia]). This species has been taken from sea level up to around 2,000 m asl.	In Southeast Asia, this species ;is hunted ;for local consumption and medicinal value.	Terrestrial	This species is present in a number of protected areas in its range. Surveys, habitat management, and population monitoring are important recommendations. Public awareness and outreach is necessary in areas where this taxon faces threat due to mining activities (Molur et al. 2002). Further studies are needed to clarify the taxonomic status of populations currently allocated to this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Hipposideros		larvatus	Horsfield	1823	1	Zool. Res. Java	6: Rhinolophus larvatus , pl. and 10 unno. pp	Intermediate Leaf-nosed Bat	 deformis Horsfield, 1823; insignis Horsfield, 1823; vulgaris Horsfield, 1823; <b> barbensis </b> Miller, 1900; <b> leptophyllus </b> Dobson, 1874; <b> neglectus </b> Sody, 1936; <b>poutensis</b> Allen, 1906.	Indonesia, Java.	N and E India and Bangladesh; Myanmar, Thailand, Cambodia, Laos, and Vietnam; W Malaysia to Sumatra, Java, Borneo, and adjacent small islands including Kangean Isls (Indonesia).	Not listed.	Least Concern	 larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018; Yuzefovich et al., 2022). Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. Does not include poutensis ; see Yuzefovich et al. (2022). See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993). Includes khasiana (Thabah et al., 2006) as a nomen nudum as no specific holotype was designated.	Hipposideros larvatus	1004613	23	Horsfield's Leaf-nosed Bat	Horsfield's Roundleaf Bat|Intermediate Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Hipposideros	NA	larvatus	Horsfield	1823	1	Rhinolophus_larvatus	Horsfield, T. (1823). Zoological researches in Java, and the neighboring islands. Printed for Kingsbury, Parbury, & Allen, 1824, London, unpaginated.	https://www.biodiversitylibrary.org/page/31111887#page/71/mode/1up	BM 1879.11.21.93		Java, Indonesia.			larvatus (Horsfield, 1823)|deformis (Horsfield, 1823)|insignis (Horsfield, 1823)|vulgaris (Horsfield, 1823)|leptophyllus (Dobson, 1874)|barbensis G. S. Miller, 1900|neglectus Sody, 1936|khasiana Thabah, Rossiter, Kingston, Zhang Shuyi, Parsons, Mya Mya Khin, Akbar, & G. Jones, 2006 [nomen nudum]	previously included H. poutensis and includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; the species limits between H. grandis, H. larvatus, and H. poutensis is currently unresolved and represents a species complex	Thabah, A., Rossiter, S. J., Kingston, T., Zhang, S., Parsons, S., Mya, K. M., ... & Jones, G. (2006). Genetic divergence and echolocation call frequency in cryptic species of Hipposideros larvatus sl.(Chiroptera: Hipposideridae) from the Indo-Malayan region. Biological Journal of the Linnean Society, 88(1), 119-130.|Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.|Yuzefovich, A. P., Artyushin, I. V., Skopin, A. E., Son, N. T., & Kruskop, S. V. (2022). Taxonomic diversity of the Hipposideros larvatus species complex (Chiroptera: Hipposideridae) in mainland Asia. Zootaxa, 5200(1), 73-95.				India|Bangladesh|Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_larvatus	0	sciname match	Hipposideros_larvatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Hipposideros_larvatus	1004613	23	Horsfield's Roundleaf Bat	Horsfield's Roundleaf Bat|Intermediate Leaf-nosed Bat|Horsfield's Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Hipposideros	NA	larvatus	Horsfield	1	Rhinolophus larvatus	Horsfield, T. 1823-06. No. VI. _Cervus Muntjak_. _Viverra Rasse_. _Rhinolophus larvatus_. _Rhinoceros sondaicus_. _Iora scapularis_. _Falco Limnæetus_. _Oriolus xanthonotus_. _Centropus Philippensis_. in Horsfield, T. 1824. Zoological Researches in Java, and the Neighbouring Islands. Kingsbury, Parbury, & Allen, London, not continuously paginated pp.	https://www.biodiversitylibrary.org/page/31111856	BMNH:Mamm:1879.11.21.93	holotype	https://data.nhm.ac.uk/object/f46fefee-278e-4dfd-b550-619dca25d2c8	Java, Indonesia.			previously included H. poutensis and includes the recently described khasiana, which is a nomen nudum as the publication in which the name was first used does not include a proper description, holotype designation, or type locality; the khasiana form likely represents a distinct species, but further studies are needed to both confirm this and name the form; the species limits between H. grandis, H. larvatus, and H. poutensis is currently unresolved and represents a species complex	Thabah, A., Rossiter, S. J., Kingston, T., Zhang, S., Parsons, S., Mya, K. M., ... & Jones, G. (2006). Genetic divergence and echolocation call frequency in cryptic species of Hipposideros larvatus sl.(Chiroptera: Hipposideridae) from the Indo-Malayan region. Biological Journal of the Linnean Society, 88(1), 119-130.|Wilson, D. E. & Mittermeier, R. A.Handbook of the Mammals of the World. 9. Bats. Lynx Edicions, Barcelona.|Yuzefovich, A. P., Artyushin, I. V., Skopin, A. E., Son, N. T., & Kruskop, S. V. (2022). Taxonomic diversity of the Hipposideros larvatus species complex (Chiroptera: Hipposideridae) in mainland Asia. Zootaxa, 5200(1), 73-95.				India|Bangladesh|Myanmar|Vietnam|Laos|Cambodia|Thailand|Malaysia|Indonesia	Asia	Indomalaya|Palearctic	LC	0	0	0	Hipposideros_larvatus	0	sciname match	Hipposideros_larvatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Hipposideros		larvatus	Horsfield	1823	1	Zool. Res. Java	6: Rhinolophus larvatus, pl. and 10 unno. pp	Intermediate Leaf-nosed Bat	deformis Horsfield, 1823; insignis Horsfield, 1823; vulgaris Horsfield, 1823; barbensis Miller, 1900; leptophyllus Dobson, 1874; neglectus Sody, 1936; poutensis Allen, 1906.	Indonesia, Java.	N and E India and Bangladesh; Myanmar, Thailand, Cambodia, Laos, and Vietnam; W Malaysia to Sumatra, Java, Borneo, and adjacent small islands including Kangean Isls (Indonesia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85646564/22091287/' target='_blank'>Least Concern</a>	larvatus species group. Appears to be a complex that includes several distinct species (S. Tsang, pers. comm., 2018; Yuzefovich et al., 2022). Does not include grandis and sumbae; see Kitchner and Maryantu (1993), who revised this complex. Does not include poutensis; see Yuzefovich et al. (2022). See also Hill(1963) and Sinha (1999). Subspecies limits and validity are uncertain. Does not include alongensis, see Topál (1993). Includes khasiana (Thabah et al., 2006) as a nomen nudum as no specific holotype was designated.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Hipposideros larvatus; Hipposideros larvatus; Hipposideros larvatus; Hipposideros larvatus; Hipposideros larvatus; larvatus; barbensis; leptophyllus; neglectus; poutensis; deformis; insignis; vulgaris; barbensis; leptophyllus; neglectus; poutensis; deformis; insignis; vulgaris; larvatus; deformis; insignis; vulgaris; leptophyllus; barbensis; poutensis; neglectus; khasiana; Phyllorhine de Horsfield; Horsfield-Rundblattnase; Hiposidérido de Horsfield; Horsfield’s Roundleaf Bat; ntermediate Leaf-nosed Bat; Horsfield's Leaf-nosed Bat; Horsfield's Roundleaf Bat; Intermediate Leaf-nosed Bat; Intermediate Leaf-nosed Bat; Intermediate Leaf-nosed Bat; H. larvatus